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100-475: Anartia jatrophae , the white peacock , is a species of butterfly found in the southeastern United States , Central America , and throughout much of South America . The white peacock's larval hosts are water hyssop ( Bacopa monnieri ), lemon bacopa ( Bacopa caroliniensis ), tropical waterhyssop ( Bacopa innominata ), frogfruit ( Phyla nodiflora ), lanceleaf frogfruit ( Phyla lanceolata ), and Carolina wild petunia ( Ruellia caroliniana ). The males of

200-631: A diapause (resting) stage, and the hatching may take place only in spring. Some temperate region butterflies, such as the Camberwell beauty , lay their eggs in the spring and have them hatch in the summer. Butterfly larvae, or caterpillars, consume plant leaves and spend practically all of their time searching for and eating food. Although most caterpillars are herbivorous, a few species are predators : Spalgis epius eats scale insects , while lycaenids such as Liphyra brassolis are myrmecophilous , eating ant larvae. Some larvae, especially those of

300-441: A schwa ( /ə/ ). A short phrase from the gospel book of Munsterbilzen Abbey , written around 1130, still shows several unstressed vowels distinguished: That was a late monument, however, as the merging of all unstressed short vowels was already well underway by that time. Most likely, the difference was maintained only in spelling traditions, but it had been mostly lost in speech. With the introduction of new scribal traditions in

400-476: A North sea Germanic substrate . Linguists typically date this transition to around the 5th century. Old Dutch is divided into Old West Low Franconian and Old East Low Franconian ( Limburgian ); however, these varieties are very closely related, the divergence being that the latter shares more traits with neighboring historical forms of Central Franconian dialects such as Ripuarian and Moselle Franconian . While both forms of Low Franconian were instrumental to

500-582: A West Flemish monk in a convent in Rochester , England . For a long time, the sentence was commonly but erroneously considered to be the earliest in Dutch. However, it could be considered the oldest Dutch non-religious poetry. The text is usually considered a West Flemish dialect, but certain Ingvaeonic forms might be expected in any of the coastal dialects of Old English, Old Frisian, Old Saxon or Old Dutch. However,

600-493: A butterfly cannot fly until the wings are unfolded. A newly emerged butterfly needs to spend some time inflating its wings with hemolymph and letting them dry, during which time it is extremely vulnerable to predators. The colourful patterns on many butterfly wings tell potential predators that they are toxic. Hence, the genetic basis of wing pattern formation can illuminate both the evolution of butterflies as well as their developmental biology . The colour of butterfly wings

700-570: A butterfly through metamorphosis has held great appeal to mankind. To transform from the miniature wings visible on the outside of the pupa into large structures usable for flight, the pupal wings undergo rapid mitosis and absorb a great deal of nutrients. If one wing is surgically removed early on, the other three will grow to a larger size. In the pupa, the wing forms a structure that becomes compressed from top to bottom and pleated from proximal to distal ends as it grows, so that it can rapidly be unfolded to its full adult size. Several boundaries seen in

800-417: A common family. In some species, such as the great spangled fritillary , the eggs are deposited close to but not on the food plant. This most likely happens when the egg overwinters before hatching and where the host plant loses its leaves in winter, as do violets in this example. The egg stage lasts a few weeks in most butterflies, but eggs laid close to winter, especially in temperate regions, go through

900-411: A dialect continuum formed/existed between Old Dutch, Old Saxon and Old Frisian. Despite sharing some particular features, a number of disparities separate Old Saxon, Old Frisian, Old English and Old Dutch. One such difference is that Old Dutch used -a as its plural a-stem noun ending, while Old Saxon and Old English employed -as or -os . Much of the grammatical variation between Old Dutch and Old Saxon

1000-436: A distinction in writing: dag "day" (short vowel), thahton "they thought" (long vowel). Later on, the long vowels were sometimes marked with a macron to indicate a long vowel: ā . In some texts long vowels were indicated by simply doubling the vowel in question, as in the placename Heembeke and personal name Oodhelmus (both from charters written in 941 and 797 respectively). Old Dutch may have preserved at least four of

1100-571: A few butterflies (e.g., harvesters ) eat harmful insects, and a few are predators of ants , while others live as mutualists in association with ants. Culturally, butterflies are a popular motif in the visual and literary arts. The Smithsonian Institution says "butterflies are certainly one of the most appealing creatures in nature". The Oxford English Dictionary derives the word straightforwardly from Old English butorflēoge , butter-fly; similar names in Old Dutch and Old High German show that

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1200-444: A few species. Some butterflies have organs of hearing and some species make stridulatory and clicking sounds. Many species of butterfly maintain territories and actively chase other species or individuals that may stray into them. Some species will bask or perch on chosen perches. The flight styles of butterflies are often characteristic and some species have courtship flight displays. Butterflies can only fly when their temperature

1300-606: A hard-ridged outer layer of shell, called the chorion . This is lined with a thin coating of wax which prevents the egg from drying out before the larva has had time to fully develop. Each egg contains a number of tiny funnel-shaped openings at one end, called micropyles ; the purpose of these holes is to allow sperm to enter and fertilize the egg. Butterfly eggs vary greatly in size and shape between species, but are usually upright and finely sculptured. Some species lay eggs singly, others in batches. Many females produce between one hundred and two hundred eggs. Butterfly eggs are fixed to

1400-503: A leaf before eggs are laid on it. Many butterflies use chemical signals, pheromones ; some have specialized scent scales ( androconia ) or other structures ( coremata or "hair pencils" in the Danaidae). Vision is well developed in butterflies and most species are sensitive to the ultraviolet spectrum. Many species show sexual dimorphism in the patterns of UV reflective patches. Colour vision may be widespread but has been demonstrated in only

1500-416: A leaf with a special glue which hardens rapidly. As it hardens it contracts, deforming the shape of the egg. This glue is easily seen surrounding the base of every egg forming a meniscus. The nature of the glue has been little researched but in the case of Pieris brassicae , it begins as a pale yellow granular secretion containing acidophilic proteins. This is viscous and darkens when exposed to air, becoming

1600-547: A longer lifespan of several months as adults. The thorax of the butterfly is devoted to locomotion. Each of the three thoracic segments has two legs (among nymphalids , the first pair is reduced and the insects walk on four legs). The second and third segments of the thorax bear the wings. The leading edges of the forewings have thick veins to strengthen them, and the hindwings are smaller and more rounded and have fewer stiffening veins. The forewings and hindwings are not hooked together ( as they are in moths ) but are coordinated by

1700-543: A monk of the Abbey of Egmond , and so the manuscript's other name is Egmond Willeram . The text represents an imperfect attempt to translate the original into the local Old Dutch vernacular. The text contains many Old Dutch words as well as mistranslated words since the scribe must have been unfamiliar with some Old High German words in the original. It could nevertheless be regarded as the first book written in Old Dutch. However, since

1800-448: A necessity for their successful establishment. Many butterflies, such as the painted lady , monarch, and several danaine migrate for long distances. These migrations take place over a number of generations and no single individual completes the whole trip. The eastern North American population of monarchs can travel thousands of miles south-west to overwintering sites in Mexico . There is

1900-646: A reverse migration in the spring. It has recently been shown that the British painted lady undertakes a 9,000-mile round trip in a series of steps by up to six successive generations, from tropical Africa to the Arctic Circle — almost double the length of the famous migrations undertaken by monarch. Spectacular large-scale migrations associated with the monsoon are seen in peninsular India. Migrations have been studied in more recent times using wing tags and also using stable hydrogen isotopes . Butterflies navigate using

2000-412: A segmented palp. Adjoining these is the labium-hypopharynx which houses a tubular spinneret which is able to extrude silk. Caterpillars such as those in the genus Calpodes (family Hesperiidae) have a specialized tracheal system on the 8th segment that function as a primitive lung. Butterfly caterpillars have three pairs of true legs on the thoracic segments and up to six pairs of prolegs arising from

2100-435: A silken girdle may be spun to keep the pupa in a head-up position. Most of the tissues and cells of the larva are broken down inside the pupa, as the constituent material is rebuilt into the imago. The structure of the transforming insect is visible from the exterior, with the wings folded flat on the ventral surface and the two halves of the proboscis, with the antennae and the legs between them. The pupal transformation into

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2200-410: A time-compensated sun compass. They can see polarized light and therefore orient even in cloudy conditions. The polarized light near the ultraviolet spectrum appears to be particularly important. Many migratory butterflies live in semi-arid areas where breeding seasons are short. The life histories of their host plants also influence butterfly behaviour. Butterflies in their adult stage can live from

2300-528: A total of about 20,000 species. Traditionally, butterflies have been divided into the superfamilies Papilionoidea and the moth-like Hedyloidea . Recent work has discovered that Hedylidae, the only family within Hedyloidea, is nested within the Papilionoidea, meaning that Papilionoidea would be synonymous with Rhopalocera. The relationships between the rest of the 6 families are extremely well resolved, which

2400-470: A trend towards multivoltinism . Courtship is often aerial and often involves pheromones . Butterflies then land on the ground or on a perch to mate. Copulation takes place tail-to-tail and may last from minutes to hours. Simple photoreceptor cells located at the genitals are important for this and other adult behaviours. The male passes a spermatophore to the female; to reduce sperm competition, he may cover her with his scent, or in some species such as

2500-486: A tubular proboscis which is curled up at rest and expanded when needed to feed. The first and second maxillae bear palps which function as sensory organs. Some species have a reduced proboscis or maxillary palps and do not feed as adults. Many Heliconius butterflies also use their proboscis to feed on pollen; in these species only 20% of the amino acids used in reproduction come from larval feeding, which allow them to develop more quickly as caterpillars, and gives them

2600-437: A water-insoluble, rubbery material which soon sets solid. Butterflies in the genus Agathymus do not fix their eggs to a leaf; instead, the newly laid eggs fall to the base of the plant. Eggs are almost invariably laid on plants. Each species of butterfly has its own host plant range and while some species of butterfly are restricted to just one species of plant, others use a range of plant species, often including members of

2700-422: A week to nearly a year depending on the species. Many species have long larval life stages while others can remain dormant in their pupal or egg stages and thereby survive winters. The Melissa Arctic ( Oeneis melissa ) overwinters twice as a caterpillar. Butterflies may have one or more broods per year. The number of generations per year varies from temperate to tropical regions with tropical regions showing

2800-450: Is above 27 °C (81 °F); when it is cool, they can position themselves to expose the underside of the wings to the sunlight to heat themselves up. If their body temperature reaches 40 °C (104 °F), they can orientate themselves with the folded wings edgewise to the sun. Basking is an activity which is more common in the cooler hours of the morning. Some species have evolved dark wingbases to help in gathering more heat and this

2900-458: Is derived from tiny structures called scales, each of which have their own pigments . In Heliconius butterflies, there are three types of scales: yellow/white, black, and red/orange/brown scales. Some mechanism of wing pattern formation are now being solved using genetic techniques. For instance, a gene called cortex determines the colour of scales: deleting cortex turned black and red scales yellow. Mutations, e.g. transposon insertions of

3000-403: Is especially evident in alpine forms. As in many other insects, the lift generated by butterflies is more than can be accounted for by steady-state, non-transitory aerodynamics . Studies using Vanessa atalanta in a wind tunnel show that they use a wide variety of aerodynamic mechanisms to generate force. These include wake capture , vortices at the wing edge, rotational mechanisms and

3100-410: Is extruded and inserted into the female's vagina. A spermatophore is deposited in the female, following which the sperm make their way to a seminal receptacle where they are stored for later use. In both sexes, the genitalia are adorned with various spines, teeth, scales and bristles, which act to prevent the butterfly from mating with an insect of another species. After it emerges from its pupal stage,

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3200-451: Is good, especially in some species in the blue/violet range. The antennae are composed of many segments and have clubbed tips (unlike moths that have tapering or feathery antennae). The sensory receptors are concentrated in the tips and can detect odours. Taste receptors are located on the palps and on the feet. The mouthparts are adapted to sucking and the mandibles are usually reduced in size or absent. The first maxillae are elongated into

3300-563: Is native to the Americas, but in the nineteenth century or before, spread across the world, and is now found in Australia, New Zealand, other parts of Oceania, and the Iberian Peninsula . It is not clear how it dispersed; adults may have been blown by the wind or larvae or pupae may have been accidentally transported by humans, but the presence of suitable host plants in their new environment was

3400-590: Is now archived in the Vatican Codex pal. 577. Sometimes interpreteted as Old Saxon, a number of Dutch scholars have concluded the Baptismal Vow was actually written in the 8th century in Old Dutch. The difficulty in establishing whether the text was written in Old Saxon or Old Franconian is that those languages were very much alike. Irlôsin sol an frithe sêla mîna fan thên thia ginâcont mi, wanda under managon he

3500-468: Is now the southern Netherlands , northern Belgium , part of northern France, and parts of the Lower Rhine regions of Germany. It evolved into Middle Dutch around the 12th century. The inhabitants of northern Dutch provinces, including Groningen , Friesland , and the coast of North Holland , spoke Old Frisian , and some in the east ( Achterhoek , Overijssel , and Drenthe ) spoke Old Saxon . Within

3600-491: Is often cited as the time of the discontinuity, but it actually marks a time of profuse Dutch writing whose language is patently different from Old Dutch. The most notable difference between Old and Middle Dutch is vowel reduction . Back vowels ( a , o ) in non-stressed syllables are rather frequent in Old Dutch, but in Middle Dutch, they are reduced to a schwa : The following is a translation of Psalm 55 :18, taken from

3700-465: Is restricted to the males, and studies have suggested that the nutrients collected may be provided as a nuptial gift , along with the spermatophore, during mating. In hilltopping , males of some species seek hilltops and ridge tops, which they patrol in search for females. Since it usually occurs in species with low population density, it is assumed these landscape points are used as meeting places to find mates. Butterflies use their antennae to sense

3800-505: Is similar to that between Old Dutch and Old High German. It is also found that Old Dutch had lost the dual number for its pronouns, unlike Old English, which used wit to refer to "the two of us". Old Dutch would have used we both to refer to that and to refer to many more people in the "us" group, much like Modern Dutch and English. Old Dutch naturally evolved into Middle Dutch with some distinctions that approximate those found in most medieval West Germanic languages. The year 1150

3900-692: Is summarized in the below cladogram. Papilionidae [REDACTED] Hedylidae [REDACTED] Hesperiidae [REDACTED] Pieridae [REDACTED] Nymphalidae [REDACTED] Lycaenidae [REDACTED] Riodinidae [REDACTED] Butterfly adults are characterized by their four scale-covered wings, which give the Lepidoptera their name ( Ancient Greek λεπίς lepís, scale + πτερόν pterón, wing). These scales give butterfly wings their colour: they are pigmented with melanins that give them blacks and browns, as well as uric acid derivatives and flavones that give them yellows, but many of

4000-516: Is synonymous with Old Dutch . Depending on the author, the temporal boundary between Old Dutch and Old Frankish is either defined by the onset of the Second Germanic consonant shift in Eastern Frankish, the assimilation of an unattested coastal dialect showing North Sea Germanic -features by West Frankish during the closing of the 9th century , or a combination of both. Some linguists use

4100-405: Is taken up by the gut, but there may also be large silk glands, and special glands which secrete distasteful or toxic substances. The developing wings are present in later stage instars and the gonads start development in the egg stage. When the larva is fully grown, hormones such as prothoracicotropic hormone (PTTH) are produced. At this point the larva stops feeding, and begins "wandering" in

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4200-508: Is the subsumption of the High German consonant shift , a set of phonological changes beginning around the 5th or 6th century that partially influenced Old Dutch, and extensively influenced Central Franconian and other Old High German dialects. Old English , Old Frisian and (to a lesser degree) Old Saxon share the application of the Ingvaeonic nasal spirant law . Old Dutch was considerably less affected than those other three languages, but

4300-455: The Cretaceous . Butterflies have a four-stage life cycle , and like other holometabolous insects they undergo complete metamorphosis . Winged adults lay eggs on the food plant on which their larvae , known as caterpillars , will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in a chrysalis . When metamorphosis is complete, the pupal skin splits,

4400-478: The -n of the third-person plural hebban , which is absent in both Old English and Frisian, identifies the language as Old Dutch ( Old High German habent uses a different stem). Hagunnan and hi(c) have a prothetic h , which points also to West Flemish in which the h was frequently dropped or, in the written language, added before vowels (compare abent in the Latin version). However, it has been postulated that

4500-627: The Leiden Willeram translates as "All night long on my bed I looked for the one my heart loves; I looked for him but did not find him". The manuscript, now in the library of the Leiden University in the Netherlands, contains an Old Dutch translation of an Old High German (East Franconian) commentary on Song of Solomon , written by the German abbot Williram of Ebersberg . The translation was done by

4600-473: The Lycaenidae , form mutual associations with ants. They communicate with the ants using vibrations that are transmitted through the substrate as well as using chemical signals. The ants provide some degree of protection to these larvae and they in turn gather honeydew secretions . Large blue ( Phengaris arion ) caterpillars trick Myrmica ants into taking them back to the ant colony where they feed on

4700-581: The Wachtendonck Psalms ; it shows the evolution of Dutch, from the original Old Dutch, written c. 900, to modern Dutch, but so accurately copies the Latin word order of the original that there is little information that can be garnered on Old Dutch syntax . In Modern Dutch, recasting is necessary to form a coherent sentence. Old Dutch texts are extremely rare and much more limited than for related languages like Old English and Old High German . Most of

4800-469: The Weis-Fogh ' clap-and-fling ' mechanism. Butterflies are able to change from one mode to another rapidly. Butterflies are threatened in their early stages by parasitoids and in all stages by predators, diseases and environmental factors. Braconid and other parasitic wasps lay their eggs in lepidopteran eggs or larvae and the wasps' parasitoid larvae devour their hosts, usually pupating inside or outside

4900-452: The ZW sex-determination system where females are the heterogametic sex (ZW) and males homogametic (ZZ). Butterflies are distributed worldwide except Antarctica, totalling some 18,500 species. Of these, 775 are Nearctic ; 7,700 Neotropical ; 1,575 Palearctic ; 3,650 Afrotropical ; and 4,800 are distributed across the combined Oriental and Australian / Oceania regions. The monarch butterfly

5000-530: The lepidopteran suborder Rhopalocera , characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The group comprises the superfamilies Hedyloidea (moth-butterflies in the Americas) and Papilionoidea (all others). The oldest butterfly fossils have been dated to the Paleocene , about 56 million years ago, though molecular likely originated in

5100-402: The monarch and the painted lady , migrate over long distances. Many butterflies are attacked by parasites or parasitoids , including wasps , protozoans , flies , and other invertebrates, or are preyed upon by other organisms. Some species are pests because in their larval stages they can damage domestic crops or trees; other species are agents of pollination of some plants. Larvae of

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5200-600: The non-coding DNA around the cortex gene can turn a black-winged butterfly into a butterfly with a yellow wing band. When the butterfly Bicyclus anynana is subjected to repeated inbreeding in the laboratory, there is a dramatic decrease in egg hatching. This severe inbreeding depression is considered to be likely due to a relatively high mutation rate to recessive alleles with substantial damaging effects and infrequent episodes of inbreeding in nature that might otherwise purge such mutations. Although B. anynana experiences inbreeding depression when forcibly inbred in

5300-425: The 12th and 13th century, the practices were abandoned, and unstressed vowels were consistently written as e from that time onward. Notes: Old Dutch was spelt using the Latin alphabet. The length of a vowel was generally not represented in writing probably because the missionaries, who were the ones capable of writing and teaching how to write, tended to base the written language on Latin, which also did not make

5400-535: The Apollos ( Parnassius ) plugs her genital opening to prevent her from mating again. The vast majority of butterflies have a four-stage life cycle: egg , larva (caterpillar), pupa (chrysalis) and imago (adult). In the genera Colias , Erebia , Euchloe , and Parnassius , a small number of species are known that reproduce semi-parthenogenetically ; when the female dies, a partially developed larva emerges from her abdomen. Butterfly eggs are protected by

5500-563: The Netherlands before Old Dutch was spoken have been found, and they are sometimes called Oudnederlands (English: "Old Netherlandic" or "Old Dutch") in a geographic sense. The oldest known example, wad 'mudflat', is already mentioned c.  107–108 AD in Tacitus ' Histories (Book 5), in Latinised form as vadam (acc. sg.), as the name of a village, Vada , probably reflecting Early Germanic *wada . The word exclusively referred to

5600-533: The North of the Netherlands. In the rest of the coastal region, these dialects were mostly displaced following the withdrawal to England of the migrating Angles , Saxons and Jutes , who gave rise to Old English. It was largely replaced by Weser–Rhine Germanic dialects, spoken by the Salian Franks . It spread from northern Belgium and the southern Netherlands to the coast and evolved into Old Dutch. It has, however,

5700-517: The Old Dutch period, the distinction between the feminine ō -stems and ōn -stems began to disappear, when endings of one were transferred to the other declension and vice versa, as part of a larger process in which the distinction between the strong and weak inflection was being lost not only in feminine nouns but also in adjectives. The process is shown in a more advanced stage in Middle Dutch. Old Dutch reflects an intermediate form between Old Saxon and Old High German. Like Old High German, it preserved

5800-674: The Rhinelandic Rhyming Bible (Dutch: Rijnlandse Rijmbijbel ; German: Rheinische Reimbibel ). The verse translation of biblical histories is attested only in a series of fragments from different writers. It contains Old Dutch (Low Franconian), Low German (Low Saxon) and High German (Rhine-Franconian) elements. It was likely composed in the northwest of Germany in the early 12th century, possibly in Werden Abbey , near Essen . Phonologically, Old Dutch stands in between Old Saxon and Old High German , sharing some innovations with

5900-504: The Salic law code (the Malberg glosses ) contain several Old Dutch words and this full sentence written in the early 6th century, which is likely the earliest in the language. It translates as "I tell you: I am setting you free, serve". The phrase was used to free a serf . A lito (English: half-free ) was a form of serf in the feudal system , a half-free farmer, who was connected to the land of

6000-486: The Wachtendonck Psalms, the e and i merged in unstressed syllables, as did o and u . That led to variants like dagi and dage ("day", dative singular) and tungon and tungun ("tongue", genitive, dative, accusative singular and nominative, dative, accusative plural). The forms with e and o are generally found later on, showing the gradual reduction of the articulatory distinction, eventually merging into

6100-430: The abdomen, generally with short prolegs on segments 3–6 and 10; the three pairs of true legs on the thorax have five segments each. Many are well camouflaged; others are aposematic with bright colours and bristly projections containing toxic chemicals obtained from their food plants. The pupa or chrysalis, unlike that of moths, is not wrapped in a cocoon. Many butterflies are sexually dimorphic . Most butterflies have

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6200-405: The abdominal segments. These prolegs have rings of tiny hooks called crochets that are engaged hydrostatically and help the caterpillar grip the substrate. The epidermis bears tufts of setae , the position and number of which help in identifying the species. There is also decoration in the form of hairs, wart-like protuberances, horn-like protuberances and spines. Internally, most of the body cavity

6300-502: The adult colour pattern are marked by changes in the expression of particular transcription factors in the early pupa. The reproductive stage of the insect is the winged adult or imago . The surface of both butterflies and moths is covered by scales, each of which is an outgrowth from a single epidermal cell. The head is small and dominated by the two large compound eyes . These are capable of distinguishing flower shapes or motion but cannot view distant objects clearly. Colour perception

6400-433: The adult insect climbs out, expands its wings to dry, and flies off. Some butterflies, especially in the tropics, have several generations in a year, while others have a single generation, and a few in cold locations may take several years to pass through their entire life cycle. Butterflies are often polymorphic , and many species make use of camouflage , mimicry , and aposematism to evade their predators. Some, like

6500-467: The air for wind and scents. The antennae come in various shapes and colours; the hesperiids have a pointed angle or hook to the antennae, while most other families show knobbed antennae. The antennae are richly covered with sensory organs known as sensillae . A butterfly's sense of taste is coordinated by chemoreceptors on the tarsi , or feet, which work only on contact, and are used to determine whether an egg-laying insect's offspring will be able to feed on

6600-417: The ancestor of the modern Dutch verb root gun , through the addition of the prefix ge- . (An English cognate probably survives in to own (up) in the sense of 'to acknowledge, concede'.) Its modern meaning is roughly "to think someone deserves something, to derive satisfaction from someone else's success", and it is commonly translated as "grant" or "bestow". Maltho thi afrio lito Glosses to

6700-434: The ant eggs and larvae in a parasitic relationship. Caterpillars mature through a series of developmental stages known as instars . Near the end of each stage, the larva undergoes a process called apolysis , mediated by the release of a series of neurohormones . During this phase, the cuticle , a tough outer layer made of a mixture of chitin and specialized proteins , is released from the softer epidermis beneath, and

6800-658: The blues, greens, reds and iridescent colours are created by structural coloration produced by the micro-structures of the scales and hairs. As in all insects, the body is divided into three sections: the head, thorax , and abdomen . The thorax is composed of three segments, each with a pair of legs. In most families of butterfly the antennae are clubbed, unlike those of moths which may be threadlike or feathery. The long proboscis can be coiled when not in use for sipping nectar from flowers. Nearly all butterflies are diurnal , have relatively bright colours, and hold their wings vertically above their bodies when at rest, unlike

6900-422: The book never left the abbey, it cannot be regarded as the start of a Dutch literature and did not influence later works. Hebban olla vogala nestas hagunnan hinase hic enda thu, uuat unbidan uue nu. Arguably the most famous text containing Old Dutch, the fragment is translated as "All birds have started making nests, except me and you, what are we waiting for?" The text is dated from around 1100 and written by

7000-433: The deeds and words of the devil, Thunear, Wōden and Saxnōt, and all those fiends that are their companions". It mentions three Germanic pagan gods of the early Saxons which the reader is to forsake: Uuôden (" Woden "), Thunaer and Saxnōt . Scholar Rudolf Simek comments that the vow is of particular interest because it is the sole instance of the god Saxnōt mentioned in a religious context. One of many baptismal vows, it

7100-441: The desiccated husk. Most wasps are very specific about their host species and some have been used as biological controls of pest butterflies like the large white butterfly . When the small cabbage white was accidentally introduced to New Zealand, it had no natural enemies. In order to control it, some pupae that had been parasitised by a chalcid wasp were imported, and natural control was thus regained. Some flies lay their eggs on

7200-482: The earliest texts written in the Netherlands were written in Latin , rather than Old Dutch. Some of the Latin texts, however, contained Old Dutch words interspersed with the Latin text. Also, it is hard to determine whether a text actually was written in Old Dutch, as the Germanic languages spoken at that time were not standardised and were much more similar to one another. Several words that are known to have developed in

7300-525: The easternmost Dutch dialects, such as Limburgish . Before the advent of Old Dutch or any of the Germanic languages, Germanic dialects were mutually intelligible . The North Sea Germanic dialects were spoken in the whole of the coastal parts of the Netherlands and Belgium. Old Frisian was one of these dialects, and elements of it survived through the Frisian language , spoken in the province of Friesland in

7400-419: The epidermis begins to form a new cuticle. At the end of each instar, the larva moults , the old cuticle splits and the new cuticle expands, rapidly hardening and developing pigment. Development of butterfly wing patterns begins by the last larval instar. Caterpillars have short antennae and several simple eyes . The mouthparts are adapted for chewing with powerful mandibles and a pair of maxillae, each with

7500-501: The exception of Dutch, modern linguistic research has challenged the direct diachronical connection to Old Frankish for most of the varieties grouped under the broader "Franconian" category. Nevertheless, the traditional terminology of the West Germanic varieties along assumed Late Classical tribal lines, typical of 19th and early 20th century Germanic linguistics, remains common. Within historical linguistics Old Low Franconian

7600-710: The family Hesperiidae (skippers). Molecular clock estimates suggest that butterflies originated sometime in the Late Cretaceous , but only significantly diversified during the Cenozoic, with one study suggesting a North American origin for the group. The oldest American butterfly is the Late Eocene Prodryas persephone from the Florissant Fossil Beds , approximately 34   million years old. Butterflies are divided into seven families that contain

7700-431: The field of historical philology, the terminology for the oldest historical phase of the Dutch language traditionally includes both Old Dutch as well as Old Low Franconian . In English linguistic publications, Old Netherlandic is occasionally used in addition to the aforementioned terms. Old Low Franconian , derives from the linguistic category first devised by the German linguist Wilhelm Braune (1850–1926), who used

7800-535: The framing of Middle Dutch , Old East Low Franconian did not contribute much to Standard Dutch , which is based on the consolidated dialects of Holland and Brabant . During the Merovingian period, the Central Franconian dialects were influenced by Old Low Franconian (Old Dutch), resulting in certain linguistic loans which yielded a slight overlap of vocabulary, most of which relates to warfare . In addition

7900-409: The friction of their overlapping parts. The front two segments have a pair of spiracles which are used in respiration. The abdomen consists of ten segments and contains the gut and genital organs. The front eight segments have spiracles and the terminal segment is modified for reproduction. The male has a pair of clasping organs attached to a ring structure, and during copulation, a tubular structure

8000-544: The grass was growing. The earliest Lepidoptera fossils date to the Triassic - Jurassic boundary, around 200   million years ago. Butterflies evolved from moths, so while the butterflies are monophyletic (forming a single clade ), the moths are not. The oldest known butterfly is Protocoeliades kristenseni from the Palaeocene aged Fur Formation of Denmark, approximately 55   million years old, which belongs to

8100-1030: The laboratory it recovers within a few generation when allowed to breed freely. During mate selection, adult females do not innately avoid or learn to avoid siblings, implying that such detection may not be critical to reproductive fitness. Inbreeding may persist in B anynana because the probability of encountering close relatives is rare in nature; that is, movement ecology may mask the deleterious effect of inbreeding resulting in relaxation of selection for active inbreeding avoidance behaviors. Butterflies feed primarily on nectar from flowers. Some also derive nourishment from pollen , tree sap, rotting fruit, dung, decaying flesh, and dissolved minerals in wet sand or dirt. Butterflies are important as pollinators for some species of plants. In general, they do not carry as much pollen load as bees , but they are capable of moving pollen over greater distances. Flower constancy has been observed for at least one species of butterfly. Adult butterflies consume only liquids, ingested through

8200-437: The latter, and others with the former. The table below lists the consonantal phonemes of Old Dutch. For descriptions of the sounds and definitions of the terms, follow the links on the headings. Notes: Final-obstruent devoicing of Proto-Germanic [β] to [f] occurred across the West Germanic languages, and thus also in Old Dutch. Old Dutch spelling also reveals final devoicing of other consonants, namely: Final devoicing

8300-478: The lord for whom he worked but not owned by that lord. In contrast, a slave was fully owned by the lord. The Old Dutch word and the Modern Dutch counterpart laat are both etymologically and in meaning undoubtedly related to the verb root laat (English: 'let go', 'release'), which may indicate the fairly free status of such person in relation to that a slave. The Old Dutch word lito is particularly recognisable in

8400-506: The majority of moths which fly by night, are often cryptically coloured (well camouflaged), and either hold their wings flat (touching the surface on which the moth is standing) or fold them closely over their bodies. Some day-flying moths, such as the hummingbird hawk-moth , are exceptions to these rules. Butterfly larvae , caterpillars , have a hard ( sclerotised ) head with strong mandibles used for cutting their food, most often leaves. They have cylindrical bodies, with ten segments to

8500-401: The name is ancient, but modern Dutch and German use different words ( vlinder and Schmetterling ) and the common name often varies substantially between otherwise closely related languages. A possible source of the name is the bright yellow male of the brimstone ( Gonepteryx rhamni ); another is that butterflies were on the wing in meadows during the spring and summer butter season while

8600-404: The outside of caterpillars and the newly hatched fly larvae bore their way through the skin and feed in a similar way to the parasitoid wasp larvae. Predators of butterflies include ants, spiders, wasps, and birds. Old Dutch Old Dutch is regarded as the primary stage in the development of a separate Dutch language. It was spoken by the descendants of the Salian Franks who occupied what

8700-529: The proboscis. They sip water from damp patches for hydration and feed on nectar from flowers, from which they obtain sugars for energy, and sodium and other minerals vital for reproduction. Several species of butterflies need more sodium than that provided by nectar and are attracted by sodium in salt; they sometimes land on people, attracted by the salt in human sweat. Some butterflies also visit dung and scavenge rotting fruit or carcasses to obtain minerals and nutrients. In many species, this mud-puddling behaviour

8800-462: The psalms. They were named after a manuscript that has not survived but was the source from which scholars believe the surviving fragments must have been copied. The manuscript was once owned by Canon Arnold Wachtendonck. The surviving fragments are handwritten copies made by the Renaissance scholar Justus Lipsius in the sixteenth century. Lipsius made a number of separate copies of what appeared to be

8900-431: The quest for a suitable pupation site, often the underside of a leaf or other concealed location. There it spins a button of silk which it uses to fasten its body to the surface and moults for a final time. While some caterpillars spin a cocoon to protect the pupa, most species do not. The naked pupa, often known as a chrysalis, usually hangs head down from the cremaster, a spiny pad at the posterior end, but in some species

9000-451: The region and ground type that is now known as the Wadden Sea . However, since the word existed long before Old Dutch did (and even before its parent language, Frankish ), it cannot be considered part of the vocabulary of Old Dutch but rather of Proto-Germanic . Haþuþȳwas. Ann kusjam logūns. This sentence has been interpreted as "Haþuþyw's. I/He grant(s) a flame (i.e. brand, sword) to

9100-596: The same material, but the versions do not always agree. In addition, scholars conclude that the numerous errors and inconsistencies in the fragments point not only to some carelessness or inattentiveness by the Renaissance scholars but also to errors in the now-lost manuscript out of which the material was copied. The language of the Psalms suggests that they were originally written in the 10th century. Thes naghtes an minemo beddo vortheroda ich minen wino. Ich vortheroda hine ande ne vand sin niet. This example sentence taken from

9200-507: The select". It was discovered on a sword sheath mounting , excavated in 1996 in the Dutch village of Bergakker and is perhaps better described as Frankish than Old Dutch (Frankish was the direct parent language of Old Dutch). The text however, shows the beginning of Old Dutch morphology. The word ann , found in the partially-translated inscription is coined as the oldest Dutch by linguists Nicoline van der Sijs and Tanneke Schoonheim from Genootschap Onze Taal . They attribute that word to

9300-486: The six cases of Proto-Germanic: nominative , accusative , genitive and dative . A fifth case, the instrumental , could have also existed. The -s ending in the masculine plural was preserved in the coastal dialects, as can be seen in the Hebban Olla Vogala text where nestas is used instead of nesta . Later on, the -s ending entered Hollandic dialects and became part of the modern standard language. During

9400-441: The species display a unique territorial behavior, in which they stake out a territory typically 15 meters in diameter that contains larval host plants. They perch in this area and aggressively protect it from other insects and other male white peacocks. Seven subspecies are recognized. This Nymphalinae article is a stub . You can help Misplaced Pages by expanding it . Butterfly Butterflies are winged insects from

9500-454: The term Franconian as a wastebasket taxon for the early West Germanic texts that he could not readily classify as belonging to either Saxon , Alemannic or Bavarian and assumed to derive from the language of the Franks . He subsequently further divided this new grouping into Low , Middle and High Franconian based on the absence or presence of the Second Germanic consonant shift . With

9600-419: The terms Old Low Franconian or West Frankish to specifically refer to the (very sparsely attested) varieties of Old Dutch spoken prior its assimilation of the coastal dialect. Old Dutch itself is further divided into Old West Dutch and Old East Dutch, with the descendants of Old West Dutch forming the dominant basis of the Middle Dutch literary language and Old East Dutch forming a noticeable substrate within

9700-429: The text could equally well be Old English , more specifically Old Kentish . nu saget mir einen kuning other greven, the an uren got wille gelouven, that se sagent, that ist gelogen, thes ist thaz arme volc bedrogen. Translated as "Mention one king or earl who wants to believe in their god, what they say is a lie, that's how the people are being deceived", this fragment comes from an important source for Old Dutch:

9800-427: The verb's past tense lieten . End ec forsacho allum dioboles uuercum and uuordum, Thunær ende Uuôden ende Saxnôte ende allum thêm unholdum thê hira genôtas sint. The Utrecht Baptismal Vow , or Old Saxon Baptismal Vow , is a 9th-century baptismal vow that was found in a monastery library in the German city of Mainz but was written in the Dutch city of Utrecht . The sentence translates as "And I renounce all

9900-608: Was countered by the syllable-initial voicing of voiceless fricatives, which made [v] and [f] allophones of each other. Final devoicing appears much earlier in Old Dutch than it does Old Saxon and Old High German. In fact, by judging from the find at Bergakker , it would seem that the language already had inherited this characteristic from Old Frankish whereas Old Saxon and Old High German are known to have maintained word-final voiced obstruents much later (at least 900). Notes: In unstressed syllables, only three vowels seem to have been reliably distinguished: open, front and back. In

10000-418: Was mit mi The Wachtendonck Psalms are a collection of Latin psalms , with a translation in an eastern variety of Old Dutch (Old East Low Franconian) which contains a number of Old High German elements. The example sentence above translates as "He will deliver my soul in peace from those who attack me, for, amongst many, he was with me." Probably based on a Central Franconian original, very little remains of

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