Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.
101-524: Megarachne is a genus of eurypterid , an extinct group of aquatic arthropods . Fossils of Megarachne have been discovered in deposits of Late Carboniferous age, from the Gzhelian stage, in the Bajo de Véliz Formation of San Luis, Argentina . The fossils of the single and type species M. servinei have been recovered from deposits that had once been a freshwater environment. The generic name, composed of
202-496: A ricinuleid ". Even Hünicken himself acknowledged discrepancies in the morphology of the fossil that could not be accommodated with an arachnid identity. These discrepancies included an unusual cuticular ornamentation, the carapace being divided into frontal and rear parts by a suture, and spatulate (having a broad, rounded end) chelicerae (already noted by Hünicken as a strange feature, as no known spider possesses spatulate chelicerae); all features unknown in other spiders. However,
303-469: A body length of 54 cm (21 in). While large for an arthropod, Megarachne was dwarfed by other eurypterids , even relatively close relatives such as Hibbertopterus which could reach lengths exceeding 1.5 m (59 in). Though originally described as a giant spider , a multitude of features support the classification of Megarachne as a eurypterid. Among them, the raised lunules (the vaguely moon-shaped ornamentation , similar to scales) and
404-458: A circular structure behind the first body-segment - which he identified as the "moderately hairy" abdomen. Hünicken's identification relied heavily on X-ray microtomography of the holotype. Additional hidden structures – such as a sternum and labium , coxae and cheliceral fangs – were also extrapolated from the X-radiographs. With an estimated length of 33.9 cm (13.3 in) based on
505-718: A few in German , Dutch , Norwegian , Danish and Swedish . Latin is still spoken in Vatican City, a city-state situated in Rome that is the seat of the Catholic Church . The works of several hundred ancient authors who wrote in Latin have survived in whole or in part, in substantial works or in fragments to be analyzed in philology . They are in part the subject matter of the field of classics . Their works were published in manuscript form before
606-470: A flora dominated by different types of pteridosperms with pockets of isoetoid lycopsids . During Megarachne 's time, Argentina and the rest of South America was part of the ancient supercontinent Gondwana which was beginning to fuse with the northern continents of Euramerica , North China , Siberia and Kazakhstania to form Pangaea . In addition to Megarachne , the Bajo de Véliz Formation preserves
707-643: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as
808-621: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,
909-562: A new Classical Latin arose, a conscious creation of the orators, poets, historians and other literate men, who wrote the great works of classical literature , which were taught in grammar and rhetoric schools. Today's instructional grammars trace their roots to such schools , which served as a sort of informal language academy dedicated to maintaining and perpetuating educated speech. Philological analysis of Archaic Latin works, such as those of Plautus , which contain fragments of everyday speech, gives evidence of an informal register of
1010-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for
1111-472: A remarkable unity in phonological forms and developments, bolstered by the stabilising influence of their common Christian (Roman Catholic) culture. It was not until the Muslim conquest of Spain in 711, cutting off communications between the major Romance regions, that the languages began to diverge seriously. The spoken Latin that would later become Romanian diverged somewhat more from the other varieties, as it
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#17331144910011212-695: A small number of Latin services held in the Anglican church. These include an annual service in Oxford, delivered with a Latin sermon; a relic from the period when Latin was the normal spoken language of the university. In the Western world, many organizations, governments and schools use Latin for their mottos due to its association with formality, tradition, and the roots of Western culture . Canada's motto A mari usque ad mare ("from sea to sea") and most provincial mottos are also in Latin. The Canadian Victoria Cross
1313-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;
1414-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and
1515-425: A wide array of fossilized flying insects, such as Rigattoptera (classified in the order Protorthoptera ), but as a freshwater predator, Megarachne would probably not have fed on them. Instead, the blades on the frontal appendages of Megarachne would have allowed it to sweep-feed, raking through the soft sediment of the rivers it inhabited in order to capture and feed on small invertebrates. This feeding strategy
1616-411: Is Veritas ("truth"). Veritas was the goddess of truth, a daughter of Saturn, and the mother of Virtue. Switzerland has adopted the country's Latin short name Helvetia on coins and stamps, since there is no room to use all of the nation's four official languages . For a similar reason, it adopted the international vehicle and internet code CH , which stands for Confoederatio Helvetica ,
1717-420: Is a kind of written Latin used in the 3rd to 6th centuries. This began to diverge from Classical forms at a faster pace. It is characterised by greater use of prepositions, and word order that is closer to modern Romance languages, for example, while grammatically retaining more or less the same formal rules as Classical Latin. Ultimately, Latin diverged into a distinct written form, where the commonly spoken form
1818-626: Is a reversal of the original phrase Non terrae plus ultra ("No land further beyond", "No further!"). According to legend , this phrase was inscribed as a warning on the Pillars of Hercules , the rocks on both sides of the Strait of Gibraltar and the western end of the known, Mediterranean world. Charles adopted the motto following the discovery of the New World by Columbus, and it also has metaphorical suggestions of taking risks and striving for excellence. In
1919-596: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms
2020-638: Is highly fusional , with classes of inflections for case , number , person , gender , tense , mood , voice , and aspect . The Latin alphabet is directly derived from the Etruscan and Greek alphabets . Latin remains the official language of the Holy See and the Roman Rite of the Catholic Church at the Vatican City . The church continues to adapt concepts from modern languages to Ecclesiastical Latin of
2121-661: Is modelled after the British Victoria Cross which has the inscription "For Valour". Because Canada is officially bilingual, the Canadian medal has replaced the English inscription with the Latin Pro Valore . Spain's motto Plus ultra , meaning "even further", or figuratively "Further!", is also Latin in origin. It is taken from the personal motto of Charles V , Holy Roman Emperor and King of Spain (as Charles I), and
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#17331144910012222-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,
2323-958: Is taught at many high schools, especially in Europe and the Americas. It is most common in British public schools and grammar schools, the Italian liceo classico and liceo scientifico , the German Humanistisches Gymnasium and the Dutch gymnasium . Occasionally, some media outlets, targeting enthusiasts, broadcast in Latin. Notable examples include Radio Bremen in Germany, YLE radio in Finland (the Nuntii Latini broadcast from 1989 until it
2424-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with
2525-526: Is the most complete eurypterid discovered in Carboniferous deposits in South America so far. Megarachne was part of the stylonurine suborder, a relatively rare clade of eurypterids. Within the stylonurines, Megarachne is a member of the superfamily Mycteropoidea and its constituent family Mycteropidae; which includes the close relatives Woodwardopterus and Mycterops . Fossilized remains of
2626-531: The Corpus Inscriptionum Latinarum (CIL). Authors and publishers vary, but the format is about the same: volumes detailing inscriptions with a critical apparatus stating the provenance and relevant information. The reading and interpretation of these inscriptions is the subject matter of the field of epigraphy . About 270,000 inscriptions are known. The Latin influence in English has been significant at all stages of its insular development. In
2727-629: The Ancient Greek μέγας ( megas ) meaning "great" and Latin arachne meaning "spider", translates to "great spider". The holotype (now stored at the Museum of Paleontology at the National University of Córdoba ) was recovered from the Pallero Member of the Bajo de Véliz Formation of Argentina , which has been dated to the Gzhelian age : 303.7 to 298.9 million years ago. The specimen preserves
2828-506: The Ancient Greek μέγας ( megas ) meaning "great" and Ancient Greek ἀράχνη ( arachne ) meaning " spider ", translates to "great spider"; because the fossil was misidentified as a large, prehistoric spider. With a body length of 54 cm (21 in), Megarachne was a medium-sized eurypterid. If the original identification as a spider had been correct, Megarachne would have been the largest known spider to have ever lived. Eurypterids such as Megarachne are often called "sea scorpions", but
2929-538: The Bajo de Véliz Formation of Argentina, dated to the Gzhelian stage of the Late Carboniferous. The environment of the Bajo de Véliz formation was, unlike the typical living environments of eurypterids (especially the swimming eurypterids of the suborder Eurypterina ), a freshwater environment in a floodplain . Similar Late Carboniferous floodplains with fossilized remnants discovered in modern-day Australia suggest
3030-512: The Holy See , the primary language of its public journal , the Acta Apostolicae Sedis , and the working language of the Roman Rota . Vatican City is also home to the world's only automatic teller machine that gives instructions in Latin. In the pontifical universities postgraduate courses of Canon law are taught in Latin, and papers are written in the same language. There are
3131-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to
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3232-799: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;
3333-496: The Late Latin period, language changes reflecting spoken (non-classical) norms tend to be found in greater quantities in texts. As it was free to develop on its own, there is no reason to suppose that the speech was uniform either diachronically or geographically. On the contrary, Romanised European populations developed their own dialects of the language, which eventually led to the differentiation of Romance languages . Late Latin
3434-595: The Middle Ages as a working and literary language from the 9th century to the Renaissance , which then developed a classicizing form, called Renaissance Latin . This was the basis for Neo-Latin which evolved during the early modern period . In these periods Latin was used productively and generally taught to be written and spoken, at least until the late seventeenth century, when spoken skills began to erode. It then became increasingly taught only to be read. Latin grammar
3535-561: The Middle Ages , borrowing from Latin occurred from ecclesiastical usage established by Saint Augustine of Canterbury in the 6th century or indirectly after the Norman Conquest , through the Anglo-Norman language . From the 16th to the 18th centuries, English writers cobbled together huge numbers of new words from Latin and Greek words, dubbed " inkhorn terms ", as if they had spilled from a pot of ink. Many of these words were used once by
3636-407: The common language of international communication , science, scholarship and academia in Europe until well into the early 19th century, by which time modern languages had supplanted it in common academic and political usage. Late Latin is the literary language from the 3rd century AD onward. No longer spoken as a native language, Medieval Latin was used across Western and Catholic Europe during
3737-413: The cuticular sculpture of the mucrones (a dividing ridge continuing uninterrupted throughout the carapace , the part of the exoskeleton which covers the head) are especially important since these features are characteristic of eurypterids. Megarachne possessed blade-like structures on its appendages (limbs) which would have allowed it to engage in a feeding method known as sweep-feeding, raking through
3838-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,
3939-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,
4040-587: The strata in which Megarachne has been found indicates that it dwelled in freshwater and not in marine environments. Megarachne was similar to other eurypterids within the Mycteropoidea , a rare group known primarily from South Africa and Scotland. The mycteropoids had evolved a specialized method of feeding referred to as sweep-feeding. This involved raking through the substrate of riverbeds in order to capture and eat smaller invertebrates . Despite only two specimens having been recovered, Megarachne represents
4141-617: The British Crown. The motto is featured on all presently minted coinage and has been featured in most coinage throughout the nation's history. Several states of the United States have Latin mottos , such as: Many military organizations today have Latin mottos, such as: Some law governing bodies in the Philippines have Latin mottos, such as: Some colleges and universities have adopted Latin mottos, for example Harvard University 's motto
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4242-684: The Germanic and Slavic nations. It became useful for international communication between the member states of the Holy Roman Empire and its allies. Without the institutions of the Roman Empire that had supported its uniformity, Medieval Latin was much more liberal in its linguistic cohesion: for example, in classical Latin sum and eram are used as auxiliary verbs in the perfect and pluperfect passive, which are compound tenses. Medieval Latin might use fui and fueram instead. Furthermore,
4343-580: The Grinch Stole Christmas! , The Cat in the Hat , and a book of fairy tales, " fabulae mirabiles ", are intended to garner popular interest in the language. Additional resources include phrasebooks and resources for rendering everyday phrases and concepts into Latin, such as Meissner's Latin Phrasebook . Some inscriptions have been published in an internationally agreed, monumental, multivolume series,
4444-611: The Latin language. Contemporary Latin is more often studied to be read rather than spoken or actively used. Latin has greatly influenced the English language , along with a large number of others, and historically contributed many words to the English lexicon , particularly after the Christianization of the Anglo-Saxons and the Norman Conquest . Latin and Ancient Greek roots are heavily used in English vocabulary in theology ,
4545-461: The United States the unofficial national motto until 1956 was E pluribus unum meaning "Out of many, one". The motto continues to be featured on the Great Seal . It also appears on the flags and seals of both houses of congress and the flags of the states of Michigan, North Dakota, New York, and Wisconsin. The motto's 13 letters symbolically represent the original Thirteen Colonies which revolted from
4646-552: The University of Kentucky, the University of Oxford and also Princeton University. There are many websites and forums maintained in Latin by enthusiasts. The Latin Misplaced Pages has more than 130,000 articles. Italian , French , Portuguese , Spanish , Romanian , Catalan , Romansh , Sardinian and other Romance languages are direct descendants of Latin. There are also many Latin borrowings in English and Albanian , as well as
4747-420: The animal during its life) of each other based on their morphology and the size of the specimens. Should this interpretation be correct, the sparse mucrones of Megarachne might be because of its age; as Megarachne is significantly larger than Woodwardopterus . The smallest genus, Mycterops , has even more densely packed ornaments on its carapace and tergite; and thus might be the youngest, ontogenetic stage of
4848-800: The animal. Should Mycterops , Megarachne , and Woodwardopterus represent the same animal, the name-taking priority would be Mycterops ; as it was named first, in 1886. The cladogram below is adapted from Lamsdell and colleagues (2010) and shows the relationship of Megarachne within the suborder Stylonurina. Brachyopterus Kiaeropterus Brachyopterella Alkenopterus Rhenopterus Parastylonurus Stylonurella Pagea Stylonurus Laurieipterus Ctenopterus Lamontopterus Kokomopterus Hardieopterus Tarsopterella Hallipterus Drepanopterus Cyrtoctenus Hibbertopterus Woodwardopterus Megarachne Mycterops Both known specimens of Megarachne have been recovered from
4949-464: The assumption that the fossil was that of a spider, and with a leg-span estimated to be 50 centimetres (20 in), Megarachne servinei would have been the largest spider to have ever existed; exceeding the goliath birdeater ( Theraphosa blondi ), which has a maximum leg-span of around 30 cm (12 in). Because of its status as the "largest spider to have ever lived", Megarachne quickly became popular. Based on Hünicken's detailed description of
5050-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of
5151-535: The beginning of the Renaissance . Petrarch for example saw Latin as a literary version of the spoken language. Medieval Latin is the written Latin in use during that portion of the post-classical period when no corresponding Latin vernacular existed, that is from around 700 to 1500 AD. The spoken language had developed into the various Romance languages; however, in the educated and official world, Latin continued without its natural spoken base. Moreover, this Latin spread into lands that had never spoken Latin, such as
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#17331144910015252-425: The benefit of those who do not understand Latin. There are also songs written with Latin lyrics . The libretto for the opera-oratorio Oedipus rex by Igor Stravinsky is in Latin. Parts of Carl Orff 's Carmina Burana are written in Latin. Enya has recorded several tracks with Latin lyrics. The continued instruction of Latin is seen by some as a highly valuable component of a liberal arts education. Latin
5353-416: The carapace, the first two tergites, three partial appendages , and what is possibly a coxa (the proximal -most limb segment). Hünicken wrongly identified the specimen as a mygalomorph spider (the group that includes tarantulae ) based on the shape of the carapace; the 15-millimetre (0.59 in) wide, circular eye tubercle (round outgrowth) located in the center of the head between the two eyes; and
5454-430: The comic playwrights Plautus and Terence and the author Petronius . While often called a "dead language", Latin did not undergo language death . By the 6th to 9th centuries, natural language change eventually resulted in Latin as a vernacular language evolving into distinct Romance languages in the large areas where it had come to be natively spoken. However, even after the fall of Western Rome , Latin remained
5555-461: The country's full Latin name. Some film and television in ancient settings, such as Sebastiane , The Passion of the Christ and Barbarians (2020 TV series) , have been made with dialogue in Latin. Occasionally, Latin dialogue is used because of its association with religion or philosophy, in such film/television series as The Exorcist and Lost (" Jughead "). Subtitles are usually shown for
5656-493: The decline in written Latin output. Despite having no native speakers, Latin is still used for a variety of purposes in the contemporary world. The largest organisation that retains Latin in official and quasi-official contexts is the Catholic Church . The Catholic Church required that Mass be carried out in Latin until the Second Vatican Council of 1962–1965 , which permitted the use of the vernacular . Latin remains
5757-561: The development of European culture, religion and science. The vast majority of written Latin belongs to this period, but its full extent is unknown. The Renaissance reinforced the position of Latin as a spoken and written language by the scholarship by the Renaissance humanists . Petrarch and others began to change their usage of Latin as they explored the texts of the Classical Latin world. Skills of textual criticism evolved to create much more accurate versions of extant texts through
5858-413: The earliest extant Latin literary works, such as the comedies of Plautus and Terence . The Latin alphabet was devised from the Etruscan alphabet . The writing later changed from what was initially either a right-to-left or a boustrophedon script to what ultimately became a strictly left-to-right script. During the late republic and into the first years of the empire, from about 75 BC to AD 200,
5959-445: The fifteenth and sixteenth centuries, and some important texts were rediscovered. Comprehensive versions of authors' works were published by Isaac Casaubon , Joseph Scaliger and others. Nevertheless, despite the careful work of Petrarch, Politian and others, first the demand for manuscripts, and then the rush to bring works into print, led to the circulation of inaccurate copies for several centuries following. Neo-Latin literature
6060-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this
6161-456: The fossil specimen and various other illustrations and reconstructions made by him, reconstructions of Megarachne as a giant spider were set up in museums around the world. The identification of the specimen as a spider was doubted by some arachnologists , such as Shear and colleagues (1989), who stated that while Megarachne had been assigned to the Araneae, it "may represent an unnamed order or
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#17331144910016262-727: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,
6363-403: The genus, as a eurypterid, was only discovered well into production and by then it was far too late to update the reconstructions. The scenes were left in, but the giant spider was renamed as an unspecified species belonging to the primitive spider suborder Mesothelae , a suborder that actually exists but with genera much smaller than, and looking considerably different from, the spider featured in
6464-536: The history of Latin, and the kind of informal Latin that had begun to move away from the written language significantly in the post-Imperial period, that led ultimately to the Romance languages . During the Classical period, informal language was rarely written, so philologists have been left with only individual words and phrases cited by classical authors, inscriptions such as Curse tablets and those found as graffiti . In
6565-413: The holotype was by then deposited in a bank vault so other paleontologists only had access to plaster casts . In 2005, a second - more complete - specimen consisting of a part and counterpart (the matching halves of a compression fossil ) was recovered. This specimen had preserved parts of the front section of the body, as well as coxae possibly from the fourth pair of appendages, and was recovered from
6666-679: The invention of printing and are now published in carefully annotated printed editions, such as the Loeb Classical Library , published by Harvard University Press , or the Oxford Classical Texts , published by Oxford University Press . Latin translations of modern literature such as: The Hobbit , Treasure Island , Robinson Crusoe , Paddington Bear , Winnie the Pooh , The Adventures of Tintin , Asterix , Harry Potter , Le Petit Prince , Max and Moritz , How
6767-675: The language of the Roman Rite . The Tridentine Mass (also known as the Extraordinary Form or Traditional Latin Mass) is celebrated in Latin. Although the Mass of Paul VI (also known as the Ordinary Form or the Novus Ordo) is usually celebrated in the local vernacular language, it can be and often is said in Latin, in part or in whole, especially at multilingual gatherings. It is the official language of
6868-405: The language, Vulgar Latin (termed sermo vulgi , "the speech of the masses", by Cicero ). Some linguists, particularly in the nineteenth century, believed this to be a separate language, existing more or less in parallel with the literary or educated Latin, but this is now widely dismissed. The term 'Vulgar Latin' remains difficult to define, referring both to informal speech at any time within
6969-628: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,
7070-547: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Latin Latin ( lingua Latina , pronounced [ˈlɪŋɡʷa ɫaˈtiːna] , or Latinum [ɫaˈtiːnʊ̃] ) is a classical language belonging to the Italic branch of the Indo-European languages . Latin
7171-431: The meanings of many words were changed and new words were introduced, often under influence from the vernacular. Identifiable individual styles of classically incorrect Latin prevail. Renaissance Latin, 1300 to 1500, and the classicised Latin that followed through to the present are often grouped together as Neo-Latin , or New Latin, which have in recent decades become a focus of renewed study , given their importance for
7272-598: The modern glacial ice sheets of the Arctic and Antarctica , or smaller glaciers in dispersed centers. The spread of the ice sheets also affected sea levels, which would rise and fall throughout the period. Late Carboniferous flora was low in diversity but also developed uniformly throughout Gondwana. The plant life consisted of pteridosperm trees such as Nothorhacopteris , Triphyllopteris and Botrychiopsis , and lycopsid trees Malanzania , Lepidodendropsis and Bumbudendron . The plant fossils present also suggest that it
7373-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus
7474-455: The most complete eurypterid discovered in Carboniferous deposits in South America so far. Due to their fragmentary fossil record and similarities between the genera, some researchers have hypothesized that Megarachne and two other members of its family, Mycterops and Woodwardopterus , represent different developmental stages of a single genus. Known fossils of Megarachne indicate
7575-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this
7676-479: The parabolic shape of their prosoma (the head plate), hastate telsons (the hindmost part of the body being shaped like a gladius , a Roman sword) with paired keel -shaped projections on the underside, and heads with small compound eyes that were roughly trapezoidal in shape. Megarachne servinei was originally described in 1980 by the Argentine paleontologist Mario Hünicken. The generic name, composed of
7777-461: The program. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of
7878-526: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for
7979-466: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or
8080-460: The same locality and horizon . A research team led by the British paleontologist and arachnologist Paul A. Selden, and also consisting of Hünicken and Argentine arachnologist José A. Corronca, re-examined the holotype in light of the new discovery. They concluded that Megarachne servinei was a large eurypterid (a group also known as "sea scorpions"), not a spider. Although Hünicken had misidentified Megarachne , his identification of it as an arachnid
8181-421: The sciences , medicine , and law . A number of phases of the language have been recognized, each distinguished by subtle differences in vocabulary, usage, spelling, and syntax. There are no hard and fast rules of classification; different scholars emphasize different features. As a result, the list has variants, as well as alternative names. In addition to the historical phases, Ecclesiastical Latin refers to
8282-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in
8383-505: The second tergite of the mycteroptid Woodwardopterus were compared to the fossil remains of Megarachne by Selden and colleagues (2005), which revealed that they were virtually identical; including features previously not noted in Woodwardopterus , such as radiating lines covering the tergite. It was concluded that Megarachne and Woodwardopterus were part of the same family by Selden and colleagues (2005), with two primary differences:
8484-441: The soft sediment of aquatic environments in swamps and rivers with its frontal appendage blades to capture and feed on small invertebrates . Megarachne also possessed a large and circular second opisthosomal tergite (the second dorsal segment of the abdomen), the function of which remains unknown. Megarachne was very similar to other mycteroptid eurypterids in appearance, a group distinguished from other mycteropoids by
8585-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,
8686-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being
8787-618: The styles used by the writers of the Roman Catholic Church from late antiquity onward, as well as by Protestant scholars. The earliest known form of Latin is Old Latin, also called Archaic or Early Latin, which was spoken from the Roman Kingdom , traditionally founded in 753 BC, through the later part of the Roman Republic , up to 75 BC, i.e. before the age of Classical Latin . It is attested both in inscriptions and in some of
8888-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for
8989-560: The tergites and the mucrones on the carapace are more sparsely packed in Megarachne and the protrusion of the anteroedian (i.e. before the middle) carapace, seen prominently in Megarachne , does not occur in Woodwardopterus . It has been suggested that three of the four genera that constitute the Mycteroptidae, Mycterops , Woodwardopterus , and Megarachne might represent different ontogenetic stages (different developmental stages of
9090-566: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up
9191-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in
9292-422: The written form of Latin was increasingly standardized into a fixed form, the spoken forms began to diverge more greatly. Currently, the five most widely spoken Romance languages by number of native speakers are Spanish , Portuguese , French , Italian , and Romanian . Despite dialectal variation, which is found in any widespread language, the languages of Spain, France, Portugal, and Italy have retained
9393-730: Was also used as a convenient medium for translations of important works first written in a vernacular, such as those of Descartes . Latin education underwent a process of reform to classicise written and spoken Latin. Schooling remained largely Latin medium until approximately 1700. Until the end of the 17th century, the majority of books and almost all diplomatic documents were written in Latin. Afterwards, most diplomatic documents were written in French (a Romance language ) and later native or other languages. Education methods gradually shifted towards written Latin, and eventually concentrating solely on reading skills. The decline of Latin education took several centuries and proceeded much more slowly than
9494-539: Was common to other mycteropoids. In comparison to the comparatively warm climate of the earlier parts of the Carboniferous, the Late Carboniferous was relatively cold globally. This climate change likely occurred during the Middle Carboniferous due to falling CO 2 levels in the atmosphere and high oxygen levels. The Southern Hemisphere , where Argentina was and still is located, may even have experienced glaciation with large continental ice sheets similar to
9595-491: Was extensive and prolific, but less well known or understood today. Works covered poetry, prose stories and early novels, occasional pieces and collections of letters, to name a few. Famous and well regarded writers included Petrarch, Erasmus, Salutati , Celtis , George Buchanan and Thomas More . Non fiction works were long produced in many subjects, including the sciences, law, philosophy, historiography and theology. Famous examples include Isaac Newton 's Principia . Latin
9696-499: Was largely separated from the unifying influences in the western part of the Empire. Spoken Latin began to diverge into distinct languages by the 9th century at the latest, when the earliest extant Romance writings begin to appear. They were, throughout the period, confined to everyday speech, as Medieval Latin was used for writing. For many Italians using Latin, though, there was no complete separation between Italian and Latin, even into
9797-417: Was not entirely absurd; as the two groups are closely related. A morphological comparison with other eurypterids indicated that Megarachne most closely resembled another large Permo - Carboniferous eurypterid, the mycteroptid Woodwardopterus scabrosus , which is known only from a single specimen. Selden and colleagues (2005) concluded that, despite only being represented by two known specimens, Megarachne
9898-710: Was originally spoken by the Latins in Latium (now known as Lazio ), the lower Tiber area around Rome , Italy. Through the expansion of the Roman Republic it became the dominant language in the Italian Peninsula and subsequently throughout the Roman Empire . By the late Roman Republic , Old Latin had evolved into standardized Classical Latin . Vulgar Latin refers to the less prestigious colloquial registers , attested in inscriptions and some literary works such as those of
9999-515: Was perceived as a separate language, for instance early French or Italian dialects, that could be transcribed differently. It took some time for these to be viewed as wholly different from Latin however. After the Western Roman Empire fell in 476 and Germanic kingdoms took its place, the Germanic people adopted Latin as a language more suitable for legal and other, more formal uses. While
10100-478: Was shut down in June 2019), and Vatican Radio & Television, all of which broadcast news segments and other material in Latin. A variety of organisations, as well as informal Latin 'circuli' ('circles'), have been founded in more recent times to support the use of spoken Latin. Moreover, a number of university classics departments have begun incorporating communicative pedagogies in their Latin courses. These include
10201-450: Was subject to monsoons during certain time intervals. During the production of the 2005 British documentary Walking with Monsters , Megarachne was slated to appear as a giant tarantula -like spider hunting the cat-sized reptile Petrolacosaurus in the segment detailing the Carboniferous, with the reconstruction closely following what was thought to be known of the genus at the time the series began production. The actual identity of
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