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67-523: For prehistoric taxa, see text Galbuliformes Fürbringer, 1888 Nine families of largely arboreal birds make up the order Piciformes / ˈ p ɪ s ɪ f ɔːr m iː z / , the best-known of them being the Picidae , which includes the woodpeckers and close relatives. The Piciformes contain about 71 living genera with a little over 450 species , of which the Picidae make up about half. In general,

134-648: A Phanerozoic low in diversity by the late Oligocene, and the temperate forests and jungles of the Eocene were replaced by forest and scrubland. The closing of the Tethys Seaway destroyed its tropical biota. The Oi1 event of the Eocene-Oligocene transition covered the continent of Antarctica with ice sheets, leaving Nothofagus and mosses and ferns clinging to life around the periphery of Antarctica in tundra conditions. Angiosperms continued their expansion throughout

201-668: A broader trend of global cooling lasting from the Bartonian to the Rupelian. The transition is marked by the Oi1 event, an oxygen isotope excursion occurring approximately 33.55 million years ago, during which oxygen isotope ratios decreased by 1.3 ‰ . About 0.3–0.4 ‰ of this is estimated to be due to major expansion of Antarctic ice sheets. The remaining 0.9 to 1.0 ‰ was due to about 5 to 6 °C (9 to 10 °F) of global cooling . The transition likely took place in three closely spaced steps over

268-487: A critical threshold value. Brachiopod oxygen isotope ratios from New Zealand suggest that a proto-Subtropical Convergence developed during the Early Oligocene, with northern New Zealand being subtropical and southern and eastern New Zealand being cooled by cold, subantarctic water. Oligocene climate following the Eocene-Oligocene event is poorly known. There were several pulses of glaciation in middle Oligocene, about

335-491: A distinct Galbuliformes order. Analysis of nuclear genes confirms that they form a lineage of their own, but suggests that they are better treated as a suborder . The other families form another monophyletic group of suborder rank, but the barbets were determined to be paraphyletic with regard to the toucans and hence, the formerly all-encompassing Capitonidae have been split up. The woodpeckers and honeyguides are each other's closest relatives. According to some researchers,

402-417: A few species of three-toed woodpeckers. The jacamars aside, Piciformes do not have down feathers at any age, only true feathers. They range in size from the rufous piculet at 8 centimetres in length, and weighing 7 grams, to the toco toucan , at 63 centimetres long, and weighing 680 grams. All nest in cavities and have altricial young. The Galbulidae and Bucconidae are often separated into

469-554: A lack of widespread consensus within the scientific community for extended periods. The continual publication of new data and diverse opinions plays a crucial role in facilitating adjustments and ultimately reaching a consensus over time. The naming of families is codified by various international bodies using the following suffixes: The taxonomic term familia was first used by French botanist Pierre Magnol in his Prodromus historiae generalis plantarum, in quo familiae plantarum per tabulas disponuntur (1689) where he called

536-603: A link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion of grasslands , and a regression of tropical broad leaf forests to the equatorial belt . The start of the Oligocene is marked by a notable extinction event called the Grande Coupure ; it featured the replacement of European fauna with Asian fauna , except for

603-519: A major mountain chain in the Oligocene, as subduction became more direct into the coastline. Climate during the Oligocene reflected a general cooling trend following the Early Eocene Climatic Optimum . This transformed the Earth's climate from a greenhouse to an icehouse climate. The Eocene-Oligocene transition was a major cooling event and reorganization of the biosphere, being part of

670-460: A quarry at Massignano , Italy . However, this GSSP has been criticized as excluding the uppermost part of the type Eocene Priabonian Stage and because it is slightly earlier than important climate shifts that form natural markers for the boundary, such as the global oxygen isotope shift marking the expansion of Antarctic glaciation (the Oi1 event). The upper boundary of the Oligocene is defined by its GSSP at Carrosio , Italy , which coincides with

737-460: A semiarid prairie home and included entelodonts like Archaeotherium , camelids (such as Poebrotherium ), running rhinoceratoids , three-toed equids (such as Mesohippus ), nimravids , protoceratids , and early canids like Hesperocyon . Merycoidodonts, an endemic American group, were very diverse during this time. Australia and South America became geographically isolated and developed their own distinctive endemic fauna. These included

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804-499: Is commonly referred to as the "walnut family". The delineation of what constitutes a family— or whether a described family should be acknowledged— is established and decided upon by active taxonomists . There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to

871-617: Is discernible in pollen counts from the Tibetan Plateau, which also show that the South Asian Monsoon had already developed by the late Oligocene. Around 25.8 Ma, the South Asian Monsoon underwent an episode of major intensification brought on by the uplift of the Tibetan Plateau. A deep 400,000-year glaciated Oligocene-Miocene boundary event is recorded at McMurdo Sound and King George Island . The early Eocene climate

938-480: Is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy . It is classified between order and genus . A family may be divided into subfamilies , which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae , but that family

1005-485: The Genera Plantarum of George Bentham and Joseph Dalton Hooker this word ordo was used for what now is given the rank of family. Families serve as valuable units for evolutionary, paleontological, and genetic studies due to their relatively greater stability compared to lower taxonomic levels like genera and species. Oligocene The Oligocene is often considered an important time of transition,

1072-426: The archaeocete cetaceans began to decrease in diversity due to their lack of echolocation, which was very useful as the water became colder and cloudier. Other factors to their decline could include climate changes and competition with today's modern cetaceans and the requiem sharks , which also appeared in this epoch. Early desmostylians , like Behemotops , are known from the Oligocene. Pinnipeds appeared near

1139-651: The creodonts that had dominated the Paleocene in the Old World. Rodents and rabbits underwent tremendous diversification due to the increase in suitable habitats for ground-dwelling seed eaters, as habitats for squirrel-like nut- and fruit-eaters diminished. The primates, once present in Eurasia, were reduced in range to Africa and South America. Many groups, such as equids , entelodonts , rhinos , merycoidodonts , and camelids , became more able to run during this time, adapting to

1206-552: The first appearance of the foraminiferan Paragloborotalia kugleri and with the base of magnetic polarity chronozone C6Cn.2n. Oligocene faunal stages from youngest to oldest are: During the Oligocene Epoch, the continents continued to drift toward their present positions. Antarctica became more isolated as deep ocean channels were established between Antarctica and Australia and South America . Australia had been very slowly rifting away from West Antarctica since

1273-578: The Antarctic Circumpolar Current. There is some evidence that it occurred much earlier, during the early Eocene. The other major oceanic gateway opening during this time was the Tasman, or Tasmanian, depending on the paper, gateway between Australia and Antarctica. The time frame for this opening is less disputed than the Drake Passage and is largely considered to have occurred around 34 Ma. As

1340-689: The Arctic may have influenced this oceanic cooling, though the extent of this influence is still a matter of some significant dispute. The opening and closing of ocean gateways: the opening of the Drake Passage ; the opening of the Tasmanian Gateway and the closing of the Tethys seaway; along with the final formation of the Greenland – Iceland – Faroes Ridge; played vital parts in reshaping oceanic currents during

1407-528: The Dong Ho Formation of Oligocene age shows that the Oligocene flora of what is now Vietnam was very similar to its present flora. Kelps make their first appearance in the fossil record during the earliest Oligocene. Most extant mammal families had appeared by the end of the Oligocene. These included primitive three-toed horses, rhinoceroses, camels, deer, and peccaries. Carnivores such as dogs , nimravids , bears, weasels, and raccoons began to replace

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1474-402: The Drake Passage may also have been in place by the end of the early Oligocene. This may have been interrupted by a temporary constriction of the Drake Passage from sometime in the middle to late Oligocene (29 to 22 mya) to the middle Miocene (15 mya). The reorganization of the oceanic tectonic plates of the northeastern Pacific, which had begun in the Paleocene, culminated with the arrival of

1541-512: The Drake Passage opened and enabled the formation of the Antarctic Circumpolar Current (ACC), which would have kept the cold waters of Antarctica circulating around that continent and strengthened the formation of Antarctic Bottom Water (ABW). With the cold water concentrated around Antarctica, sea surface temperatures and, consequently, continental temperatures would have dropped. The onset of Antarctic glaciation occurred during

1608-928: The East African large igneous province , which also initiated rifting along the Red Sea and Gulf of Aden . The Alps were rapidly rising in Europe as the African Plate continued to push north into the Eurasian Plate , isolating the remnants of the Tethys Sea . Sea levels were lower in the Oligocene than in the early Eocene, exposing large coastal plains in Europe and the Gulf Coast and Atlantic Coast of North America. The Obik Sea , which had separated Europe from Asia, retreated early in

1675-555: The Eocene-Oligocene transition are not yet fully understood. The timing is wrong for this to be caused either by known impact events or by the volcanic activity on the Ethiopean Plateau. Two other possible drivers of climate change, not mutually exclusive, have been proposed. The first is thermal isolation of the continent of Antarctica by development of the Antarctic Circumpolar Current . Deep sea cores from south of New Zealand suggest that cold deep-sea currents were present by

1742-655: The Grande Coupure. The lowering of sea levels closed the Turgai Strait across the Obik Sea, which had previously separated Asia from Europe. This allowed Asian mammals, such as rhinoceroses and ruminants , to enter Europe and drive endemic species to extinction. Lesser faunal turnovers occurred simultaneously with the Oi2 event and towards the end of the Oligocene. There was significant diversification of mammals in Eurasia, including

1809-499: The Jurassic, but the exact timing of the establishment of ocean channels between the two continents remains uncertain. However, one estimate is that a deep channel was in place between the two continents by the end of the early Oligocene. The timing of the formation of the Drake Passage between South America and Antarctica is also uncertain, with estimates ranging from 49 to 17 mya (early Eocene to Miocene), but oceanic circulation through

1876-717: The Murray and Mendocino Fracture Zones at the North American subduction zone in the Oligocene. This initiated strike-slip movement along the San Andreas Fault and extensional tectonics in the Basin and Range province , ended volcanism south of the Cascades, and produced clockwise rotation of many western North American terranes. The Rocky Mountains were at their peak. A new volcanic arc was established in western North America, far inland from

1943-501: The New World and Old World monkeys. The South American continent was home to animals such as pyrotheres and astrapotheres , as well as litopterns and notoungulates . Sebecosuchians , terror birds , and carnivorous metatheres , like the borhyaenids remained the dominant predators. Africa was also relatively isolated and retained its endemic fauna. These included mastodonts, hyraxes, arsinoitheres, and other archaic forms. Egypt in

2010-702: The Norwegian-Greenland Sea indicate a drop in winter temperatures at high latitudes of about 5 °C (9.0 °F) just prior to the Oi1 event. Borehole dating from the Southeast Faroes drift indicates that deep-ocean circulation from the Arctic Ocean to the North Atlantic Ocean began in the early Oligocene. The best terrestrial record of Oligocene climate comes from North America, where temperatures dropped by 7 to 11 °C (13 to 20 °F) in

2077-594: The Norwegian-Greenland Sea indicated that glaciers had appeared in Greenland by the start of the Oligocene. Continental ice sheets in Antarctica reached sea level during the transition. Glacially rafted debris of early Oligocene age in the Weddell Sea and Kerguelen Plateau , in combination with Oi1 isotope shift, provides unambiguous evidence of a continental ice sheet on Antarctica by the early Oligocene. The causes of

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2144-401: The Oligocene exists mostly in isotopic proxies. Patterns of extinction and patterns of species migration can also be studied to gain insight into ocean conditions. For a while, it was thought that the glaciation of Antarctica may have significantly contributed to the cooling of the ocean, however, recent evidence tends to deny this. Isotopic evidence suggests that during the early Oligocene,

2211-507: The Oligocene was an environment of lush forested deltas. Nevertheless, the Early Oligocene saw a major reduction in the diversity of many Afro-Arabian mammal clades, including hyaenodonts, primates, and hystricognath and anomaluroid rodents. During the Oligocene, the Tethyan marine biodiversity hotspot collapsed as the Tethys Ocean contracted. The seas around Southeast Asia and Australia became

2278-516: The Oligocene, creating a persistent land connection between the continents. The Paratethys Sea stretched from what is now the Balkan Peninsula across Central Asia to the Tian Shan region of what is now Xinjiang . There appears to have been a land bridge in the early Oligocene between North America and Europe, since the faunas of the two regions are very similar. However, towards the end of

2345-492: The Oligocene, there was a brief marine incursion in Europe. The rise of the Himalayas during the Oligocene remains poorly understood. One recent hypothesis is that a separate microcontinent collided with south Asia in the early Eocene, and India itself did not collide with south Asia until the end of the Oligocene. The Tibetan Plateau may have reached nearly its present elevation by the late Oligocene. The Andes first became

2412-516: The Oligocene. The Oligocene saw the emergence of parrotfishes, as the centre of marine biodiversity shifted from the Central Tethys eastward into the Indo-Pacific. The fossil record of marine mammals is a little spotty during this time, and not as well known as the Eocene or Miocene, but some fossils have been found. The baleen whales and toothed whales had just appeared, and their ancestors,

2479-579: The Oligocene. As the continents shifted to a more modern configuration, so too did ocean circulation. The Drake Passage is located between South America and Antarctica . Once the Tasmanian Gateway between Australia and Antarctica opened, all that kept Antarctica from being completely isolated by the Southern Ocean was its connection to South America. As the South American continent moved north,

2546-577: The Pici. The modern families are known to exist since the mid-late Oligocene to early Miocene; consequently, the older forms appear to be more basal . A large part of Piciform evolution seems to have occurred in Europe where only Picidae occur today; perhaps even some now exclusively Neotropical families have their origin in the Old World . Order: PICIFORMES [REDACTED] Family (biology) Family ( Latin : familia , pl. : familiae )

2613-477: The Piciformes are insectivorous, although the barbets and toucans mostly eat fruit and the honeyguides are unique among birds in being able to digest beeswax (although insects make up the bulk of their diet). Nearly all Piciformes have parrot-like zygodactyl feet—two toes forward and two back, an arrangement that has obvious advantages for birds that spend much of their time on tree trunks. An exception are

2680-538: The atmosphere, contributing to global cooling. The gradual separation of the clump of continental crust and the deepening of the tectonic ridge in the North Atlantic that would become Greenland, Iceland, and the Faroe Islands helped to increase the deep water flow in that area. More information about the evolution of North Atlantic Deep Water will be given a few sections down. Evidence for ocean-wide cooling during

2747-708: The coast, reaching from central Mexico through the Mogollon-Datil volcanic field to the San Juan volcanic field , then through Utah and Nevada to the ancestral Northern Cascades. Huge ash deposits from these volcanoes created the White River and Arikaree Groups of the High Plains, with their excellent fossil beds. Between 31 and 26 mya, the Ethiopia-Yemen Continental Flood Basalts were emplaced by

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2814-446: The difficulty may be that there were strong regional variations in the response to climate shifts. Evidence of a relatively warm Oligocene suggests an enigmatic climate state, neither hothouse nor icehouse. The late Oligocene (26.5 to 24 mya) likely saw a warming trend in spite of low pCO2 levels, though this appears to vary by region. However, Antarctica remained heavily glaciated during this warming period. The late Oligocene warming

2881-417: The earliest Oligocene. This change is seen from Alaska to the Gulf Coast. Upper Eocene paleosols reflect annual precipitation of over a meter of rain, but early Oligocene precipitation was less than half this. In central North America, the cooling was by 8.2 ± 3.1 °C over a period of 400,000 years, though there is little indication of significant increase in aridity during this interval. Ice-rafted debris in

2948-487: The early Oligocene, and the effect of the Drake Passage opening on this glaciation has been the subject of much research. However, some controversy still exists as to the exact timing of the passage opening, whether it occurred at the start of the Oligocene or nearer the end. Even so, many theories agree that at the Eocene/Oligocene (E/O) boundary, a yet shallow flow existed between South America and Antarctica, permitting

3015-496: The early Oligocene. However, the timing of this event remains controversial. The other possibility, for which there is considerable evidence, is a drop in atmospheric carbon dioxide levels ( pCO2 ) during the transition. The pCO2 is estimated to have dropped just before the transition, to 760 ppm at the peak of ice sheet growth, then rebounded slightly before resuming a more gradual fall. Climate modeling suggests that glaciation of Antarctica took place only when pCO2 dropped below

3082-432: The end of the epoch from an otter -like ancestor. The Oligocene sees the beginnings of modern ocean circulation, with tectonic shifts causing the opening and closing of ocean gateways. Cooling of the oceans had already commenced by the Eocene/Oligocene boundary, and they continued to cool as the Oligocene progressed. The formation of permanent Antarctic ice sheets during the early Oligocene and possible glacial activity in

3149-484: The endemic rodent and marsupial families. By contrast, the Oligocene–Miocene boundary is not set at an easily identified worldwide event but rather at regional boundaries between the warmer late Oligocene and the relatively cooler Miocene. The lower boundary of the Oligocene (its Global Boundary Stratotype Section and Point or GSSP) is placed at the last appearance of the foraminiferan genus Hantkenina in

3216-944: The entire order Piciformes should be included as a subgroup in Coraciiformes . The phylogenetic relationship between the nine families that make up the order Piciformes is shown in the cladogram below. The number of species in each family is taken from the list maintained by Frank Gill , Pamela C. Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC). Galbulidae – jacamars (18 species) Bucconidae – puffbirds (38 species) Indicatoridae – honeyguides (16 species) Picidae – woodpeckers (240 species) Megalaimidae – Asian barbets (35 species) Lybiidae – African barbets (42 species) Capitonidae – New World barbets (15 species) Semnornithidae – toucan barbets (2 species) Ramphastidae – toucans (43 species) Reconstruction of

3283-810: The evolutionary history of the Piciformes has been hampered by poor understanding of the evolution of the zygodactyl foot. A number of prehistoric families and genera, from the Early Eocene Neanis and Hassiavis , the Zygodactylidae / Primoscenidae , Gracilitarsidae , Sylphornithidae , and " Homalopus " , to the Miocene "Picus" gaudryi and the Pliocene Bathoceleus are sometimes tentatively assigned to this order. There are some extinct ancestral Piciformes known from fossils which have been difficult to place but at least in part probably belong to

3350-530: The family as a rank intermediate between order and genus was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes, disposés dans un ordre naturel (1796). He used families (some of them were not named) in some but not in all his orders of "insects" (which then included all arthropods ). In nineteenth-century works such as the Prodromus of Augustin Pyramus de Candolle and

3417-783: The gateway widened, the Antarctic Circumpolar Current strengthened. The Tethys Seaway was not a gateway, but rather a sea in its own right. Its closing during the Oligocene had significant impact on both ocean circulation and climate. The collisions of the African plate with the European plate and of the Indian subcontinent with the Asian plate, cut off the Tethys Seaway that had provided a low-latitude ocean circulation. The closure of Tethys built some new mountains (the Zagros range) and drew down more carbon dioxide from

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3484-526: The giant indricotheres , that grew up to 6 meters (20 ft) at the shoulder and weighed up to 20 tons. Paraceratherium was one of the largest land mammals ever to walk the Earth. However, the indricotheres were an exception to a general tendency for Oligocene mammals to be much smaller than their Eocene counterparts. The earliest deer, giraffes, pigs, and cattle appeared in the mid-Oligocene in Eurasia. The first felid , Proailurus , originated in Asia during

3551-456: The late Oligocene and spread to Europe. There was only limited migration between Asia and North America. The cooling of central North America at the Eocene-Oligocene transition resulted in a large turnover of gastropods, amphibians, and reptiles. Mammals were much less affected. Crocodilians and pond turtles replaced by dry land tortoises. Molluscs shifted to more drought-tolerant forms. The White River Fauna of central North America inhabited

3618-769: The main source of deep water was the North Pacific and the Southern Ocean . As the Greenland-Iceland-Faroe Ridge sank and thereby connected the Norwegian–Greenland sea with the Atlantic Ocean, the deep water of the North Atlantic began to come into play as well. Computer models suggest that once this occurred, a more modern in appearance thermo-haline circulation started. Evidence for the early Oligocene onset of chilled North Atlantic deep water lies in

3685-448: The new dominant hotspot of marine biodiversity. At sea, 97% of marine snail species, 89% of clams, and 50% of echinoderms of the Gulf Coast did not survive past the earliest Oligocene. New species evolved, but the overall diversity diminished. Cold-water mollusks migrated around the Pacific Rim from Alaska and Siberia. The marine animals of Oligocene oceans resembled today's fauna, such as the bivalves . Calcareous cirratulids appeared in

3752-500: The opening of the Drake Passage is around 30 Ma. One of the possible issues with this timing was the continental debris cluttering up the seaway between the two plates in question. This debris, along with what is known as the Shackleton Fracture Zone , has been shown in a recent study to be fairly young, only about 8 million years old. The study concludes that the Drake Passage would be free to allow significant deep water flow by around 31 Ma. This would have facilitated an earlier onset of

3819-468: The opening of the Drake Passage is during the early Miocene. Despite the shallow flow between South America and Antarctica, there was not enough of a deep water opening to allow for significant flow to create a true Antarctic Circumpolar Current. If the opening occurred as late as hypothesized, then the Antarctic Circumpolar Current could not have had much of an effect on early Oligocene cooling, as it would not have existed. The earliest hypothesized time for

3886-517: The period from 33.8 to 33.5 mya. By the end of the transition, sea levels had dropped by 105 meters (344 ft), and ice sheets were 25% greater in extent than in the modern world. The effects of the transition can be seen in the geological record at many locations around the world. Ice volumes rose as temperature and sea levels dropped. Playa lakes of the Tibetan Plateau disappeared at the transition, pointing to cooling and aridification of central Asia. Pollen and spore counts in marine sediments of

3953-568: The period, grass was not quite common enough for modern savannas . In North America, much of the dense forest was replaced by patchy scrubland with riparian forests. Subtropical species dominated with cashews and lychee trees present, and temperate woody plants such as roses , beeches , and pines were common. The legumes spread, while sedges and ferns continued their ascent. In Europe, floral assemblages became increasingly affected by strengthening seasonality as it related to wildfire activity. The Ha Long megafossil flora from

4020-459: The plains that were spreading as the Eocene rainforests receded. Brontotheres died out in the Earliest Oligocene, and creodonts died out outside Africa and the Middle East at the end of the period. Multituberculates , an ancient lineage of primitive mammals that originated back in the Jurassic , also became extinct in the Oligocene, aside from the gondwanatheres . The Eocene-Oligocene transition in Europe and Asia has been characterized as

4087-512: The seventy-six groups of plants he recognised in his tables families ( familiae ). The concept of rank at that time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera , which is far from how the term is used today. In his work Philosophia Botanica published in 1751, Carl Linnaeus employed the term familia to categorize significant plant groups such as trees , herbs , ferns , palms , and so on. Notably, he restricted

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4154-432: The start of an Antarctic Circumpolar Current. Stemming from the issue of when the opening of the Drake Passage took place, is the dispute over how great of an influence the opening of the Drake Passage had on the global climate. While early researchers concluded that the advent of the ACC was highly important, perhaps even the trigger, for Antarctic glaciation and subsequent global cooling, other studies have suggested that

4221-537: The time of the Oi2 oxygen isotope shift. This led to the largest drop of sea level in past 100 million years, by about 75 meters (246 ft). This is reflected in a mid-Oligocene incision of continental shelves and unconformities in marine rocks around the world. Some evidence suggests that the climate remained warm at high latitudes even as ice sheets experienced cyclical growth and retreat in response to orbital forcing and other climate drivers. Other evidence indicates significant cooling at high latitudes. Part of

4288-444: The use of this term solely within the book's morphological section, where he delved into discussions regarding the vegetative and generative aspects of plants. Subsequently, in French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille was used as a French equivalent of the Latin ordo (or ordo naturalis ). In zoology ,

4355-401: The world as tropical and sub- tropical forests were replaced by temperate deciduous forests . Open plains and deserts became more common and grasses expanded from their water-bank habitat in the Eocene moving out into open tracts. The decline in pCO2 favored C4 photosynthesis , which is found only in angiosperms and is particularly characteristic of grasses. However, even at the end of

4422-414: The δ O signature is too strong for glaciation to be the main trigger for cooling. Through study of Pacific Ocean sediments, other researchers have shown that the transition from warm Eocene ocean temperatures to cool Oligocene ocean temperatures took only 300,000 years, which strongly implies that feedbacks and factors other than the ACC were integral to the rapid cooling. The latest hypothesized time for

4489-475: Was very warm, with crocodilians and temperate plants thriving north of the Arctic Circle . The cooling trend that began in the middle Eocene continued into the Oligocene, bringing both poles well below freezing for the first time in the Phanerozoic . The cooling climate, together with the opening of some land bridges and the closing of others, led to a profound reorganization of the biosphere and loss of taxonomic diversity. Land animals and marine organisms reached

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