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83-559: Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are a part of the subphylum Crustacea ( / k r ə ˈ s t eɪ ʃ ə / ), a large, diverse group of mainly aquatic arthropods including decapods ( shrimps , prawns , crabs , lobsters and crayfish ), seed shrimp , branchiopods , fish lice , krill , remipedes , isopods , barnacles , copepods , opossum shrimps , amphipods and mantis shrimp . The crustacean group can be treated as

166-526: A faster pace. It is characterised by greater use of prepositions, and word order that is closer to modern Romance languages, for example, while grammatically retaining more or less the same formal rules as Classical Latin. Ultimately, Latin diverged into a distinct written form, where the commonly spoken form was perceived as a separate language, for instance early French or Italian dialects, that could be transcribed differently. It took some time for these to be viewed as wholly different from Latin however. After

249-718: A few in German , Dutch , Norwegian , Danish and Swedish . Latin is still spoken in Vatican City, a city-state situated in Rome that is the seat of the Catholic Church . The works of several hundred ancient authors who wrote in Latin have survived in whole or in part, in substantial works or in fragments to be analyzed in philology . They are in part the subject matter of the field of classics . Their works were published in manuscript form before

332-404: A few. Famous and well regarded writers included Petrarch, Erasmus, Salutati , Celtis , George Buchanan and Thomas More . Non fiction works were long produced in many subjects, including the sciences, law, philosophy, historiography and theology. Famous examples include Isaac Newton 's Principia . Latin was also used as a convenient medium for translations of important works first written in

415-464: A larger Pancrustacea clade . The traditional classification of Crustacea based on morphology recognised four to six classes. Bowman and Abele (1982) recognised 652 extant families and 38 orders, organised into six classes: Branchiopoda , Remipedia , Cephalocarida , Maxillopoda, Ostracoda , and Malacostraca . Martin and Davis (2001) updated this classification, retaining the six classes but including 849 extant families in 42 orders. Despite outlining

498-463: A limbless abdomen, except from a telson and caudal rami which is present in many groups. The abdomen in malacostracans bears pleopods , and ends in a telson, which bears the anus , and is often flanked by uropods to form a tail fan . The number and variety of appendages in different crustaceans may be partly responsible for the group's success. Crustacean appendages are typically biramous , meaning they are divided into two parts; this includes

581-548: A native language, Medieval Latin was used across Western and Catholic Europe during the Middle Ages as a working and literary language from the 9th century to the Renaissance , which then developed a classicizing form, called Renaissance Latin . This was the basis for Neo-Latin which evolved during the early modern period . In these periods Latin was used productively and generally taught to be written and spoken, at least until

664-551: A result, the list has variants, as well as alternative names. In addition to the historical phases, Ecclesiastical Latin refers to the styles used by the writers of the Roman Catholic Church from late antiquity onward, as well as by Protestant scholars. The earliest known form of Latin is Old Latin, also called Archaic or Early Latin, which was spoken from the Roman Kingdom , traditionally founded in 753 BC, through

747-407: A separate language, existing more or less in parallel with the literary or educated Latin, but this is now widely dismissed. The term 'Vulgar Latin' remains difficult to define, referring both to informal speech at any time within the history of Latin, and the kind of informal Latin that had begun to move away from the written language significantly in the post-Imperial period, that led ultimately to

830-695: A small number of Latin services held in the Anglican church. These include an annual service in Oxford, delivered with a Latin sermon; a relic from the period when Latin was the normal spoken language of the university. In the Western world, many organizations, governments and schools use Latin for their mottos due to its association with formality, tradition, and the roots of Western culture . Canada's motto A mari usque ad mare ("from sea to sea") and most provincial mottos are also in Latin. The Canadian Victoria Cross

913-429: A sort of informal language academy dedicated to maintaining and perpetuating educated speech. Philological analysis of Archaic Latin works, such as those of Plautus , which contain fragments of everyday speech, gives evidence of an informal register of the language, Vulgar Latin (termed sermo vulgi , "the speech of the masses", by Cicero ). Some linguists, particularly in the nineteenth century, believed this to be

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996-485: A spiral format. Structures that function as kidneys are located near the antennae. A brain exists in the form of ganglia close to the antennae, and a collection of major ganglia is found below the gut. In many decapods , the first (and sometimes the second) pair of pleopods are specialised in the male for sperm transfer. Many terrestrial crustaceans (such as the Christmas Island red crab ) mate seasonally and return to

1079-565: A spoken and written language by the scholarship by the Renaissance humanists . Petrarch and others began to change their usage of Latin as they explored the texts of the Classical Latin world. Skills of textual criticism evolved to create much more accurate versions of extant texts through the fifteenth and sixteenth centuries, and some important texts were rediscovered. Comprehensive versions of authors' works were published by Isaac Casaubon , Joseph Scaliger and others. Nevertheless, despite

1162-432: A strictly left-to-right script. During the late republic and into the first years of the empire, from about 75 BC to AD 200, a new Classical Latin arose, a conscious creation of the orators, poets, historians and other literate men, who wrote the great works of classical literature , which were taught in grammar and rhetoric schools. Today's instructional grammars trace their roots to such schools , which served as

1245-575: A subphylum under the clade Mandibulata . It is now well accepted that the hexapods ( insects and entognathans ) emerged deep in the Crustacean group, with the completed pan-group referred to as Pancrustacea . The three classes Cephalocarida , Branchiopoda and Remipedia are more closely related to the hexapods than they are to any of the other crustaceans ( oligostracans and multicrustaceans ). The 67,000 described species range in size from Stygotantulus stocki at 0.1 mm (0.004 in), to

1328-426: A tergite are known as paranota (Greek for "alongside the back") or carinae (Latin for "keel"), exemplified by the flat-backed millipedes of the order Polydesmida . Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods. Tergo-tergal is a stridulatory mechanism in which fine spines of the abdominal tergites are rubbed together to produce sound. This process

1411-689: A vernacular, such as those of Descartes . Latin education underwent a process of reform to classicise written and spoken Latin. Schooling remained largely Latin medium until approximately 1700. Until the end of the 17th century, the majority of books and almost all diplomatic documents were written in Latin. Afterwards, most diplomatic documents were written in French (a Romance language ) and later native or other languages. Education methods gradually shifted towards written Latin, and eventually concentrating solely on reading skills. The decline of Latin education took several centuries and proceeded much more slowly than

1494-411: Is Veritas ("truth"). Veritas was the goddess of truth, a daughter of Saturn, and the mother of Virtue. Switzerland has adopted the country's Latin short name Helvetia on coins and stamps, since there is no room to use all of the nation's four official languages . For a similar reason, it adopted the international vehicle and internet code CH , which stands for Confoederatio Helvetica ,

1577-626: Is a reversal of the original phrase Non terrae plus ultra ("No land further beyond", "No further!"). According to legend , this phrase was inscribed as a warning on the Pillars of Hercules , the rocks on both sides of the Strait of Gibraltar and the western end of the known, Mediterranean world. Charles adopted the motto following the discovery of the New World by Columbus, and it also has metaphorical suggestions of taking risks and striving for excellence. In

1660-436: Is an open circulatory system , where blood is pumped into the haemocoel by a heart located near the dorsum. Malacostraca have haemocyanin as the oxygen-carrying pigment, while copepods, ostracods, barnacles and branchiopods have haemoglobins . The alimentary canal consists of a straight tube that often has a gizzard-like "gastric mill" for grinding food and a pair of digestive glands that absorb food; this structure goes in

1743-548: Is found in any widespread language, the languages of Spain, France, Portugal, and Italy have retained a remarkable unity in phonological forms and developments, bolstered by the stabilising influence of their common Christian (Roman Catholic) culture. It was not until the Muslim conquest of Spain in 711, cutting off communications between the major Romance regions, that the languages began to diverge seriously. The spoken Latin that would later become Romanian diverged somewhat more from

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1826-661: Is modelled after the British Victoria Cross which has the inscription "For Valour". Because Canada is officially bilingual, the Canadian medal has replaced the English inscription with the Latin Pro Valore . Spain's motto Plus ultra , meaning "even further", or figuratively "Further!", is also Latin in origin. It is taken from the personal motto of Charles V , Holy Roman Emperor and King of Spain (as Charles I), and

1909-517: Is produced in Asia, with China alone producing nearly half the world's total. Non-decapod crustaceans are not widely consumed, with only 118,000 tons of krill being caught, despite krill having one of the greatest biomasses on the planet. Latin Latin ( lingua Latina , pronounced [ˈlɪŋɡʷa ɫaˈtiːna] , or Latinum [ɫaˈtiːnʊ̃] ) is a classical language belonging to

1992-666: Is some debate as to whether or not Cambrian animals assigned to Ostracoda are truly ostracods , which would otherwise start in the Ordovician . The only classes to appear later are the Cephalocarida , which have no fossil record, and the Remipedia , which were first described from the fossil Tesnusocaris goldichi , but do not appear until the Carboniferous . Most of the early crustaceans are rare, but fossil crustaceans become abundant from

2075-958: Is taught at many high schools, especially in Europe and the Americas. It is most common in British public schools and grammar schools, the Italian liceo classico and liceo scientifico , the German Humanistisches Gymnasium and the Dutch gymnasium . Occasionally, some media outlets, targeting enthusiasts, broadcast in Latin. Notable examples include Radio Bremen in Germany, YLE radio in Finland (the Nuntii Latini broadcast from 1989 until it

2158-419: Is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. A given tergum may be divided into hardened plates or sclerites commonly referred to as tergites . In a thoracic segment, for example, the tergum may be divided into an anterior notum and a posterior scutellum . Lateral extensions of

2241-518: Is thought to be just 1 ⁄ 10 to 1 ⁄ 100 of the total number as most species remain as yet undiscovered . Although most crustaceans are small, their morphology varies greatly and includes both the largest arthropod in the world – the Japanese spider crab with a leg span of 3.7 metres (12 ft) – and the smallest, the 100- micrometre -long (0.004 in) Stygotantulus stocki . Despite their diversity of form, crustaceans are united by

2324-531: The Corpus Inscriptionum Latinarum (CIL). Authors and publishers vary, but the format is about the same: volumes detailing inscriptions with a critical apparatus stating the provenance and relevant information. The reading and interpretation of these inscriptions is the subject matter of the field of epigraphy . About 270,000 inscriptions are known. The Latin influence in English has been significant at all stages of its insular development. In

2407-608: The Marmorkrebs crayfish. In many crustaceans, the fertilised eggs are released into the water column , while others have developed a number of mechanisms for holding on to the eggs until they are ready to hatch. Most decapods carry the eggs attached to the pleopods , while peracarids , notostracans , anostracans , and many isopods form a brood pouch from the carapace and thoracic limbs. Female Branchiura do not carry eggs in external ovisacs but attach them in rows to rocks and other objects. Most leptostracans and krill carry

2490-690: The Carboniferous , as are the first true mantis shrimp. In the Decapoda , prawns and polychelids appear in the Triassic, and shrimp and crabs appear in the Jurassic . The fossil burrow Ophiomorpha is attributed to ghost shrimps, whereas the fossil burrow Camborygma is attributed to crayfishes. The Permian–Triassic deposits of Nurra preserve the oldest (Permian: Roadian) fluvial burrows ascribed to ghost shrimps (Decapoda: Axiidea, Gebiidea) and crayfishes (Decapoda: Astacidea, Parastacidea), respectively. However,

2573-461: The Carboniferous period onwards. Within the Malacostraca, no fossils are known for krill , while both Hoplocarida and Phyllopoda contain important groups that are now extinct as well as extant members (Hoplocarida: mantis shrimp are extant, while Aeschronectida are extinct; Phyllopoda: Canadaspidida are extinct, while Leptostraca are extant). Cumacea and Isopoda are both known from

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2656-562: The Holy See , the primary language of its public journal , the Acta Apostolicae Sedis , and the working language of the Roman Rota . Vatican City is also home to the world's only automatic teller machine that gives instructions in Latin. In the pontifical universities postgraduate courses of Canon law are taught in Latin, and papers are written in the same language. There are

2739-818: The Italic branch of the Indo-European languages . Latin was originally spoken by the Latins in Latium (now known as Lazio ), the lower Tiber area around Rome , Italy. Through the expansion of the Roman Republic it became the dominant language in the Italian Peninsula and subsequently throughout the Roman Empire . By the late Roman Republic , Old Latin had evolved into standardized Classical Latin . Vulgar Latin refers to

2822-401: The Japanese spider crab with a leg span of up to 3.8 m (12.5 ft) and a mass of 20 kg (44 lb). Like other arthropods , crustaceans have an exoskeleton , which they moult to grow. They are distinguished from other groups of arthropods, such as insects , myriapods and chelicerates , by the possession of biramous (two-parted) limbs, and by their larval forms , such as

2905-561: The Middle Ages , borrowing from Latin occurred from ecclesiastical usage established by Saint Augustine of Canterbury in the 6th century or indirectly after the Norman Conquest , through the Anglo-Norman language . From the 16th to the 18th centuries, English writers cobbled together huge numbers of new words from Latin and Greek words, dubbed " inkhorn terms ", as if they had spilled from a pot of ink. Many of these words were used once by

2988-559: The Roman Rite of the Catholic Church at the Vatican City . The church continues to adapt concepts from modern languages to Ecclesiastical Latin of the Latin language. Contemporary Latin is more often studied to be read rather than spoken or actively used. Latin has greatly influenced the English language , along with a large number of others, and historically contributed many words to

3071-553: The Romance languages . During the Classical period, informal language was rarely written, so philologists have been left with only individual words and phrases cited by classical authors, inscriptions such as Curse tablets and those found as graffiti . In the Late Latin period, language changes reflecting spoken (non-classical) norms tend to be found in greater quantities in texts. As it

3154-622: The Western Roman Empire fell in 476 and Germanic kingdoms took its place, the Germanic people adopted Latin as a language more suitable for legal and other, more formal uses. While the written form of Latin was increasingly standardized into a fixed form, the spoken forms began to diverge more greatly. Currently, the five most widely spoken Romance languages by number of native speakers are Spanish , Portuguese , French , Italian , and Romanian . Despite dialectal variation, which

3237-628: The nauplius stage of branchiopods and copepods . Most crustaceans are free-living aquatic animals , but some are terrestrial (e.g. woodlice , sandhoppers ), some are parasitic (e.g. Rhizocephala , fish lice , tongue worms ) and some are sessile (e.g. barnacles ). The group has an extensive fossil record , reaching back to the Cambrian . More than 7.9 million tons of crustaceans per year are harvested by fishery or farming for human consumption, consisting mostly of shrimp and prawns . Krill and copepods are not as widely fished, but may be

3320-409: The pereon or thorax , and the pleon or abdomen . The head and thorax may be fused together to form a cephalothorax , which may be covered by a single large carapace . The crustacean body is protected by the hard exoskeleton , which must be moulted for the animal to grow. The shell around each somite can be divided into a dorsal tergum , ventral sternum and a lateral pleuron. Various parts of

3403-512: The zoea (pl. zoeæ or zoeas). This name was given to it when naturalists believed it to be a separate species. It follows the nauplius stage and precedes the post-larva . Zoea larvae swim with their thoracic appendages , as opposed to nauplii, which use cephalic appendages, and megalopa, which use abdominal appendages for swimming. It often has spikes on its carapace , which may assist these small organisms in maintaining directional swimming. In many decapods , due to their accelerated development,

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3486-617: The British Crown. The motto is featured on all presently minted coinage and has been featured in most coinage throughout the nation's history. Several states of the United States have Latin mottos , such as: Many military organizations today have Latin mottos, such as: Some law governing bodies in the Philippines have Latin mottos, such as: Some colleges and universities have adopted Latin mottos, for example Harvard University 's motto

3569-599: The English lexicon , particularly after the Christianization of the Anglo-Saxons and the Norman Conquest . Latin and Ancient Greek roots are heavily used in English vocabulary in theology , the sciences , medicine , and law . A number of phases of the language have been recognized, each distinguished by subtle differences in vocabulary, usage, spelling, and syntax. There are no hard and fast rules of classification; different scholars emphasize different features. As

3652-580: The Grinch Stole Christmas! , The Cat in the Hat , and a book of fairy tales, " fabulae mirabiles ", are intended to garner popular interest in the language. Additional resources include phrasebooks and resources for rendering everyday phrases and concepts into Latin, such as Meissner's Latin Phrasebook . Some inscriptions have been published in an internationally agreed, monumental, multivolume series,

3735-681: The Hexapoda is distinctly closer to e.g. a Multicrustacean than an Oligostracan is. Crustaceans have a rich and extensive fossil record , which begins with animals such as Canadaspis and Perspicaris from the Middle Cambrian age Burgess Shale . Most of the major groups of crustaceans appear in the fossil record before the end of the Cambrian, namely the Branchiopoda , Maxillopoda (including barnacles and tongue worms ) and Malacostraca ; there

3818-461: The United States the unofficial national motto until 1956 was E pluribus unum meaning "Out of many, one". The motto continues to be featured on the Great Seal . It also appears on the flags and seals of both houses of congress and the flags of the states of Michigan, North Dakota, New York, and Wisconsin. The motto's 13 letters symbolically represent the original Thirteen Colonies which revolted from

3901-552: The University of Kentucky, the University of Oxford and also Princeton University. There are many websites and forums maintained in Latin by enthusiasts. The Latin Misplaced Pages has more than 130,000 articles. Italian , French , Portuguese , Spanish , Romanian , Catalan , Romansh , Sardinian and other Romance languages are direct descendants of Latin. There are also many Latin borrowings in English and Albanian , as well as

3984-406: The animals with the greatest biomass on the planet, and form a vital part of the food chain. The scientific study of crustaceans is known as carcinology (alternatively, malacostracology , crustaceology or crustalogy ), and a scientist who works in carcinology is a carcinologist . The body of a crustacean is composed of segments, which are grouped into three regions: the cephalon or head ,

4067-501: The author and then forgotten, but some useful ones survived, such as 'imbibe' and 'extrapolate'. Many of the most common polysyllabic English words are of Latin origin through the medium of Old French . Romance words make respectively 59%, 20% and 14% of English, German and Dutch vocabularies. Those figures can rise dramatically when only non-compound and non-derived words are included. Tergum A tergum (Latin for "the back"; pl. : terga , associated adjective tergal )

4150-425: The benefit of those who do not understand Latin. There are also songs written with Latin lyrics . The libretto for the opera-oratorio Oedipus rex by Igor Stravinsky is in Latin. Parts of Carl Orff 's Carmina Burana are written in Latin. Enya has recorded several tracks with Latin lyrics. The continued instruction of Latin is seen by some as a highly valuable component of a liberal arts education. Latin

4233-409: The careful work of Petrarch, Politian and others, first the demand for manuscripts, and then the rush to bring works into print, led to the circulation of inaccurate copies for several centuries following. Neo-Latin literature was extensive and prolific, but less well known or understood today. Works covered poetry, prose stories and early novels, occasional pieces and collections of letters, to name

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4316-415: The classicised Latin that followed through to the present are often grouped together as Neo-Latin , or New Latin, which have in recent decades become a focus of renewed study , given their importance for the development of European culture, religion and science. The vast majority of written Latin belongs to this period, but its full extent is unknown. The Renaissance reinforced the position of Latin as

4399-461: The country's full Latin name. Some film and television in ancient settings, such as Sebastiane , The Passion of the Christ and Barbarians (2020 TV series) , have been made with dialogue in Latin. Occasionally, Latin dialogue is used because of its association with religion or philosophy, in such film/television series as The Exorcist and Lost (" Jughead "). Subtitles are usually shown for

4482-493: The decline in written Latin output. Despite having no native speakers, Latin is still used for a variety of purposes in the contemporary world. The largest organisation that retains Latin in official and quasi-official contexts is the Catholic Church . The Catholic Church required that Mass be carried out in Latin until the Second Vatican Council of 1962–1965 , which permitted the use of the vernacular . Latin remains

4565-570: The educated and official world, Latin continued without its natural spoken base. Moreover, this Latin spread into lands that had never spoken Latin, such as the Germanic and Slavic nations. It became useful for international communication between the member states of the Holy Roman Empire and its allies. Without the institutions of the Roman Empire that had supported its uniformity, Medieval Latin

4648-447: The eggs between their thoracic limbs; some copepods carry their eggs in special thin-walled sacs, while others have them attached together in long, tangled strings. Crustaceans exhibit a number of larval forms, of which the earliest and most characteristic is the nauplius . This has three pairs of appendages , all emerging from the young animal's head, and a single naupliar eye. In most groups, there are further larval stages, including

4731-672: The evidence that Maxillopoda was non-monophyletic, they retained it as one of the six classes, although did suggest that Maxillipoda could be replaced by elevating its subclasses to classes. Since then phylogenetic studies have confirmed the polyphyly of Maxillipoda and the paraphyletic nature of Crustacea with respect to Hexapoda. Recent classifications recognise ten to twelve classes in Crustacea or Pancrustacea, with several former maxillopod subclasses now recognised as classes (e.g. Thecostraca , Tantulocarida , Mystacocarida , Copepoda , Branchiura and Pentastomida ). The following cladogram shows

4814-440: The exoskeleton may be fused together. Each somite , or body segment can bear a pair of appendages : on the segments of the head, these include two pairs of antennae , the mandibles and maxillae ; the thoracic segments bear legs , which may be specialised as pereiopods (walking legs) and maxillipeds (feeding legs). Malacostraca and Remipedia (and the hexapods) have abdominal appendages. All other classes of crustaceans have

4897-451: The females are algae-fed instead of yeast-fed. A small number are hermaphrodites , including barnacles , remipedes , and Cephalocarida . Some may even change sex during the course of their life. Parthenogenesis is also widespread among crustaceans, where viable eggs are produced by a female without needing fertilisation by a male. This occurs in many branchiopods , some ostracods , some isopods , and certain "higher" crustaceans, such as

4980-659: The great radiation of crustaceans occurred in the Cretaceous , particularly in crabs, and may have been driven by the adaptive radiation of their main predators, bony fish . The first true lobsters also appear in the Cretaceous. Many crustaceans are consumed by humans, and nearly 10,700,000  tons were harvested in 2007; the vast majority of this output is of decapod crustaceans : crabs , lobsters , shrimp , crawfish , and prawns . Over 60% by weight of all crustaceans caught for consumption are shrimp and prawns, and nearly 80%

5063-679: The invention of printing and are now published in carefully annotated printed editions, such as the Loeb Classical Library , published by Harvard University Press , or the Oxford Classical Texts , published by Oxford University Press . Latin translations of modern literature such as: The Hobbit , Treasure Island , Robinson Crusoe , Paddington Bear , Winnie the Pooh , The Adventures of Tintin , Asterix , Harry Potter , Le Petit Prince , Max and Moritz , How

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5146-675: The language of the Roman Rite . The Tridentine Mass (also known as the Extraordinary Form or Traditional Latin Mass) is celebrated in Latin. Although the Mass of Paul VI (also known as the Ordinary Form or the Novus Ordo) is usually celebrated in the local vernacular language, it can be and often is said in Latin, in part or in whole, especially at multilingual gatherings. It is the official language of

5229-440: The large areas where it had come to be natively spoken. However, even after the fall of Western Rome , Latin remained the common language of international communication , science, scholarship and academia in Europe until well into the early 19th century, by which time modern languages had supplanted it in common academic and political usage. Late Latin is the literary language from the 3rd century AD onward. No longer spoken as

5312-462: The late seventeenth century, when spoken skills began to erode. It then became increasingly taught only to be read. Latin grammar is highly fusional , with classes of inflections for case , number , person , gender , tense , mood , voice , and aspect . The Latin alphabet is directly derived from the Etruscan and Greek alphabets . Latin remains the official language of the Holy See and

5395-431: The later part of the Roman Republic , up to 75 BC, i.e. before the age of Classical Latin . It is attested both in inscriptions and in some of the earliest extant Latin literary works, such as the comedies of Plautus and Terence . The Latin alphabet was devised from the Etruscan alphabet . The writing later changed from what was initially either a right-to-left or a boustrophedon script to what ultimately became

5478-420: The layer allow light to reach the retina. As the larvae mature into adults, the layer migrates to a new position behind the retina where it works as a backscattering mirror that increases the intensity of light passing through the eyes, as seen in many nocturnal animals. In an effort to understand whether DNA repair processes can protect crustaceans against DNA damage , basic research was conducted to elucidate

5561-421: The less prestigious colloquial registers , attested in inscriptions and some literary works such as those of the comic playwrights Plautus and Terence and the author Petronius . While often called a "dead language", Latin did not undergo language death . By the 6th to 9th centuries, natural language change eventually resulted in Latin as a vernacular language evolving into distinct Romance languages in

5644-399: The oceans as insects are on land. Most crustaceans are also motile , moving about independently, although a few taxonomic units are parasitic and live attached to their hosts (including sea lice , fish lice , whale lice , tongue worms , and Cymothoa exigua , all of which may be referred to as "crustacean lice"), and adult barnacles live a sessile life – they are attached headfirst to

5727-596: The opening of the Suez Canal , close to 100 species of crustaceans from the Red Sea and the Indo-Pacific realm have established themselves in the eastern Mediterranean sub-basin, with often significant impact on local ecosystems. Most crustaceans have separate sexes , and reproduce sexually . In fact, a recent study explains how the male T. californicus decide which females to mate with by dietary differences, preferring when

5810-413: The other varieties, as it was largely separated from the unifying influences in the western part of the Empire. Spoken Latin began to diverge into distinct languages by the 9th century at the latest, when the earliest extant Romance writings begin to appear. They were, throughout the period, confined to everyday speech, as Medieval Latin was used for writing. For many Italians using Latin, though, there

5893-428: The repair mechanisms used by Penaeus monodon (black tiger shrimp). Repair of DNA double-strand breaks was found to be predominantly carried out by accurate homologous recombinational repair. Another, less accurate process, microhomology-mediated end joining , is also used to repair such breaks. The expression pattern of DNA repair related and DNA damage response genes in the intertidal copepod Tigriopus japonicus

5976-445: The sea to release the eggs. Others, such as woodlice , lay their eggs on land, albeit in damp conditions. In most decapods, the females retain the eggs until they hatch into free-swimming larvae. Most crustaceans are aquatic, living in either marine or freshwater environments, but a few groups have adapted to life on land, such as terrestrial crabs , terrestrial hermit crabs , and woodlice . Marine crustaceans are as ubiquitous in

6059-507: The second pair of antennae, but not the first, which is usually uniramous , the exception being in the Class Malacostraca where the antennules may be generally biramous or even triramous. It is unclear whether the biramous condition is a derived state which evolved in crustaceans, or whether the second branch of the limb has been lost in all other groups. Trilobites , for instance, also possessed biramous appendages. The main body cavity

6142-471: The special larval form known as the nauplius . The exact relationships of the Crustacea to other taxa are not completely settled as of April 2012. Studies based on morphology led to the Pancrustacea hypothesis, in which Crustacea and Hexapoda ( insects and allies) are sister groups . More recent studies using DNA sequences suggest that Crustacea is paraphyletic , with the hexapods nested within

6225-599: The substrate and cannot move independently. Some branchiurans are able to withstand rapid changes of salinity and will also switch hosts from marine to non-marine species. Krill are the bottom layer and most important part of the food chain in Antarctic animal communities. Some crustaceans are significant invasive species , such as the Chinese mitten crab, Eriocheir sinensis , and the Asian shore crab, Hemigrapsus sanguineus . Since

6308-411: The updated relationships between the different extant groups of the paraphyletic Crustacea in relation to the class Hexapoda . Ostracoda Mystacocarida Branchiura Pentastomida Malacostraca Copepoda Tantulocarida Thecostraca Cephalocarida Branchiopoda   Remipedia Hexapoda According to this diagram, the Hexapoda are deep in the Crustacea tree, and any of

6391-426: The zoea is the first larval stage. In some cases, the zoea stage is followed by the mysis stage, and in others, by the megalopa stage, depending on the crustacean group involved. Providing camouflage against predators, the otherwise black eyes in several forms of swimming larvae are covered by a thin layer of crystalline isoxanthopterin that gives their eyes the same color as the surrounding water, while tiny holes in

6474-446: Was analyzed after ultraviolet irradiation. This study revealed increased expression of proteins associated with the DNA repair processes of non-homologous end joining , homologous recombination , base excision repair and DNA mismatch repair . The name "crustacean" dates from the earliest works to describe the animals, including those of Pierre Belon and Guillaume Rondelet , but the name

6557-413: Was free to develop on its own, there is no reason to suppose that the speech was uniform either diachronically or geographically. On the contrary, Romanised European populations developed their own dialects of the language, which eventually led to the differentiation of Romance languages . Late Latin is a kind of written Latin used in the 3rd to 6th centuries. This began to diverge from Classical forms at

6640-496: Was much more liberal in its linguistic cohesion: for example, in classical Latin sum and eram are used as auxiliary verbs in the perfect and pluperfect passive, which are compound tenses. Medieval Latin might use fui and fueram instead. Furthermore, the meanings of many words were changed and new words were introduced, often under influence from the vernacular. Identifiable individual styles of classically incorrect Latin prevail. Renaissance Latin, 1300 to 1500, and

6723-441: Was no complete separation between Italian and Latin, even into the beginning of the Renaissance . Petrarch for example saw Latin as a literary version of the spoken language. Medieval Latin is the written Latin in use during that portion of the post-classical period when no corresponding Latin vernacular existed, that is from around 700 to 1500 AD. The spoken language had developed into the various Romance languages; however, in

6806-408: Was not used by some later authors, including Carl Linnaeus , who included crustaceans among the " Aptera " in his Systema Naturae . The earliest nomenclatural valid work to use the name "Crustacea" was Morten Thrane Brünnich 's Zoologiæ Fundamenta in 1772, although he also included chelicerates in the group. The subphylum Crustacea comprises almost 67,000 described species , which

6889-478: Was shut down in June 2019), and Vatican Radio & Television, all of which broadcast news segments and other material in Latin. A variety of organisations, as well as informal Latin 'circuli' ('circles'), have been founded in more recent times to support the use of spoken Latin. Moreover, a number of university classics departments have begun incorporating communicative pedagogies in their Latin courses. These include

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