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145-432: A mastodon ( mastós 'breast' + odoús 'tooth') is a member of the genus Mammut (German for 'mammoth'), which was endemic to North America and lived from the late Miocene to the early Holocene . Mastodons belong to the order Proboscidea , the same order as elephants and mammoths (which belong to the family Elephantidae ). Mammut is the type genus of the extinct family Mammutidae , which diverged from

290-526: A broader aridification trend. The EMCI ended 18 million years ago, giving way to the Middle Miocene Warm Interval (MMWI), the warmest part of which was the MMCO that began 16 million years ago. As the world transitioned into the MMCO, carbon dioxide concentrations varied between 300 and 500 ppm. Global annual mean surface temperature during the MMCO was about 18.4 °C. MMCO warmth was driven by

435-531: A change in the use of coastal resources and advancements in marine technology. The reasons for these changes have not been confirmed; various triggering mechanisms have been theorized such as climate change , the arrival of new people, or the struggle for resources. The South American land mammal age , the Lujanian , corresponds with the late Pleistocene. The Lujanian is a geologic period from 0.8 - 0.11Ma specifically for prehistoric South American fauna. There

580-519: A clade of large terrestrial predatory crocodyliformes distantly related to modern crocodilians, from which they likely diverged over 180 million years ago, are known from the Miocene of South America. The last Desmostylians thrived during this period before becoming the only extinct marine mammal order. The pinnipeds , which appeared near the end of the Oligocene, became more aquatic. A prominent genus

725-561: A cooler, drier climate. C 4 grasses, which are able to assimilate carbon dioxide and water more efficiently than C 3 grasses, expanded to become ecologically significant near the end of the Miocene between 6 and 7 million years ago, although they did not expand northward during the Late Miocene. The expansion of grasslands and radiations among terrestrial herbivores correlates to fluctuations in CO 2 . One study, however, has attributed

870-679: A family of the Elephantimorpha clade, it is only distantly related to the Deinotheriidae due to major differences in dentition and emergence of adult teeth. The Mammutidae is identified as a monophyletic clade, meaning that it did not leave any derived descendant groups in its evolutionary history. The monophyly of the Mammutidae makes it differ from the Elephantida, where the Gomphotheriidae

1015-473: A good analogue for future warmer climates caused by anthropogenic global warming , with this being especially true of the global climate during the Middle Miocene Climatic Optimum (MMCO), because the last time carbon dioxide levels were comparable to projected future atmospheric carbon dioxide levels resulting from anthropogenic climate change was during the MMCO. The Ross Sea margin of

1160-435: A good model for a "living fossil". Eucalyptus fossil leaves occur in the Miocene of New Zealand , where the genus is not native today, but have been introduced from Australia . Both marine and continental fauna were fairly modern, although marine mammals were less numerous. Only in isolated South America and Australia did widely divergent fauna exist. In Eurasia, genus richness shifted southward to lower latitudes from

1305-632: A greater understanding of extinctions. Taxonomically, it was first recognized as a distinct species by Robert Kerr in 1792 then classified to its own genus Mammut by Johann Friedrich Blumenbach in 1799, thus making it amongst the first fossil mammal genera to be erected with undisputed taxonomic authority. The genus served as a wastebasket taxon for proboscidean species with superficially similar molar teeth morphologies but today includes 7 definite species, 1 of questionable affinities, and 4 other species from Eurasia that are pending reassessments to other genera. Mastodons are considered to have had

1450-565: A letter dating to 1713, Edward Hyde, 3rd Earl of Clarendon (known also as Lord Cornbury) from New York reported to the Royal Society learned society of Great Britain that in 1705, a large-sized tooth was found near the side of the Hudson River by a Dutch country-fellow and was sold to New York General Assembly member Van Bruggen for a gill of rum, and Bruggen eventually gave it to Cornbury. He then stated that he sent Johannis Abeel,

1595-644: A long and complex paleontological history spanning all the way back to 1705 when the first fossils were uncovered from Claverack , New York in the American colonies. Because of the uniquely shaped molars with no modern analogues in terms of large animals, the species caught wide attention of European researchers and influential Americans before and after the American Revolution to the point of, according to American historians Paul Semonin and Keith Stewart Thomson , bolstering American nationalism and contributing to

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1740-579: A major expansion of Antarctic glaciers. This severed the connection between the Indian Ocean and the Mediterranean Sea and formed the present land connection between Afro-Arabia and Eurasia. The subsequent uplift of mountains in the western Mediterranean region and a global fall in sea levels combined to cause a temporary drying up of the Mediterranean Sea (known as the Messinian salinity crisis ) near

1885-480: A major expansion of grass-grazer ecosystems . Herds of large, swift grazers were hunted by predators across broad sweeps of open grasslands , displacing desert, woodland, and browsers . The higher organic content and water retention of the deeper and richer grassland soils , with long-term burial of carbon in sediments, produced a carbon and water vapor sink. This, combined with higher surface albedo and lower evapotranspiration of grassland, contributed to

2030-474: A majority known from endocasts , including M. americanum : Phosphatherium esculliei [REDACTED] Numidotherium koholense [REDACTED] Moeritherium lyonsi [REDACTED] Deinotheriidae [REDACTED] Palaeomastodon beadnelli [REDACTED] Miocene As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards

2175-643: A predominantly browsing -based diet on leaves, fruits, and woody parts of plants. This allowed mastodons to niche partition with other members of Proboscidea in North America, like gomphotheres and the Columbian mammoth , who had shifted to mixed feeding or grazing by the late Neogene - Quaternary . It is thought that mastodon behaviors were not much different from elephants and mammoths, with females and juveniles living in herds and adult males living largely solitary lives plus entering phases of aggression similar to

2320-626: A recorder of Albany , New York to dig near the original site of the tooth to find more bones. Abeel reported in a later that he went to the town of Claverack , New York where the original bones were found. American historian Paul Semonin said that the accounts written by Cornbury and Abeel match up with that written by in the July 30, 1705 entry in The Boston News-Letter . The account reported skeletal evidence of an antediluvian (or biblical) " giant " uncovered from Claverack. The femur and one of

2465-525: A second species for Pliomastodon named P. vexillarius based on fossil material from the locality of Elephant Hill in California , determining that it differs from Mammut by differences in the skull and that the etymology of the species name was made in honor of paleontological contributions by the Standard Oil Company of California. In 1933, Childs Frick named the species Mastodon raki from

2610-574: A series of ice ages . The Miocene boundaries are not marked by distinct global events but by regionally defined transitions from the warmer Oligocene to the cooler Pliocene Epoch. During the Early Miocene, Afro-Arabia collided with Eurasia, severing the connection between the Mediterranean and Indian Oceans, and allowing the interchange of fauna between Eurasia and Africa, including the dispersal of proboscideans and hominoids into Eurasia. During

2755-522: A significant drop in atmospheric carbon dioxide levels. Both continental and oceanic thermal gradients in mid-latitudes during the Early Miocene were very similar to those in the present. Global cooling caused the East Asian Summer Monsoon (EASM) to begin to take on its modern form during the Early Miocene. From 22.1 to 19.7 Ma, the Xining Basin experienced relative warmth and humidity amidst

2900-695: A significant local decline along the northeastern coast of Australia during the Tortonian, most likely due to warming seawater. Cetaceans attained their greatest diversity during the Miocene, with over 20 recognized genera of baleen whales in comparison to only six living genera. This diversification correlates with emergence of gigantic macro-predators such as megatoothed sharks and raptorial sperm whales . Prominent examples are O. megalodon and L. melvillei . Other notable large sharks were O. chubutensis , Isurus hastalis , and Hemipristis serra . Crocodilians also showed signs of diversification during

3045-604: A spring on the Pomme de Terre River to assemble a mounted skeleton of the " Missouri Leviathan " and briefly exhibited it at St. Louis . After exhibiting the skeleton throughout Europe, he sold the skeleton to the British Museum of Natural History . Richard Owen then properly reassembled the skeleton, and it today is on display there. In 1845, another skeleton was excavated from Newburgh by laborers hired by Nathaniel Brewster initially to remove lacustrine deposits to fertilize

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3190-561: A true domestic dog have been dated to 14,200 years ago. Domestication first happened in Eurasia but could have been anywhere from Western Europe to East Asia. Domestication of other animals such as cattle, goats, pigs, and sheep did not begin until the Holocene when settled farming communities became established in the Near East. The cat was probably not domesticated before c.  7500 BC at

3335-426: A view followed also by George William Featherstonhaugh . Isaac Hays comparatively defended Godman's taxon, which led to a bitter debate regarding the validity of the genus amongst American naturalists. The validities of both Tetracaulodon and Missourium were rejected by Owen in 1842, although he retained the former name informally. By 1869, American paleontologist Joseph Leidy determined that Mastodon americanus

3480-407: A view later followed by Franklin. In 1768, Scottish anatomist William Hunter recorded that he and his brother John Hunter observed that the teeth were not like those of modern elephants. He determined that the "grinders" from Ohio were of a carnivorous animal but believed that the tusks belonged to the same animal. After examining fossils from Franklin and Lord Shelburne, Hunter was convinced that

3625-627: A zone of low rainfall in the Late Miocene. The Indian Plate continued to collide with the Eurasian Plate , creating new mountain ranges and uplifting the Tibetan Plateau , resulting in the rain shadowing and aridification of the Asian interior. The Tian Shan experienced significant uplift in the Late Miocene, blocking westerlies from coming into the Tarim Basin and drying it as a result. At

3770-648: Is Z. proavus , which occurs in the Barstovian and Clarendonian stages. M? furlongi from the Black Butte in Oregon also dates back to the Clarendonian stage, but the affinities of the species remains unclear. If it truly is a species of Mammut , then its earliest temporal range is recorded at about 10 Ma. The earliest undisputed appearance of Mammut is of M. nevadanum from Thousand Creek Beds, dating back to

3915-469: Is paraphyletic (or ancestral to more derived descendant groups in the cladistic sense) in relation to the derived elephantoid families Stegodontidae and Elephantidae (elephants, mammoths, and relatives). Although the separation of the Mammutida and Elephantida is strongly supported based on morphological differences, their origins within the late Paleogene remain uncertain. One hypothesis asserts that

4060-553: Is conducive for preserving fossils . Neanderthal hominins ( Homo neanderthalensis ) inhabited Eurasia until becoming extinct between 40 and 30 ka, towards the end of the Pleistocene and possibly into the early Holocene and were replaced with modern humans ( Homo sapiens ) who emerged from East Africa about 195,000 years ago. Neanderthals co-existed with the Homo sapiens until they died out. In Eurasia, extinction happened throughout

4205-591: Is due to the lack of chronological information. The resemblance of Late Pleistocene species in Northern Africa to modern animals is the same as in Southern Africa but it's extremely difficult to date when these fauna came into place because of the lack of reliable samples from the mid-Pleistocene. Most of the significant fossil records are from the Maghreb because of its geology which helps to create deep caves which

4350-491: Is evidence of human habitation in mainland Australia , Indonesia , New Guinea and Tasmania from c. 45,000 BC. The finds include rock engravings, stone tools and evidence of cave habitation. In Australia, there are sites which show evidence of pollen records from the Late Pleistocene and they are mostly found in more temperate regions of the continent. Some megafauna decreased in size over time, while others remained

4495-599: Is extinct and has left no living descendants. He established that it had an overall body form similar to elephants but had molars more similar to hippopotamuses and pigs that did not serve to grind meat. The first species he erected within Mastodon was Mastodon giganteum , giving it the informal name "great mastodon" and writing that that it is designated to the Ohio proboscidean with abundant fossil evidence, equal size but greater proportions to modern elephants, and diamond-shaped points of

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4640-541: Is still heavily debated what caused the extinctions. Bison occidentalis and Bison antiquus , an extinct subspecies of the smaller present-day bison, survived the late Pleistocene period, between about 12 and 11 ka ago. Clovis people depended on these bison as their major food source. Earlier kills of camels, horses, and muskoxen found at Wally's beach were dated to 13.1–13.3 ka B.P. Over 50 genera (~ 83%) of megafauna in South and North America went extinct during

4785-472: Is the type genus of the Mammutidae , the sole family of the elephantimorph clade Mammutida (the other elephantimorph clade is Elephantida ). The Mammutidae is characterized by molars with zygodont-form crests, which have remained morphologically conservative throughout the evolutionary history of the family. Mammut is considered to be a derived genus of the family because of strong zygodont development. As

4930-401: Is the senior species synonym and listed M. giganteum as a junior synonym. He also listed Mammut , Harpagmotherium , Mastotherium , Missourium , and Leviathan as synonyms of Mastodon . He also noted that M. americanum as a species was highly variable in morphology. In 1902, American paleontologist Oliver Perry Hay listed Mammut as the prioritized genus name given its status as

5075-701: The American Museum of Natural History . Other skeletons of Mammut americanum were excavated within the United States in the first half of the 19th century. One of them was collected by American showman Albert C. Koch in what is today the Mastodon State Historic Site at Missouri in 1839. He hypothesized in 1840 that the proboscidean, which he classified as Missourium , was much larger than an elephant, had horizontal tusks plus trunks, and occupied aquatic habitats. He acquired additional fossils from

5220-641: The Bering land bridge which joined Alaska to Siberia . The last Ice Age was followed by the Late Glacial Interstadial , a period of global warming to 12.9 ka, and the Younger Dryas , a return to glacial conditions until 11.7 ka. Paleoclimatology holds that there was a sequence of stadials and interstadials from about 16 ka until the end of the Pleistocene. These were the Oldest Dryas (stadial),

5365-852: The Bølling oscillation (interstadial), the Older Dryas (stadial), the Allerød oscillation (interstadial) and finally the Younger Dryas. The end of the Younger Dryas marks the boundary between the Pleistocene and Holocene Epochs. Hominids in all parts of the world were still culturally and technologically in the Palaeolithic (Old Stone) Age . Tools and weapons were basic stone or wooden implements. Nomadic tribes followed moving herds. Non-nomadics acquired their food by gathering and hunting . Its present physical geography and climate have changed over time caused by

5510-558: The Clovis culture ) hunted mastodons based on the finding of mastodon remains with cut marks and/or with lithic artifacts. Mastodons disappeared along with many other North American animals, including most of its largest animals ( megafauna ), as part of the end-Pleistocene extinction event around the end of the Late Pleistocene-early Holocene, the causes typically being attributed to human hunting, severe climatic phases like

5655-660: The Coso Mountains in Inyo County , California where skull fossils were recovered. In 1963, J. Arnold Shotwell and Donald E. Russell created another species Mammut (Pliomastodon) furlongi , assigning it to fossils collected from the Juntura Formation of Oregon. The species name was created in honor of Eustace L. Furlong , who made early fossil collections from the western side of the Juntura Basin. The genus Pliomastodon

5800-772: The Holocene Epoch began. The term Upper Pleistocene is currently in use as a provisional or "quasi-formal" designation by the International Union of Geological Sciences (IUGS). Although the three oldest ages of the Pleistocene (the Gelasian , the Calabrian and the Chibanian ) have been officially defined, the late Pleistocene has yet to be formally defined. Following the brief Last Interglacial warm period (~130–115,000 years ago), where temperatures were comparable to or warmer than

5945-536: The Irvingtonian to the Rancholabrean (from around 1.6 million to 11,000 years ago), only M. americanum and the newly appearing M. pacificum are recorded, the former having an exceptional level of diversity based on abundant skeletal evidences from the late Pleistocene that is unusual for the typical mammutid fossil record. The following cladogram defines the phylogeny of certain proboscideans,

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6090-622: The Japanese archipelago has been traced to prehistoric times between 40,000 BC and 30,000 BC. The earliest fossils are radiocarbon dated to c. 35,000 BC. An archeological record of Neanderthals has been found in Asia along with records of two other hominin populations, the Denisovans and Homo floresiensis . Japan was once linked to the Asian mainland by land bridges via Hokkaido and Sakhalin Island to

6235-539: The Middle Miocene Climate Transition (MMCT). Abrupt increases in opal deposition indicate this cooling was driven by enhanced drawdown of carbon dioxide via silicate weathering. The MMCT caused a sea surface temperature (SST) drop of approximately 6 °C in the North Atlantic. The drop in benthic foraminiferal δ O values was most noticeable in the waters around Antarctica, suggesting cooling

6380-525: The Nile Valley as the Sahara was transformed from grassland to desert. The Nazlet Khater skeleton was found in 1980 and has been radiocarbon dated to between 30,360 and 35,100 years ago. Most of the knowledge of the Late Pleistocene is obtained from regions like Morocco , Algeria , Tunisia , some coastal regions of Maghreb , Libya and Egypt . The only issue with interpreting the data from this region

6525-709: The North American land mammal age scale, the Rancholabrean spans the time from c. 240,000 years ago to c. 11,000 years ago. It is named after the Rancho La Brea fossil site in California , characterized by extinct forms of bison in association with other Pleistocene species such as the mammoth . During the Late Pleistocene about 35 genera of megafauna went extinct including species such as mastodons , saber-toothed cats and giant ground sloths . Some other species went extinct in North America but not globally. it

6670-637: The Snake Creek Formation of western Nebraska , naming it in honor of William Diller Matthew . He also erected another species M. merriami from the Thousand Creek Formation in Nevada , which was eventually synonymized with Zygolophodon proavus . Osborn in 1926 followed up for Mastodon matthewi by establishing the genus Pliomastodon for the species based on cranial differences from " Miomastodon " (= Zygolophodon ). In 1930, Matthew erected

6815-420: The Younger Dryas , or some combination of the two. The American mastodon had its last recorded occurrence in the earliest Holocene around 11,000 years ago, which is considerably later than other North American megafauna species. Today, the American mastodon is one of the most well-known fossil species in both academic research and public perception, the result of its inclusion in American popular culture. In

6960-550: The carbon cycle occurred approximately 6 Ma, causing continental carbon reservoirs to no longer expand during cold spells, as they had done during cold periods in the Oligocene and most of the Miocene. At the end of the Miocene, global temperatures rose again as the amplitude of Earth's obliquity increased, which caused increased aridity in Central Asia. Around 5.5 Ma, the EAWM underwent a period of rapid intensification. Life during

7105-517: The mammoth , mastodon , and Irish elk became extinct. Upper Paleolithic people also made paintings and engravings on walls. Cave paintings have been found at Lascaux in the Dordogne which may be more than 17,000 years old. These are mainly buffalo , deer , and other animals hunted by humans. Later paintings occur in caves throughout the world, including Altamira , Spain, and in India, Australia, and

7250-520: The musth exhibited by modern elephants. Mammut achieved maximum species diversity in the Pliocene , though the genus is known from abundant fossil evidence in the Late Pleistocene . Mastodons for at least a few thousand years prior to their extinction coexisted with Paleoindians , who were the first humans to have inhabited North America. Evidence has been found that Paleoindians (including those of

7395-495: The " pseudo-elephant ", or " animal incognitum " (shortened as " incognitum "), was an animal species separate from elephants that might have also been the same as the proboscideans found in Siberia. He concluded his article with the opinion that although regrettable to philosophers, humanity should be thankful to heaven that the animal, if truly carnivorous, was extinct. In 1785, Reverend Robert Annan wrote an account recalling an event in which workers discovered bones in his farm near

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7540-484: The 19th century before eventually being reclassified into distinct genera. In addition to still-valid species names, several synonymous or dubious species names ultimately belonging to different genera were erected within the Americas as well throughout the 19th century. Also, many species names erected based on M. americanum remains were erected. As a result, M. americanum has many synonymous names. The issue of synonymous species names were especially apparent in

7685-431: The Alps and interglacials (temperate phase). The evidence of the changes in climatic conditions was from fragmentary sequences in formerly glaciated areas in northern Europe . The only domesticated animal in the Pleistocene was the dog , which evolved from the grey wolf into its many modern breeds . It is believed that the grey wolf became associated with hunter-gatherer tribes around 15 Ka. The earliest remains of

7830-429: The Americas. He stated that the bones that Buffon previously described from North America were not of elephants but another animal that he referred to as the " mastodonte ," or the " animal of Ohio ." He reinforced the idea that the extinct "mastodon" was an animal close in relationship to elephants that differed by jaws with large tubercles. He suggested that "mammoth" and "carnivorous elephant" be discontinued as names for

7975-452: The Aquitanian and Burdigalian Stages), the apes first evolved, began diversifying, and became widespread throughout the Old World . Around the end of this epoch, the ancestors of humans had split away from the ancestors of the chimpanzees and had begun following their own evolutionary path during the final Messinian Stage (7.5–5.3 Ma) of the Miocene. As in the Oligocene before it, grasslands continued to expand, and forests to dwindle. In

8120-617: The Big Bone Lick locality. He stated that the tusks were similar to elephants while the molars were completely different because they were covered with enamel and had a double row of high conical cusp processes. Kerr was unsure about the taxonomic affinities of the molars and referenced that Thomas Pennant supposed that they belong to an unknown species within the genus Elephas , giving the common name "American elephant." German naturalist Johann Friedrich Blumenbach also followed up with more taxonomic descriptions of fossil proboscideans in 1799. The first fossil species, recovered from Germany,

8265-479: The Blancan, and M. raki specifically is thought to not be synonymous with M. pacificum . M. americanum (known popularly as an "American mastodon" or simply "mastodon") is also stratigraphically recorded first from the early Blancan of the Ringold Formation , Washington . The age of the formation where the mammutid specimen was found dates to about 3.75 Ma. It is also known from multiple other Blancan sites such as Fish Springs Flat in Nevada. From

8410-417: The Central Paratethys, cut off from sources of freshwater input by its separation from the Eastern Paratethys. From 13.36 to 12.65 Ma, the Central Paratethys was characterised by open marine conditions, before the reopening of the Bârlad Strait resulted in a shift to brackish-marine conditions in the Central Paratethys, causing the Badenian-Sarmatian Extinction Event. As a result of the Bârlad Strait's reopening,

8555-721: The Early to the Middle Miocene. Europe's large mammal diversity significantly declined during the Late Miocene. In the Early Miocene, several Oligocene groups were still diverse, including nimravids , entelodonts , and three-toed equids. As in the previous Oligocene Epoch, oreodonts were still diverse, only to disappear in the earliest Pliocene. During the later Miocene mammals were more modern, with easily recognizable canids , bears , red pandas , procyonids , equids , beavers , deer , camelids , and whales , along with now-extinct groups like borophagine canids , certain gomphotheres , three-toed horses , and hornless rhinos like Teleoceras and Aphelos . The late Miocene also marks

8700-402: The East Antarctic Ice Sheet (EAIS) was highly dynamic during the Early Miocene. The Miocene began with the Early Miocene Cool Event (Mi-1) around 23 million years ago, which marked the start of the Early Miocene Cool Interval (EMCI). This cool event occurred immediately after the Oligocene-Miocene Transition (OMT) during a major expansion of Antarctica's ice sheets, but was not associated with

8845-399: The East Asian Winter Monsoon (EAWM) became stronger synchronously with a southward shift of the subarctic front. Greenland may have begun to have large glaciers as early as 8 to 7 Ma, although the climate for the most part remained warm enough to support forests there well into the Pliocene. Zhejiang, China was noticeably more humid than today. In the Great Rift Valley of Kenya , there

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8990-427: The Elephantimorpha is monophyletic if the primitive Elephantiformes genus Phiomia was truly ancestral to both the Elephantida and Mammutida. An alternate hypothesis suggests that the Elephantimorpha is diphyletic because Phiomia is ancestral to gomphotheres while Palaeomastodon is ancestral to mammutids. The earliest undisputed mammutid genus Losodokodon is recorded in Kenya , Africa and firmly establishes

9135-435: The Holocene, the Late Pleistocene was dominated by the cool Last Glacial Period , with temperatures gradually lowering throughout the period, reaching their lowest during the Last Glacial Maximum around 26-20,000 years ago. Most of the world's large ( megafaunal ) animals became extinct during the Late Pleistocene as part of the Late Pleistocene extinctions , a trend that continued into the Holocene. In palaeoanthropology ,

9280-418: The Hudson River in New York in fall of 1780. The workers found four molars in addition to another that was broken and thrown away. They also uncovered bones, including vertebrae that broke shortly thereafter. Annan expressed his confusion at what the animal could be but speculated based on its "grinders" that it was carnivorous in diet. He speculated also that it was probably extinct due to some catastrophe within

9425-411: The LMC; extratropical sea surface temperatures dropped substantially by approximately 7–9 °C. 41 kyr obliquity cycles became the dominant orbital climatic control 7.7 Ma and this dominance strengthened 6.4 Ma. Benthic δ O values show significant glaciation occurred from 6.26 to 5.50 Ma, during which glacial-interglacial cycles were governed by the 41 kyr obliquity cycle. A major reorganisation of

9570-405: The Late Cretaceous, are known from the Miocene of Patagonia, represented by the mole-like Necrolestes . The youngest known representatives of metatherians (the broader grouping to which marsupials belong) in Europe, Asia and Africa are known from the Miocene, including the European herpetotheriid Amphiperatherium , the peradectids Siamoperadectes and Sinoperadectes from Asia, and

9715-591: The Late Miocene, the Earth's climate began to display a high degree of similarity to that of the present day . The 173 kyr obliquity modulation cycle governed by Earth's interactions with Saturn became detectable in the Late Miocene. By 12 Ma, Oregon was a savanna akin to that of the western margins of the Sierra Nevada of northern California . Central Australia became progressively drier, although southwestern Australia experienced significant wettening from around 12 to 8 Ma. The South Asian Winter Monsoon (SAWM) underwent strengthening ~9.2–8.5 Ma. From 7.9 to 5.8 Ma,

9860-491: The Late Pleistocene. Some species which went extinct at the end of the Late Pleistocene in Southern Africa are the giant warthog , long-horn buffalo, Southern springbok , etc. These species were common because their distribution changed in response to climatic influences on vegetation. Carnivores were more widespread due to their varying habitat requirements. In Egypt , the Late (or Upper) Palaeolithic began sometime after 30,000 BC. People in North Africa had relocated to

10005-511: The Miocene Epoch was mostly supported by the two newly formed biomes , kelp forests and grasslands . Grasslands allow for more grazers, such as horses , rhinoceroses , and hippos . Ninety-five percent of modern plants existed by the end of this epoch . Modern bony fish genera were established. A modern-style latitudinal biodiversity gradient appeared ~15 Ma. The coevolution of gritty , fibrous, fire-tolerant grasses and long-legged gregarious ungulates with high-crowned teeth , led to

10150-449: The Miocene. The largest form among them was a gigantic caiman Purussaurus which inhabited South America. Another gigantic form was a false gharial Rhamphosuchus , which inhabited modern age India . A strange form, Mourasuchus also thrived alongside Purussaurus . This species developed a specialized filter-feeding mechanism, and it likely preyed upon small fauna despite its gigantic size. The youngest members of Sebecidae ,

10295-413: The North American Great Plains and in Argentina . The global trend was towards increasing aridity caused primarily by global cooling reducing the ability of the atmosphere to absorb moisture, particularly after 7 to 8 million years ago. Uplift of East Africa in the late Miocene was partly responsible for the shrinking of tropical rain forests in that region, and Australia got drier as it entered

10440-569: The North American species of Zygolophodon and Mammut . They synonymized P. adamsi and P. sellardsi with Mammut matthewi and emended M. nevadanus and M. pacificus to M. nevadanum and M. pacificum , respectively. They also said that they were uncertain of the taxonomic status of M. furlongi , specifically whether or not it was a variant of sexual dimorphism of Z. proavus . Some authors have considered M. nevadanum to be synonymous with M. matthewi while others had retained validity of

10585-705: The Oligocene–Miocene transgression. As the southern Andes rose in the Middle Miocene (14–12 million years ago) the resulting rain shadow originated the Patagonian Desert to the east. Far northern Australia was monsoonal during the Miocene. Although northern Australia is often believed to have been much wetter during the Miocene, this interpretation may be an artefact of preservation bias of riparian and lacustrine plants; this finding has itself been challenged by other papers. Western Australia, like today,

10730-658: The Pleistocene but those that happened during the Later Pleistocene were of megafauna and there were no replacements for the extinct species. Some Molluscan species went extinct but not on the same scale as the mammals living during the time. Some examples of species which extinct without replacements include the Straight-tusked elephant ( Palaeoloxodon antiquus ), Giant deer ( Megaloceros giganteus ), cave bear ( Ursus spelaeus ) and woolly rhinoceros ( Coelodonta antiquitatis ). Several large mammalian species including

10875-430: The Pleistocene. most mega mammals (>1000kg) and large mammals (>40kg) went extinct by the end of the Late Pleistocene. During this period there was a major cooling event called the Younger Dryas and the Clovis culture of capturing game became more prominent. Diverse factors such as climate change may have triggered this extinction but it's still in debate what the major factors were. The Late Pleistocene saw

11020-598: The Qiongdongnan Basin in the northern South China Sea indicates the Pearl River was a major source of sediment flux into the sea during the Early Miocene and was a major fluvial system as in the present. During the Oligocene and Early Miocene, the coast of northern Brazil, Colombia, south-central Peru , central Chile and large swathes of inland Patagonia were subject to a marine transgression . The transgressions in

11165-664: The Sahara. Magdalenian hunter-gatherers were widespread in western Europe about 20 -12.500 cal BP years ago until the end of the Pleistocene. An example of this is the antler-working done by the human groups who lived in the Santimamine cave in the Magdalenian. They invented the earliest known harpoons using reindeer horn. Climatic conditions during the Late Pleistocene in Eurasia were predominantly cold with glaciation events happening in northern Europe , northwest Siberia and

11310-487: The activity of the Columbia River Basalts and enhanced by decreased albedo from the reduction of deserts and expansion of forests. Climate modelling suggests additional, currently unknown, factors also worked to create the warm conditions of the MMCO. The MMCO saw the expansion of the tropical climatic zone to much larger than its current size. The July ITCZ, the zone of maximal monsoonal rainfall, moved to

11455-462: The ancestors of modern elephants at least 27–25 million years ago, during the Oligocene . Like other members of Mammutidae, the molar teeth of mastodons have zygodont morphology (where parallel pairs of cusps are merged into sharp ridges), which strongly differ from those of elephantids. In comparison to its likely ancestor Zygolophodon , Mammut is characterized by particularly long and upward curving upper tusks, reduced or absent tusks on

11600-472: The aridity of the former. Unequivocally-recognizable dabbling ducks , plovers , typical owls , cockatoos and crows appear during the Miocene. By the epoch's end, all or almost all modern bird groups are believed to have been present; the few post-Miocene bird fossils which cannot be placed in the evolutionary tree with full confidence are simply too badly preserved, rather than too equivocal in character. Marine birds reached their highest diversity ever in

11745-619: The beginning of the Miocene, the northern margin of the Arabian plate, then part of the African landmass, collided with Eurasia; as a result, the Tethys seaway continued to shrink and then disappeared as Africa collided with Eurasia in the Turkish – Arabian region. The first step of this closure occurred 20 Ma, reducing water mass exchange by 90%, while the second step occurred around 13.8 Ma, coincident with

11890-460: The bones were discovered by Native Americans (probably Abenaki hunter-warriors). He came to the conclusion that the femur and tusk belonged to an elephant while the molars (or cheek teeth) came from a separate giant hippopotamus . In Shawnee tradition, the proboscideans roamed in herds and were hunted by giants, who both eventually died out. The accounts told by the Shawnee individuals in 1762 are

12035-566: The course of this epoch . The youngest representatives of Choristodera , an extinct order of aquatic reptiles that first appeared in the Middle Jurassic , are known from the Miocene of Europe, belonging to the genus Lazarussuchus , which had been the only known surviving genus of the group since the beginning of the Eocene. The last known representatives of the archaic primitive mammal order Meridiolestida , which dominated South America during

12180-458: The earliest Miocene (~23-20 Ma) of Africa after Losodokodon . Eozygodon was subsequently succeeded by Zygolophodon by the early Miocene, and the latter dispersed into Eurasia by around 19-18 million years ago, and into North America by the middle Miocene. The dispersal of mammutids between Africa and Eurasia may have occurred multiple times. The Mammutidae eventually went extinct in Africa prior to

12325-415: The earliest presence of mammutids in the late Oligocene (~27-24 Ma). The Mammutidae, like other Paleogene proboscideans, was therefore an endemic radiation within the continent akin to other endemic mammals like arsinoitheres , hyracoids , and catarrhine primates plus non-endemics such as anthracotheres and hyaenodonts . In the early Neogene phase of evolution, Eozygodon made an appearance in

12470-662: The earliest, again in the Near East . A butchered brown bear patella found in Alice and Gwendoline Cave in County Clare and dated to 10,860 to 10,641 BC indicates the first known human activity in Ireland . The topography and geography of Asia were subject to frequent changes such as the creation of land bridges when sea levels dropped which helped with the expansion and migration of human populations . The first human habitation in

12615-463: The early Hemphillian , or 8.0-7.1 Ma. Historically, North American paleontologists considered that North American Zygolophodon evolved into Mammut in an endemic fashion while European workers generally thought that Mammut was a Eurasian immigrant that replaced North American Zygolophodon during the Miocene or Pliocene . Current evidence supports an endemic origin of North American Mammut from Zygolophodon without later migration because of

12760-530: The early to mid Miocene (23–15 Ma). Oceans cooled partly due to the formation of the Antarctic Circumpolar Current , and about 15 million years ago the ice cap in the southern hemisphere started to grow to its present form. The Greenland ice cap developed later, in the Middle Pliocene time, about 3 million years ago. Late Pleistocene The Late Pleistocene is an unofficial age in

12905-451: The end of the Miocene due to increased habitat uniformity. The expansion of grasslands in North America also led to an explosive radiation among snakes. Previously, snakes were a minor component of the North American fauna, but during the Miocene, the number of species and their prevalence increased dramatically with the first appearances of vipers and elapids in North America and the significant diversification of Colubridae (including

13050-539: The end of the Miocene. The Paratethys underwent a significant transgression during the early Middle Miocene. Around 13.8 Ma, during a global sea level drop, the Eastern Paratethys was cut off from the global ocean by the closure of the Bârlad Strait, effectively turning it into a saltwater lake. From 13.8 to 13.36 Ma, an evaporite period similar to the later Messinian salinity crisis in the Mediterranean ensued in

13195-612: The enigmatic Saint Bathans Mammal . Microbial life in the igneous crust of the Fennoscandian Shield shifted from being dominated by methanogens to being primarily composed of sulphate-reducing prokaryotes . The change resulted from fracture reactivation during the Pyrenean-Alpine orogeny, enabling sulphate-reducing microbes to permeate into the Fennoscandian Shield via descending surficial waters. Diatom diversity

13340-461: The evolution of both groups into modern representatives. The early Miocene Saint Bathans Fauna is the only Cenozoic terrestrial fossil record of the landmass, showcasing a wide variety of not only bird species, including early representatives of clades such as moa , kiwi and adzebills , but also a diverse herpetofauna of sphenodontians , crocodiles and turtles as well as a rich terrestrial mammal fauna composed of various species of bats and

13485-499: The expansion of grasslands not to a CO 2 drop but to the increasing seasonality and aridity, coupled with a monsoon climate, which made wildfires highly prevalent compared to before. The Late Miocene expansion of grasslands had cascading effects on the global carbon cycle, evidenced by the imprint it left in carbon isotope records. Cycads between 11.5 and 5 million years ago began to rediversify after previous declines in variety due to climatic changes, and thus modern cycads are not

13630-760: The extinction of the last-surviving members of the hyaenodonts . Islands began to form between South and North America in the Late Miocene, allowing ground sloths like Thinobadistes to island-hop to North America. The expansion of silica-rich C 4 grasses led to worldwide extinctions of herbivorous species without high-crowned teeth . Mustelids diversified into their largest forms as terrestrial predators like Ekorus , Eomellivora , and Megalictis and bunodont otters like Enhydriodon and Sivaonyx appeared. Eulipotyphlans were widespread in Europe, being less diverse in Southern Europe than farther north due to

13775-604: The findings of marine invertebrate fossils of both Atlantic and Pacific affinity in La Cascada Formation . Connection would have occurred through narrow epicontinental seaways that formed channels in a dissected topography . The Antarctic Plate started to subduct beneath South America 14 million years ago in the Miocene, forming the Chile Triple Junction . At first the Antarctic Plate subducted only in

13920-769: The first half of the 19th century. Today, the genera that include species formerly classified into Mastodon include Gomphotherium ( G. angustidens , G. pyrenaicum , G. productum , G. libycum , G. subtapiroideum , G. steinheimense ), Zygolophodon ( Z. turicensis , Z. proavus ), Cuvieronius ( C. hyodon ), Stegodon ( S. elephantoides ), Stegolophodon ( S. latidens , S. cautleyi ), Anancus ( A. avernensis , A. sivalensis , A. perimensis ), Tetralophodon ( T. longirostris ), Choerolophodon ( C. pentelici ), Stegomastodon ( S. mirificus ), Rhynchotherium (" R. " euhypodon ), Stenobelodon ( S. floridanus ), and Notiomastodon ( N. platensis ). In 1830, American naturalist John Davidson Godman created

14065-440: The genus Tetracaulodon plus its species T. Mastodontoideum based on what he determined to be differences between it and Mastodon based on the skull and dentition. Both Richard Harlan and William Cooper pointed out that except for the tusks, all other characteristics of the specimens were consistent with M. giganteum . They therefore argued that there was no reason to assume that the tusks were not just individual variations,

14210-714: The globe. American statesman Thomas Jefferson stated his thoughts on Notes on the State of Virginia (published by 1785) that the fossil proboscideans may have been carnivorous, still exist in the northern parts of North America, and are related to mammoths whose remains were found in Siberia. Jefferson referenced the theory of American social degeneracy by Georges-Louis Leclerc, Comte de Buffon , countering it by using extant and extinct animal measurements, including those of "mammoths," as proof that North America faunas were not "degenerative" in size. Semonin pointed out that social degeneracy

14355-600: The gradual appearance of Mammut morphologies and a lack of solid evidence that Mammut sensu stricto (in a strict sense) ever dispersed outside of North America. M. matthewi is recorded from the late Hemphillian to early Blancan stages. Mammutid specimens of the Hemphillian and Blancan had typically previously been assigned to M. matthewi , but this is seemingly the result of overreliance on stratigraphic positions to define taxa. M. vexillarius , M. raki , and M. cosoensis are definitively recorded from

14500-450: The historic plus taxonomically correct name Mammut over Mastodon . He continued prioritizing Mammut in 1945, stating that people were generally aware of its taxonomic priorities over Mastodon and that people had refused to use it. He stated that he did not want to either but reluctantly set aside his personal preferences to follow taxonomic rules. In 1921, Osborn created the species name Mastodon matthewi based on distinct molars from

14645-411: The human lineage) appeared in Africa at the very end of the Miocene, including Sahelanthropus , Orrorin , and an early form of Ardipithecus ( A. kadabba ). The chimpanzee–human divergence is thought to have occurred at this time. The evolution of bipedalism in apes at the end of the Miocene instigated an increased rate of faunal turnover in Africa. In contrast, European apes met their end at

14790-546: The international geologic timescale in chronostratigraphy , also known as the Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently defined as the time between c. 129,000 and c. 11,700 years ago. The late Pleistocene equates to the proposed Tarantian Age of the geologic time scale , preceded by

14935-587: The lake levels of the Eastern Paratethys dropped as it once again became a sea. The Fram Strait opened during the Miocene and acted as the only throughflow for Atlantic Water into the Arctic Ocean until the Quaternary period. Due to regional uplift of the continental shelf, this water could not move through the Barents Seaway in the Miocene. The modern day Mekong Delta took shape after 8 Ma. Geochemistry of

15080-669: The land bridge between South America and North America was absent, although South America was approaching the western subduction zone in the Pacific Ocean , causing both the rise of the Andes and a southward extension of the Meso-American peninsula. Mountain building took place in western North America , Europe , and East Asia . Both continental and marine Miocene deposits are common worldwide with marine outcrops common near modern shorelines. Well studied continental exposures occur in

15225-487: The last Eurasian mammutid, became extinct during the earliest Pleistocene , around 2.5-2 million years ago. The oldest evidence of mammutids in North America is of a fragmentary molar of Zygolophodon sp. from Massacre Lake , Nevada, dating to 16.5-16.4 Ma (during the Hemingfordian stage of the North American land mammal ages (NALMA)). The only definitively defined species of Zygolophodon from North America

15370-571: The late Miocene, the connections between the Atlantic and Mediterranean closed, causing the Mediterranean Sea to almost completely evaporate. This event is referred to as the " Messinian salinity crisis ". Then, at the Miocene–Pliocene boundary, the Strait of Gibraltar opened, and the Mediterranean refilled. That event is referred to as the " Zanclean flood ". Also during the early Miocene (specifically

15515-466: The late Miocene. Mammut as currently defined sensu lato (in a broad sense) is most likely polyphyletic (comprising several unrelated groups). This is because the inclusion of Eurasian mammutid species into Mammut implies that they share a common origin with North American Mammut , but this relationship has been doubted. As a result, these Eurasian species may belong to either other existing mammutid genera or entirely new genera. "Mammut" borsoni ,

15660-591: The late Pleistocene contains the Upper Palaeolithic stage of human development, including the early human migrations of modern humans outside of Africa, and the extinction of all archaic human species. Paleolithic Epipalaeolithic Mesolithic Neolithic The proposed beginning of the late Pleistocene is the end of the Penultimate Glacial Period (PGP) 126 ka when the Riß glaciation (Alpine)

15805-594: The locality in 1801, where he first sketched the fossils then purchased excavation privileges and full ownership of the fossils from Masten and borrowed a loan from the American Philosophical Society (APS) in Philadelphia , Pennsylvania . In addition to the first skeleton, the second was excavated using a mill-like device to drain a 12 ft (3.7 m) deep marl pit. Peale assembled a complete skeleton in his Philadelphia Museum in 1804, and its exhibit

15950-478: The locality of Truth or Consequences , New Mexico based on differences on the heel and M 3 tooth from M. americanus , otherwise having proportions similar to it. In 1936, Chester Stock published the species name Pliomastodon nevadanus based on fossils from the Thousand Creek Beds of northwestern Nevada. In 1937, John R. Schultz created the species name Pliomastodon? cosoensis , naming it after

16095-909: The lower jaw, as well as the shortening of the mandibular symphysis (the frontmost part of the lower jaw), the latter two traits also having evolved in parallel separately in elephantids. Mastodons had an overall stockier skeletal build, a lower-domed skull, and a longer tail compared to elephantids. Fully grown male M. americanum are thought to have been have been 275 cm (9.02 ft) to 305 cm (10.01 ft) at shoulder height and from 6.8 t (6.7 long tons; 7.5 short tons) to 9.2 t (9.1 long tons; 10.1 short tons) in body mass on average. The size estimates suggest that American mastodon males were on average heavier than any living elephant species; they were typically larger than Asian elephants and African forest elephants of both sexes but shorter than male African bush elephants . M. americanum , known as an "American mastodon" or simply "mastodon," had

16240-471: The molars. The naturalist also created the second species name Mastodon angustidens and gave it the informal name "narrow-toothed mastodon," diagnosing it as having narrower molars, smaller sizes compared to M. giganteum , and range distributions in Europe and South America. Cuvier also erected several other species of Mastodon originating from other continents in 1824. Despite Cuvier's genus name being younger than multiple other genus names, Mastodon became

16385-423: The most commonly used genus name for the 19th century. " Mastodon " was riddled with major taxonomic problems since species now determined as belonging to other proboscidean genera were classified to Mastodon on the basis of similar dentitions to that of " Mastodon giganteum " (= Mammut americanum ), effectively making it a wastebasket taxon . Various fossil proboscidean species were classified into Mastodon in

16530-455: The movement of tectonic plates and volcanoes but glacial cycles and sea level variation have a more significant effect on the vertebrate communities during the Late Pleistocene. The Late Pleistocene was the time when most animals evolved to resemble modern-day animals and they managed to live through the Late mid-Pleistocene since there were no extinction events of megafauna until the end of

16675-503: The neighboring fields. They were observed by a large amount of spectators and uncovered relatively complete fossil evidence of M. americanum . The skeleton was exhibited in New York City and other New England towns then was acquired by John Collins Warren for study. After Warren's death in 1856, the skeleton was sent to Warren's family but was traded to Harvard Medical School for John Warren's skeleton. The "Warren mastodon", under

16820-561: The north but was unconnected at this time when the main islands of Hokkaido, Honshu , Kyushu and Shikoku were all separate entities. Human migrations happened during this time with people coming in from Eurasia . From about 28 ka, there were migrations across the Bering land bridge from Siberia to Alaska . The people became the Native Americans . It is believed that the original tribes subsequently moved down to Central and South America under pressure from later migrations. In

16965-543: The north, increasing precipitation over southern China whilst simultaneously decreasing it over Indochina during the EASM. Western Australia was at this time characterised by exceptional aridity. In Antarctica, average summer temperatures on land reached 10 °C. In the oceans, the lysocline shoaled by approximately half of a kilometre during warm phases that corresponded to orbital eccentricity maxima. The MMCO ended around 14 million years ago, when global temperatures fell in

17110-488: The northern hemisphere. The Miocene faunal stages from youngest to oldest are typically named according to the International Commission on Stratigraphy : Regionally, other systems are used, based on characteristic land mammals; some of them overlap with the preceding Oligocene and following Pliocene Epochs: Continents continued to drift toward their present positions. Of the modern geologic features, only

17255-543: The officially ratified Chibanian (commonly known as the Middle Pleistocene). The beginning of the Late Pleistocene is the transition between the end of the Penultimate Glacial Period and the beginning of the Last Interglacial around 130,000 years ago (corresponding with the beginning of Marine Isotope Stage 5 ). The Late Pleistocene ends with the termination of the Younger Dryas , some 11,700 years ago when

17400-404: The oldest genus name, making Mastodon , Tetracaulodon , and Missourium classified as junior synonyms. He also established M. americanum as the type species. The genus name Mastodon was subsequently abandoned by many American paleontologists in favor of Mammut within the early 20th century. In 1942, American paleontologist George Gaylord Simpson said that for his study, he prioritized

17545-519: The oldest known documented interpretations of the " Ohio " fossils, although the traditions may have had been told for generations. In 1767, Peter Collinson credited Irish trader George Croghan for having sent him and Benjamin Franklin fossil evidence of the mysterious proboscideans, using them for his studies. He concluded that the peculiar grinders (the molars) were built for herbivorous diets of branches of trees and shrubs as well as other vegetation,

17690-523: The origin of many modern genera such as Nerodia , Lampropeltis , Pituophis and Pantherophis ). Arthropods were abundant, including in areas such as Tibet where they have traditionally been thought to be undiverse. Neoisopterans diversified and expanded into areas they previously were absent from, such as Madagascar and Australia. In the oceans, brown algae , called kelp , proliferated, supporting new species of sea life, including otters , fish and various invertebrates . Corals suffered

17835-463: The possible herpetotheriid Morotodon from the late Early Miocene of Uganda. Approximately 100 species of apes lived during this time , ranging throughout Africa, Asia and Europe and varying widely in size, diet, and anatomy. Due to scanty fossil evidence it is unclear which ape or apes contributed to the modern hominid clade, but molecular evidence indicates this ape lived between 18 and 13 million years ago. The first hominins ( bipedal apes of

17980-600: The previously established species " Elephas americanus " and argued that the species was different from elephants and mammoths and cannot be found amongst living animals due to extinction from catastrophism . The proboscidean species was subject to several other species names given by other taxonomists within the earliest 18th century as well as the genus name Harpagmotherium by the Russian naturalist Gotthelf Fischer von Waldheim in 1808. In 1806, Cuvier wrote multiple extended research articles on fossil proboscideans of Eurasia and

18125-488: The promotion of the "mastodon" skeleton made it a symbol of the strength of American nationalism and that "mammoth" as a term became associated with gigantism. Decades later, the museum bankrupted, and the first skeleton's specimens were sold to some German spectators in around 1848, who eventually sold it to Hessisches Landesmuseum Darmstadt in Germany where it is now displayed. The second skeleton's specimens landed eventually at

18270-499: The request of American paleontologist Henry Fairfield Osborn , was purchased by the American financier J. P. Morgan for $ 30,000 in 1906 and donated to the American Museum of Natural History where it is exhibited today. In the 1790s, the " American incognitum " was subject to research by multiple taxonomists. Scottish writer Robert Kerr erected the species name Elephas americanus in 1792 based on fossil tusks and "grinders" from

18415-577: The seas of the Miocene, kelp forests made their first appearance and soon became one of Earth's most productive ecosystems. The plants and animals of the Miocene were recognizably modern. Mammals and birds were well established. Whales , pinnipeds , and kelp spread. The Miocene is of particular interest to geologists and palaeoclimatologists because major phases of the geology of the Himalaya occurred during that epoch, affecting monsoonal patterns in Asia, which were interlinked with glacial periods in

18560-695: The southernmost tip of Patagonia, meaning that the Chile Triple Junction lay near the Strait of Magellan . As the southern part of Nazca Plate and the Chile Rise became consumed by subduction the more northerly regions of the Antarctic Plate begun to subduct beneath Patagonia so that the Chile Triple Junction advanced to the north over time. The asthenospheric window associated to the triple junction disturbed previous patterns of mantle convection beneath Patagonia inducing an uplift of ca. 1 km that reversed

18705-473: The species and that it receive a new genus name instead. Cuvier said that for " mastodonte ," he derived the name's etymology (compound μαστός ( mastós , "breast") + ὀδούς ( odoús , "tooth") from Ancient Greek to mean "nipple tooth," since he thought that it expressed the characteristic form of the teeth. In 1817, the French naturalist officially established the genus name Mastodon , reaffirming that it

18850-406: The species name. Several mammutid species outside of North America are classified to Mammut (or " Pliomastodon "), namely M. borsoni , M. obliquelophus , M. zhupengensis , and M. lufugense (possibly synonymous with M. obliquelophus ). Recent research such as that of von Koenigswald et al. in 2023 warned that the genus Mammut should be carefully used for non-North American species. Mammut

18995-537: The species was distinguished from other animals of the prehistoric world based on the unusual shapes of the large molars. The genus name "Mammut" refers to the German translation for "mammoth." The naming of the genus Mammut in 1799 makes it the second or third genus to be recognized with taxonomic authority given that Megalonyx had been named the same year. French naturalist Georges Cuvier also described known fossil proboscidean species back in 1796, although his account

19140-575: The teeth both dissolved before they could be further observed, however. In 1739, a French military expedition under the command of Charles III Le Moyne (known also as "Longueil") explored the locality of " Big Bone Lick " (located in what is now the US state of Kentucky ) and gathered fossil bones and teeth there. The French naturalist Louis Jean-Marie Daubenton examined the fossil collection brought by Longueuil and compared it with specimens of extant elephants and Siberian mammoths in 1762. Daubenton said that

19285-590: The west coast of South America are thought to be caused by a regional phenomenon while the steadily rising central segment of the Andes represents an exception. While there are numerous registers of Oligocene–Miocene transgressions around the world it is doubtful that these correlate. It is thought that the Oligocene–Miocene transgression in Patagonia could have temporarily linked the Pacific and Atlantic Oceans, as inferred from

19430-423: Was Allodesmus . A ferocious walrus , Pelagiarctos may have preyed upon other species of pinnipeds including Allodesmus . Furthermore, South American waters witnessed the arrival of Megapiranha paranensis , which were considerably larger than modern age piranhas . New Zealand 's Miocene fossil record is particularly rich. Marine deposits showcase a variety of cetaceans and penguins , illustrating

19575-610: Was a gradual and progressive trend of increasing aridification, though it was not unidirectional, and wet humid episodes continued to occur. Between 7 and 5.3 Ma, temperatures dropped sharply again in the Late Miocene Cooling (LMC), most likely as a result of a decline in atmospheric carbon dioxide and a drop in the amplitude of Earth's obliquity, and the Antarctic ice sheet was approaching its present-day size and thickness. Ocean temperatures plummeted to near-modern values during

19720-526: Was an offensive concept to Anglo-American naturalists and that the American proboscidean fossils were used as political tools to inspire American nationalism and counter against the theory of American degeneracy. In 1799, laborers recovered a thighbone while digging a marl pit at John Masten's farm in Newburgh , New York, and subsequent excavations were observed by a crowd of over a hundred people. American painter and exhibitionist Charles Willson Peale visited

19865-458: Was arid, particularly so during the Middle Miocene. Climates remained moderately warm, although the slow global cooling that eventually led to the Pleistocene glaciations continued. Although a long-term cooling trend was well underway, there is evidence of a warm period during the Miocene when the global climate rivalled that of the Oligocene . The climate of the Miocene has been suggested as

20010-624: Was being succeeded by the Eemian (Riß-Würm) interglacial period . The Riß-Würm ended 115 ka with the onset of the Last Glacial Period (LGP) which is known in Europe as the Würm (Alpine) or Devensian (Great Britain) or Weichselian glaciation (northern Europe); these are broadly equated with the Wisconsin glaciation (North America), though technically that began much later. The Last Glacial Maximum

20155-400: Was described as belonging to the newly erected species Elephas primigenius ? (now known as Mammuthus primigenius ). The second was what he considered to be an unknown "colossal land monster of the prehistoric world," considering it to be the "mammoth." He created the genus Mammut and erected the species Mammut ohioticum based on fossil bones dug up from Ohio in North America. He said that

20300-671: Was inversely correlated with carbon dioxide levels and global temperatures during the Miocene. Most modern lineages of diatoms appeared by the Late Miocene. There is evidence from oxygen isotopes at Deep Sea Drilling Program sites that ice began to build up in Antarctica about 36 Ma during the Eocene . Further marked decreases in temperature during the Middle Miocene at 15 Ma probably reflect increased ice growth in Antarctica. It can therefore be assumed that East Antarctica had some glaciers during

20445-456: Was later published in 1799. He considered that the remains uncovered from Siberia were true "mammoths" that had similar dentitions to extant elephants but had some morphological differences. He mentioned the fossil remains that were brought back by Longueil from Ohio back in 1739 and several researchers from previous decades who noted the unusual molars and thought that they belonged to different animals like hippopotamuses. He followed recognition in

20590-627: Was most intense there. Around this time the Mi3b glacial event (a massive expansion of Antarctic glaciers) occurred. The East Antarctic Ice Sheet (EAIS) markedly stabilised following the MMCT. The intensification of glaciation caused a decoherence of sediment deposition from the 405 kyr eccentricity cycle. The MMWI ended about 11 Ma, when the Late Miocene Cool Interval (LMCI) started. A major but transient warming occurred around 10.8-10.7 Ma. During

20735-665: Was open first to invited members of the American Philosophical Society on December 24 then to the general public on December 25 for an exhibit admission fee in addition to the general admission fee. The special exhibition attracted thousands of visitors, and the skeleton became a US national symbol. Charles Peale's son Rembrandt Peale took the skeleton to Europe used to promote the fossil proboscidean and have it used as support for Jefferson's final rebuttals against Buffon's arguments for supposed inferiority of American faunas. Author Keith Stewart Thomson argued that

20880-686: Was reached during the later millennia of the Würm/Weichselian, estimated between 26 ka and 19 ka when deglaciation began in the Northern Hemisphere. The Würm/Weichselian endured until 16 ka with Northern Europe, including most of Great Britain , covered by an ice sheet. The glaciers reached the Great Lakes in North America. Sea levels fell and two land bridges were temporarily in existence that had significance for human migration : Doggerland , which connected Great Britain to mainland Europe; and

21025-693: Was synonymized with Mammut while Miomastodon was synonymized with Zygolophodon by Jeheskel Shoshani and Pascal Tassy in a 1996 appendix, a view that was followed by other authors in later years. In 2019, Alton C. Dooley Jr. et al. established Mammut pacificus based on fossils collected from the Diamond Valley Lake in Hemet , California. They also stated that M. oregonense is a nomen dubium and that further analysis needs to be done to confirm whether or not M. furlongi belongs to Zygolophodon instead. In 2023, Wighart von Koenigswald et al. reviewed

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