A zoo (short for zoological garden ; also called an animal park or menagerie ) is a facility in which animals are kept within enclosures for public exhibition and often bred for conservation purposes.
204-664: The term zoological garden refers to zoology , the study of animals. The term is derived from the Ancient Greek ζῷον , zōion , 'animal', and the suffix -λογία , -logia , 'study of'. The abbreviation zoo was first used of the London Zoological Gardens , which was opened for scientific study in 1828, and to the public in 1847. The first modern zoo was the Tierpark Hagenbeck by Carl Hagenbeck in Germany . In
408-591: A legatus legionis in Hispania Tarraconensis , he supported the emperor against a revolt on the Rhine led by Antonius Saturninus . He then served as governor of Germania and Pannonia . In September 96, Domitian was succeeded by the elderly and childless Nerva , who proved to be unpopular with the army. After a revolt by members of the Praetorian Guard , Nerva decided to adopt as his heir and successor
612-532: A Traia . Their son, Trajan's namesake father Marcus Ulpius Traianus , was born at Italica during the reign of Tiberius and became a prominent senator and general, commanding the Legio X Fretensis under Vespasian in the First Jewish-Roman War . Trajan's mother was Marcia , a Roman noblewoman of the gens Marcia and a sister-in-law of the second Flavian Emperor Titus . Little is known of her. Her father
816-532: A bear and a bull , chained together, rolled in fierce combat across the sand ... Four hundred bears were killed in a single day under Caligula ... Under Nero , four hundred tigers fought with bulls and elephants. In a single day, at the dedication of the Colosseum by Titus , five thousand animals perished. Under Trajan ... lions , tigers, elephants, rhinoceroses , hippopotami, giraffes, bulls, stags, even crocodiles and serpents were employed to give novelty to
1020-448: A client kingdom in the first war (101–102), followed by a second war that ended in actual incorporation into the Empire of the trans-Danube border group of Dacia. According to the provisions of Decebalus's earlier treaty with Rome, made in the time of Domitian, Decebalus was acknowledged as rex amicus , that is, client king; in exchange for accepting client status, he received from Rome both
1224-508: A "house of animals" with a large collection of birds, mammals and reptiles in a garden tended by more than 600 employees. The garden was described by several Spanish conquerors, including Hernán Cortés in 1520. After the Aztec revolt against the Spanish rule, and during the subsequent battle for the city, Cortés reluctantly ordered the zoo to be destroyed. The oldest zoo in the world still in existence
1428-441: A Congolese pygmy , displayed in a cage with the chimpanzees, then with an orangutan named Dohong, and a parrot. The exhibit was intended as an example of the "missing link" between the orangutan and white man. It triggered protests from the city's clergymen, but the public reportedly flocked to see Benga. Humans were also displayed at various events, especially colonial expositions such as the 1931 Paris Colonial Exposition , with
1632-444: A Dacian nobleman called Bikilis was captured. Decebalus’ treasures had been buried under a temporarily diverted river and the captive workers executed to retain the secret. Staggering amounts of gold and silver were found and packed off to fill Rome's coffers. Trajan built a new city, Colonia Ulpia Traiana Augusta Dacica Sarmizegetusa , on another site (north of the hill citadel holding the previous Dacian capital), although bearing
1836-538: A banquet. The details of Trajan's early military career are obscure, save for the fact that in 89, as legate of Legio VII Gemina in Hispania Tarraconensis , he supported Domitian against an attempted coup by Lucius Antonius Saturninus , the governor of Germania Superior . Trajan probably remained in the region after the revolt was quashed, to engage with the Chatti who had sided with Saturninus, before returning
2040-626: A basis for further expansion within Eastern Europe, as the Romans believed the region to be much more geographically "flattened", and thus easier to traverse, than it actually was; they also underestimated the distance from those vaguely defined borders to the ocean. Defence of the province was entrusted to a single legion, the XIII Gemina , stationed at Apulum , which functioned as an advance guard that could, in case of need, strike either west or east at
2244-460: A change of mores that began with the Severan dynasty , Trajan's putative lovers included the future emperor, Hadrian, pages of the imperial household, the actor Pylades, a dancer called Apolaustus, Lucius Licinius Sura, and Trajan's predecessor Nerva. Cassius Dio also relates that Trajan made an ally out of Abgar VII on account of the latter's beautiful son, Arbandes, who would then dance for Trajan at
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#17328368842912448-503: A clear area first established by Domitian. Apollodorus of Damascus ' "magnificent" design incorporated a Triumphal arch entrance, a forum space approximately 120 m long and 90m wide, surrounded by peristyles: a monumentally sized basilica : and later, Trajan's Column and libraries. It was started in AD 107, dedicated on 1 January 112, and remained in use for at least 500 years. It still drew admiration when Emperor Constantius II visited Rome in
2652-404: A coin. In reality, Trajan did not share power in any meaningful way with the senate, something that Pliny admits candidly: "[E]verything depends on the whims of a single man who, on behalf of the common welfare, has taken upon himself all functions and all tasks". One of the most significant trends of his reign was his encroachment on the senate's sphere of authority, such as his decision to make
2856-480: A consequence, various management tools are used to preserve the space for the genetically most important individuals and to reduce the risk of inbreeding . Management of animal populations is typically through international organizations such as AZA and EAZA . Zoos have several different ways of managing the animal populations, such as moves between zoos, contraception , sale of excess animals and euthanization (culling). Contraception can be an effective way to limit
3060-586: A difference in wildlife conservation and education. Humans were occasionally displayed in cages at zoos along with non-human animals, to illustrate the differences between people of European and non-European origin. In September 1906, William Hornaday , director of the Bronx Zoo in New York—;with the agreement of Madison Grant , head of the New York Zoological Society —had Ota Benga ,
3264-645: A gas station. The animals may be trained to perform tricks, and visitors are able to get closer to them than in larger zoos. Since they are sometimes less regulated, roadside zoos are often subject to accusations of neglect and cruelty . In June 2014 the Animal Legal Defense Fund filed a lawsuit against the Iowa-based roadside Cricket Hollow Zoo for violating the Endangered Species Act by failing to provide proper care for its animals. Since filing
3468-609: A generous stipend and a steady supply of technical experts. The treaty seems to have allowed Roman troops the right of passage through the Dacian kingdom in order to attack the Marcomanni , Quadi and Sarmatians . However, senatorial opinion never forgave Domitian for paying what was seen as tribute to a barbarian king. Unlike the Germanic tribes, the Dacian kingdom was an organized state capable of developing alliances of its own, thus making it
3672-504: A grand scale. Trajan's reconstruction, completed by 103, was modestly described by Trajan himself as "adequate" for the Roman people. It replaced flammable wooden seating tiers with stone, and increased the Circus' already vast capacity by about 5,000 seats. Its lofty, elevated Imperial viewing box was rebuilt among the seating tiers, so that spectators could see their emperor sharing their enjoyment of
3876-492: A greenhouse with an aviary, aquarium, and museum of stuffed animals and birds), a pond, a bridge over a waterfall, an artificial lake with a fountain, a wood-ornamented greenhouse, a forest area, and enclosures and buildings. The first zoological garden in Australia was Melbourne Zoo in 1860. In German states leading roles came Berlin (1841), Frankfurt (1856), and Hamburg (1863). In 1907, the entrepreneur Carl Hagenbeck founded
4080-601: A laboratory setting. Ethologists have been particularly concerned with the evolution of behavior and the understanding of behavior in terms of the theory of natural selection . In one sense, the first modern ethologist was Charles Darwin , whose book, The Expression of the Emotions in Man and Animals , influenced many future ethologists. A subfield of ethology is behavioral ecology which attempts to answer Nikolaas Tinbergen 's four questions with regard to animal behavior: what are
4284-501: A lack of natural wild habitat in which to reintroduce animals. This highlights the importance of in situ conservation, or preservation of natural spaces, in addition to the utility of zoo captive breeding and reintroduction programs. In situ conservation and reintroduction programs are key elements to obtaining certification by reputable organisations such as the Association of Zoos and Aquariums (AZA). Efforts to clone endangered species in
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#17328368842914488-697: A lengthy tour of inspection on the Rhine and Danube frontiers, may suggest that he was unsure of his position, both in Rome and with the armies at the front. Alternatively, Trajan's keen military mind understood the importance of strengthening the empire's frontiers. His vision for future conquests required the diligent improvement of surveillance networks, defences and transport along the Danube . Prior to his frontier tours, Trajan ordered his Prefect Aelianus to attend him in Germany, where he
4692-412: A lump sum from the imperial treasury, and in return were expected to repay an annual sum to support the alimentary fund. The earliest of Trajan's conquests were Rome's two wars against Dacia , an area that had troubled Roman politics for over a decade in regard to the unstable peace negotiated by Domitian 's ministers with the powerful Dacian king Decebalus . Dacia would be reduced by Trajan's Rome to
4896-455: A matter of public interest in the 1970s, a few zoos began to consider making conservation their central role, with Gerald Durrell of the Jersey Zoo , George Rabb of Brookfield Zoo , and William Conway of the Bronx Zoo ( Wildlife Conservation Society ) leading the discussion. From then on, zoo professionals became increasingly aware of the need to engage themselves in conservation programs, and
5100-448: A model of cooperation for conservation. Loaned animals usually remain the property of the original park, and any offspring yielded by loaned animals are usually divided between the lending and holding institutions. For decades the capture of wild animals or purchasing of animals has been broadly considered unethical and has not been practiced by reputable zoos. Especially in large animals, a limited number of spaces are available in zoos. As
5304-487: A network of local notables who act as mediators between the ruled and the ruler. Dio's notion of being "friend" to Trajan (or any other Roman emperor), however, was that of an informal arrangement, that involved no formal entry of such "friends" into the Roman administration. Trajan ingratiated himself with the Greek intellectual elite by recalling to Rome many (including Dio) who had been exiled by Domitian, and by returning (in
5508-413: A new footing, by explaining the processes by which it can occur, and providing observational evidence that it had done so. Darwin's theory was rapidly accepted by the scientific community and soon became a central axiom of the rapidly developing science of biology. The basis for modern genetics began with the work of Gregor Mendel on peas in 1865, although the significance of his work was not realized at
5712-656: A new name is a strategy used by some zoo professionals to distance their institutions from the stereotypical and nowadays criticized zoo concept of the 19th century. The term "bio park" was first coined and developed by the National Zoo in Washington D.C. in the late 1980s. In 1993, the New York Zoological Society changed its name to the Wildlife Conservation Society and re branded the zoos under its jurisdiction as "wildlife conservation parks". The predecessor of
5916-406: A new perspective on the fundamental basis of life. Having previously been the realm of gentlemen naturalists, over the 18th, 19th and 20th centuries, zoology became an increasingly professional scientific discipline . Explorer-naturalists such as Alexander von Humboldt investigated the interaction between organisms and their environment, and the ways this relationship depends on geography, laying
6120-416: A new strategy that sets out the aims and mission of zoological gardens of the 21st century. When studying behaviour of captive animals, several things should however be taken into account before drawing conclusions about wild populations. Including that captive populations are often smaller than wild ones and that the space available to each animal is often less than in the wild. Conservation programs all over
6324-551: A noblewoman from the Roman settlement at Nîmes ; the marriage ultimately remained childless. The historian Cassius Dio later noted that Trajan was a lover of young men , in contrast to the usual bisexual activity that was common among upper-class Roman men of the period. The emperor Julian also made a sardonic reference to his predecessor's sexual preference, stating that Zeus himself would have had to be on guard had his Ganymede come within Trajan's vicinity. This distaste reflected
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6528-506: A pack of fools, yes, they treat you just like children, for we often offer children the most trivial things in place of things of greatest worth [...] In place of justice, in place of the freedom of the cities from spoliation or from the seizure of the private possessions of their inhabitants, in place of their refraining from insulting you [...] your governors hand you titles, and call you 'first' either by word of mouth or in writing; that done, they may thenceforth with impunity treat you as being
6732-433: A place of refuge . Today, many zoos are improving enclosures by including tactile and sensory features in the habitat that allow animals to encourage natural behaviors. These additions can prove to be effective in improving the lives of animals in captivity. The tactile and sensory features will vary depending on the species of animal. There are animals that are injured in the wild and are unable to survive on their own, but in
6936-601: A plan that failed. Decebalus also took prisoner Trajan's legate Longinus, who eventually poisoned himself while in custody. Finally, in 105, Decebalus undertook an invasion of Roman-occupied territory north of the Danube. Prior to the campaign, Trajan had raised two entirely new legions: II Traiana – which, however, may have been posted in the East, at the Syrian port of Laodicea – and XXX Ulpia Victrix , which
7140-438: A political intent, enabling planned increases in civil and military spending. Trajan formalised the alimenta, a welfare program that helped orphans and poor children throughout Italy by providing cash, food and subsidized education. The program was supported out of Dacian War booty, estate taxes and philanthropy. The alimenta also relied indirectly on mortgages secured against Italian farms ( fundi ). Registered landowners received
7344-479: A population's breeding. However it may also have health repercussions and can be difficult or even impossible to reverse in some animals. Additionally, some species may lose their reproductive capability entirely if prevented from breeding for a period (whether through contraceptives or isolation), but further study is needed on the subject. Sale of surplus animals from zoos was once common and in some cases animals have ended up in substandard facilities. In recent decades
7548-499: A privileged position. As Pliny said in one of his letters at the time, it was official policy that Greek civic elites be treated according to their status as notionally free but not put on an equal footing with their Roman rulers. When the city of Apamea complained of an audit of its accounts by Pliny, alleging its "free" status as a Roman colony, Trajan replied by writing that it was by his own wish that such inspections had been ordered. Concern about independent local political activity
7752-522: A process begun by Nerva) a great deal of private property that Domitian had confiscated. He also had good dealings with Plutarch , who, as a notable of Delphi , seems to have been favoured by the decisions taken on behalf of his home-place by one of Trajan's legates, who had arbitrated a boundary dispute between Delphi and its neighbouring cities. However, it was clear to Trajan that Greek intellectuals and notables were to be regarded as tools for local administration, and not be allowed to fancy themselves in
7956-709: A process involving the union of a male and female haploid gamete , each gamete formed by meiosis . Ordinarily, gametes produced by separate individuals unite by a process of fertilization to form a diploid zygote that can then develop into a genetically unique individual progeny. However, some animals are also capable, as an alternative reproductive process, to reproduce parthenogenetically. Parthenogenesis has been described in snakes and lizards (see Misplaced Pages Parthenogenesis in squamates ), in amphibians (see Misplaced Pages Parthenogenesis in amphibians ) and in numerous other species (see Misplaced Pages Parthenogenesis ). Generally, meiosis in parthanogenetically reproducing animals occurs by
8160-456: A public bath was built with the proceeds from the entrance fees paid by "supernumerary" members of the council, enrolled with Trajan's permission. According to the Digest , Trajan decreed that when a city magistrate promised to achieve a particular public building, his heirs inherited responsibility for its completion. Trajan was a prolific builder. Many of his buildings were designed and erected by
8364-656: A reversed light-dark cycle, i.e. only dim white or red lights are on during the day so the animals are active during visitor hours, and brighter lights on at night when the animals sleep. Special climate conditions may be created for animals living in extreme environments, such as penguins. Special enclosures for birds , mammals , insects , reptiles , fish , and other aquatic life forms have also been developed. Some zoos have walk-through exhibits where visitors enter enclosures of non-aggressive species, such as lemurs , marmosets , birds, lizards , and turtles . Visitors are asked to keep to paths and avoid showing or eating foods that
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8568-476: A role model, for, according to Pliny, "men learn better from examples". Eventually, Trajan's popularity among his peers was such that the Roman Senate bestowed upon him the honorific of optimus , meaning "the best", which appears on coins from 105 on. This title had mostly to do with Trajan's role as benefactor, such as in the case of his returning confiscated property. Pliny states that Trajan's ideal role
8772-461: A show". A side effect of such extravagant spending was that junior and thus less wealthy members of the local oligarchies felt disinclined to present themselves to fill posts as local magistrates, positions that involved ever-increasing personal expense. Roman authorities liked to play the Greek cities against one another – something of which Dio of Prusa was fully aware: [B]y their public acts [the Roman governors] have branded you as
8976-445: A similar process to that in sexually reproducing animals, but the diploid zygote nucleus is generated by the union of two haploid genomes from the same individual rather than from different individuals. Trajan Trajan ( / ˈ t r eɪ dʒ ən / TRAY -jən ; born Marcus Ulpius Traianus , 18 September 53 – c. 9 August 117 ) was a Roman emperor from AD 98 to 117, remembered as
9180-710: A small number of zoos. To solve this organizations like EAZA and AZA have begun to develop husbandry manuals. Many modern zoos attempt to improve animal welfare by providing more space and behavioural enrichments . This often involves housing the animals in naturalistic enclosures that allow the animals to express more of their natural behaviours, such as roaming and foraging. Whilst many zoos have been working hard on this change, in some zoos, some enclosures still remain barren concrete enclosures or other minimally enriched cages. Zoology Zoology ( UK : / z u ˈ ɒ l ə dʒ i / zoo- OL -ə-jee , US : / z oʊ ˈ ɒ l ə dʒ i / zoh- OL -ə-jee )
9384-428: A state of disorder", Pliny once wrote to Trajan, plans for unnecessary works made in collusion with local contractors being identified as one of the main problems. One of the compensatory measures proposed by Pliny expressed a thoroughly Roman conservative position: as the cities' financial solvency depended on the councilmen's purses, it was necessary to have more councilmen on the local city councils. According to Pliny,
9588-535: A strategic threat and giving Trajan a strong motive to attack it. In May of 101, Trajan launched his first campaign into the Dacian kingdom, crossing to the northern bank of the Danube and defeating the Dacian army at Tapae (see Second Battle of Tapae ), near the Iron Gates of Transylvania . It was not a decisive victory, however. Trajan's troops took heavy losses in the encounter, and he put off further campaigning for
9792-950: A strong local power base, caused by the size of the town from which they came, made it necessary for the Ulpii (and for the Aelii , the other important senatorial family of Italica with whom they were allied) to weave local alliances, in the Baetica (with the Annii , the Ucubi and perhaps the Dasumii from Corduba), the Tarraconense and the Narbonense , here above all through Pompeia Plotina , Trajan's wife. Many of these alliances were made not in Spain, but in Rome. The family home in Rome,
9996-547: A suitable alternative. Off-site conservation relies on zoos, national parks, or other care facilities to support the rehabilitation of the animals and their populations. Zoos benefit conservation by providing suitable habitats and care to endangered animals. When properly regulated, they present a safe, clean environment for the animals to increase populations sizes. A study on amphibian conservation and zoos addressed these problems by writing, Whilst addressing in situ threats, particularly habitat loss, degradation and fragmentation,
10200-484: A suitable environment for wild native animals such as herons to live in or visit. A colony of black-crowned night herons has regularly summered at the National Zoo in Washington, D.C. for more than a century. Some zoos may provide information to visitors on wild animals visiting or living in the zoo, or encourage them by directing them to specific feeding or breeding platforms. In modern, well-regulated zoos, breeding
10404-453: A very narrow territory under its direct administration. Trajan's year of birth is not reliably attested and may instead have been AD 56. The epitome of Cassius Dio's Roman history describes Trajan as "an Iberian and neither an Italian nor even an Italiote", but this claim is contradicted by other ancient sources and rejected by modern scholars, who have reconstructed Trajan's Italic lineage. Appian states that Trajan's hometown of Italica
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#173283688429110608-576: A year and have special programs for schools. They are organized by the Association of Zoos and Aquariums . Japan's first modern zoo, Tokyo's Ueno Imperial Zoological Gardens , opened in 1882 based on European models. In World War II it was used to teach the Japanese people about the lands recently conquered by the Army. In 1943, fearing American bombing attacks, the government ordered the zoo to euthanize dangerous animals that might escape. When ecology emerged as
10812-937: Is "the advancement of Zoology and Animal Physiology and the introduction of new and curious subjects of the Animal Kingdom." It maintains two research institutes, the Nuffield Institute of Comparative Medicine and the Wellcome Institute of Comparative Physiology. In the United States, the Penrose Research Laboratory of the Philadelphia Zoo focuses on the study of comparative pathology . The World Association of Zoos and Aquariums produced its first conservation strategy in 1993, and in November 2004, it adopted
11016-453: Is a form of scientific taxonomy . Modern biological classification has its root in the work of Carl Linnaeus , who grouped species according to shared physical characteristics. These groupings have since been revised to improve consistency with the Darwinian principle of common descent . Molecular phylogenetics , which uses nucleic acid sequence as data, has driven many recent revisions and
11220-657: Is administered by the International Code of Zoological Nomenclature . A merging draft, BioCode, was published in 1997 in an attempt to standardize nomenclature, but has yet to be formally adopted. Vertebrate zoology is the biological discipline that consists of the study of vertebrate animals, that is animals with a backbone , such as fish , amphibians , reptiles , birds and mammals . The various taxonomically oriented disciplines i.e. mammalogy , biological anthropology , herpetology , ornithology , and ichthyology seek to identify and classify species and study
11424-480: Is an integrative field of study, uniting concepts and information from evolutionary biology , taxonomy , ecology , physical geography , geology , paleontology and climatology . The origin of this field of study is widely accredited to Alfred Russel Wallace , a British biologist who had some of his work jointly published with Charles Darwin . Molecular biology studies the common genetic and developmental mechanisms of animals and plants, attempting to answer
11628-402: Is being studied. For example, what is learned about the physiology of yeast cells can also apply to human cells. The field of animal physiology extends the tools and methods of human physiology to non-human species. Physiology studies how, for example, the nervous , immune , endocrine , respiratory , and circulatory systems function and interact. Developmental biology is the study of
11832-512: Is believed to be Quintus Marcius Barea Sura . Her mother was Antonia Furnilla , daughter of Aulus Antonius Rufus and Furnia . Trajan owned some lands called Figlinae Marcianae in Ameria , another Umbrian town, located near both Tuder and Reate (the home of the Flavian dynasty) and believed to be the home of Marcia's family. The line of the Ulpii continued long after Trajan's death. His elder sister
12036-413: Is controlled to maintain a self-sustaining, global captive population. This is not the case in some less well-regulated zoos, often based in poorer regions. Overall "stock turnover" of animals during a year in a select group of poor zoos was reported as 20%-25% with 75% of wild caught apes dying in captivity within the first 20 months. The authors of the report stated that before successful breeding programs,
12240-406: Is done on both the microscopic and molecular levels for single-celled organisms such as bacteria as well as the specialized cells in multicellular organisms such as humans . Understanding the structure and function of cells is fundamental to all of the biological sciences. The similarities and differences between cell types are particularly relevant to molecular biology. Anatomy considers
12444-404: Is generally agreed that Pliny, being part of the emperor's inner circle, provides a unique and valuable source of information through his letters with Trajan, the only surviving correspondence between a governor and his emperor. However, it has been argued that Pliny's correspondence with Trajan is neither intimate nor candid, but rather an exchange of official mail in which Pliny's stance borders on
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#173283688429112648-443: Is likely to continue to do so. Biological classification belongs to the science of zoological systematics . Many scientists now consider the five-kingdom system outdated. Modern alternative classification systems generally start with the three-domain system : Archaea (originally Archaebacteria); Bacteria (originally Eubacteria); Eukaryota (including protists , fungi , plants , and animals ) These domains reflect whether
12852-507: Is lost with the exception of one sentence. Only fragments remain of the Getica , a book by Trajan's personal physician Titus Statilius Criton . The Parthica , a 17-volume account of the Parthian Wars written by Arrian , has met a similar fate. Book 68 in Greek author Cassius Dio 's Roman History , which survives mostly as Byzantine abridgements and epitomes , is the main source for
13056-462: Is of primary importance; for many amphibian species in situ conservation alone will not be enough, especially in light of current un-mitigatable threats that can impact populations very rapidly such as chytridiomycosis [an infectious fungal disease]. Ex situ programmes can complement in situ activities in a number of ways including maintaining genetically and demographically viable populations while threats are either better understood or mitigated in
13260-534: Is one of the few rulers whose reputation has survived 19 centuries. Every new emperor after him was honoured by the Senate with the wish felicior Augusto, melior Traiano (that he be "luckier than Augustus and better than Trajan"). Among medieval Christian theologians, Trajan was considered a virtuous pagan . In the Renaissance , Machiavelli , speaking on the advantages of adoptive succession over heredity, mentioned
13464-415: Is partly based on paleontology , which uses the fossil record to answer questions about the mode and tempo of evolution, and partly on the developments in areas such as population genetics and evolutionary theory. Following the development of DNA fingerprinting techniques in the late 20th century, the application of these techniques in zoology has increased the understanding of animal populations. In
13668-406: Is quantitative, and recently a significant amount of work has been done using computer science techniques such as bioinformatics and computational biology . Molecular genetics , the study of gene structure and function, has been among the most prominent sub-fields of molecular biology since the early 2000s. Other branches of biology are informed by molecular biology, by either directly studying
13872-698: Is reflected in zoology by the presence of animals in Egyptian hieroglyphics. Although the concept of zoology as a single coherent field arose much later, the zoological sciences emerged from natural history reaching back to the biological works of Aristotle and Galen in the ancient Greco-Roman world . In the fourth century BC, Aristotle looked at animals as living organisms, studying their structure, development and vital phenomena. He divided them into two groups: animals with blood, equivalent to our concept of vertebrates , and animals without blood, invertebrates . He spent two years on Lesbos , observing and describing
14076-462: Is said to have informed Hadrian in 108 that he had been chosen as Trajan's imperial heir. As governor of Upper Germany (Germania Superior) during Nerva's reign, Trajan received the impressive title of Germanicus for his skilful management and rule of the volatile Imperial province. When Nerva died on 28 January 98, Trajan succeeded to the role of emperor without any outward adverse incident. The fact that he chose not to hasten towards Rome, but made
14280-427: Is seen in Trajan's decision to forbid Nicomedia from having a corps of firemen ("If people assemble for a common purpose ... they soon turn it into a political society", Trajan wrote to Pliny) as well as in his and Pliny's fears about excessive civic generosities by local notables such as distribution of money or gifts. Pliny's letters suggest that Trajan and his aides were as much bored as they were alarmed by
14484-566: Is the Tiergarten Schönbrunn in Vienna , Austria. It was constructed by Adrian van Stekhoven in 1752 at the order of Emperor Francis I , to serve as an imperial menagerie as part of Schönbrunn Palace . The menagerie was initially reserved for the viewing pleasure of the imperial family and the court, but was made accessible to the public in 1765. In 1775, a zoo was founded in Madrid, and in 1795,
14688-559: Is the genus, and sapiens the specific epithet, both of them combined make up the species name. When writing the scientific name of an organism, it is proper to capitalize the first letter in the genus and put all of the specific epithet in lowercase. Additionally, the entire term may be italicized or underlined. The dominant classification system is called the Linnaean taxonomy . It includes ranks and binomial nomenclature . The classification, taxonomy , and nomenclature of zoological organisms
14892-569: Is the oldest zoo in Poland, opened in 1865 when the city was part of Prussia , and was home to about 10,500 animals representing about 1,132 species (in terms of the number of animal species, it is the third largest in the world). In 2014 the Wrocław Zoo opened the Africarium , the only themed oceanarium devoted solely to exhibiting the fauna of Africa , comprehensively presenting selected ecosystems from
15096-448: Is the scientific study of animals . Its studies include the structure , embryology , classification , habits , and distribution of all animals, both living and extinct , and how they interact with their ecosystems . Zoology is one of the primary branches of biology . The term is derived from Ancient Greek ζῷον , zōion ('animal'), and λόγος , logos ('knowledge', 'study'). Although humans have always been interested in
15300-614: Is the subfield of biology that studies the evolutionary processes (natural selection, common descent, speciation) that produced the diversity of life on Earth. Evolutionary research is concerned with the origin and descent of species , as well as their change over time, and includes scientists from many taxonomically oriented disciplines. For example, it generally involves scientists who have special training in particular organisms such as mammalogy , ornithology , herpetology , or entomology , but use those organisms as systems to answer general questions about evolution. Evolutionary biology
15504-517: The municipium of Italica in the present-day Andalusian province of Seville in southern Spain, an Italic settlement in Hispania Baetica ; his gens Ulpia came from the town of Tuder in the Umbria region of central Italy. His namesake father, Marcus Ulpius Traianus , was a general and distinguished senator. Trajan rose to prominence during the reign of Domitian ; in AD 89, serving as
15708-529: The American Zoo Association soon said that conservation was its highest priority. In order to stress conservation issues, many large zoos stopped the practice of having animals perform tricks for visitors. The Detroit Zoo , for example, stopped its elephant show in 1969, and its chimpanzee show in 1983, acknowledging that the trainers had probably abused the animals to get them to perform. Mass destruction of wildlife habitat has yet to cease all over
15912-695: The Epitome de Caesaribus , was the town of Tuder ( Todi ) in the Umbria region of central Italy . This is confirmed by archeology, with epigraphic evidence placing both the Ulpii and the Traii in Umbria generally and Tuder specifically, and by linguistic studies of the family names Ulpius and Traius which show that both are of Osco-Umbrian origin. It is unknown whether Trajan's ancestors were Roman citizens or not at their arrival in Spain. They would have certainly possessed Roman citizenship in case they arrived after
16116-519: The Roman currency , decreasing the silver content of the denarius from 93.5% to 89.0% – the actual silver weight dropping from 3.04 grams to 2.88 grams. This devaluation, along with the massive amounts of gold and silver acquired through his Dacian wars , allowed Trajan to mint many more denarii than his predecessors. He also withdrew from circulation silver denarii minted before Nero's devaluation. Trajan's devaluation may have had
16320-468: The Second Sophistic ; this "cultural patriotism" acted as a kind of substitute for the loss of political independence, and as such was shunned by Roman authorities. As Trajan himself wrote to Pliny: "These poor Greeks all love a gymnasium ... they will have to content with one that suits their real needs". The first known corrector was charged with a commission "to deal with the situation of
16524-472: The Social War (91–87 BC) , when Tuder became a municipium of Roman citizens. In Spain they may well have intermarried with native Iberians, in which case they would have lost their citizenship. Had they lacked or lost the status of Roman citizens, they would have achieved it or recovered it when Italica became a municipium with Latin rights in the mid-1st century BC. Trajan's paternal grandfather Ulpius married
16728-560: The Tierpark Hagenbeck in Stellingen, now a quarter of Hamburg . His zoo was a radical departure from the layout of the zoo that had been established in 1828. It was the first zoo to use open enclosures surrounded by moats, rather than barred cages, to better approximate animals' natural environments. He also set up mixed-species exhibits and based the layout on the different organizing principle of geography, as opposed to taxonomy. The Wrocław Zoo ( Polish : Ogród Zoologiczny we Wrocławiu )
16932-634: The United States alone, zoos are visited by over 181 million people annually. The London Zoo , which was opened in 1828, was initially known as the "Gardens and Menagerie of the Zoological Society of London", and it described itself as a menagerie or "zoological forest". The abbreviation "zoo" first appeared in print in the United Kingdom around 1847, when it was used for the Clifton Zoo , but it
17136-907: The Zoological Society of London in 1931 which today (2014) covers 600 acres (2.4 km). Since the early 1970s, an 1,800 acre (7 km) park in the San Pasqual Valley near San Diego has featured the San Diego Zoo Safari Park , run by the Zoological Society of San Diego. One of two state-supported zoo parks in North Carolina is the 2,000-acre (8.1 km) North Carolina Zoo in Asheboro. The 500-acre (2.0 km) Werribee Open Range Zoo in Melbourne, Australia, displays animals living in an artificial savannah . The first public aquarium
17340-545: The anatomy of different groups. It is closely related to evolutionary biology and phylogeny (the evolution of species). Physiology studies the mechanical, physical, and biochemical processes of living organisms by attempting to understand how all of the structures function as a whole. The theme of "structure to function" is central to biology. Physiological studies have traditionally been divided into plant physiology and animal physiology , but some principles of physiology are universal, no matter what particular organism
17544-550: The proximate causes of the behavior, the developmental history of the organism, the survival value and phylogeny of the behavior? Another area of study is animal cognition , which uses laboratory experiments and carefully controlled field studies to investigate an animal's intelligence and learning. Biogeography studies the spatial distribution of organisms on the Earth , focusing on topics like dispersal and migration , plate tectonics , climate change , and cladistics . It
17748-399: The taxonomist . Originally it was thought that species were immutable, but with the arrival of Darwin's theory of evolution, the field of cladistics came into being, studying the relationships between the different groups or clades . Systematics is the study of the diversification of living forms, the evolutionary history of a group is known as its phylogeny , and the relationship between
17952-415: The "contagion" of Christianity threatened everyone, regardless of gender, age, or rank. Pliny gave those accused of being Christians opportunity to deny it, and those who would not, he executed. Any who cursed Christ or recited a prayer to the gods or to Trajan’s statue were released. Pliny acknowledged that these were things that "those who are really Christians cannot be made to do." In 107, Trajan devalued
18156-441: The "tyrant" Domitian – attributes to him, at the time, various (and unspecified) feats of arms. Domitian's successor, Nerva , was unpopular with the army, and had been forced by his Praetorian Prefect Casperius Aelianus to execute Domitian's killers. Nerva needed the army's support to avoid being ousted. He accomplished this in the summer of 97 by naming Trajan as his adoptive son and successor, claiming that this
18360-573: The 11th century BCE. In the 2nd century BCE, the Chinese Empress Tanki had a "house of deer" built, and King Wen of Zhou kept a 1,500-acre (6.1 km) zoo called Ling-Yu, or the Garden of Intelligence. Other well-known collectors of animals included King Solomon of the Kingdom of Israel and Judah , Queen Semiramis and King Ashurbanipal of Assyria , and King Nebuchadnezzar of Babylonia . By
18564-489: The 16th century. During the Renaissance and early modern period, zoological thought was revolutionized in Europe by a renewed interest in empiricism and the discovery of many novel organisms. Prominent in this movement were Andreas Vesalius and William Harvey , who used experimentation and careful observation in physiology , and naturalists such as Carl Linnaeus , Jean-Baptiste Lamarck , and Buffon who began to classify
18768-470: The 1980s, developmental biology re-entered evolutionary biology from its initial exclusion from the modern synthesis through the study of evolutionary developmental biology . Related fields often considered part of evolutionary biology are phylogenetics , systematics , and taxonomy . Ethology is the scientific and objective study of animal behavior under natural conditions, as opposed to behaviorism , which focuses on behavioral response studies in
18972-463: The 4th century BCE, zoos existed in most of the Greek city states; Alexander the Great is known to have sent animals that he found on his military expeditions back to Greece. The Roman emperors kept private collections of animals for study or for use in the arena, the latter faring notoriously poorly. The 19th-century historian W. E. H. Lecky wrote of the Roman games , first held in 366 BCE: At one time,
19176-626: The Dacians, devoid of manoeuvring room, kept to their network of fortresses, which the Romans sought systematically to storm (see also Second Dacian War ). The Romans gradually tightened their grip around Decebalus' stronghold in Sarmizegetusa Regia , which they finally took and destroyed. A controversial scene on Trajan's column just before the fall of Sarmizegetusa Regia suggests that Decebalus may have offered poison to his remaining men as an alternative option to capture or death while trying to flee
19380-725: The Danubian lands; when Rome was weak, as during the Crisis of the Third Century , the province became a liability and was eventually abandoned. Trajan resettled Dacia with Romans and annexed it as a province of the Roman Empire. Aside from their enormous booty (over half a million slaves, according to John Lydus ), Trajan's Dacian campaigns benefited the Empire's finances through the acquisition of Dacia's gold mines, managed by an imperial procurator of equestrian rank ( procurator aurariarum ). On
19584-1073: The Domus Traiana, was on the Aventine Hill ; excavations under the Piazza del Tempio di Diana found remains thought to be of the family's large suburban villa, with evidence of highly decorated rooms. As a young man Trajan rose through the ranks of the Roman army , serving in some of the most contested parts of the empire's frontier. In 76–77, his father was Governor of Syria ( Legatus pro praetore Syriae ), where Trajan himself remained as Tribunus legionis . From there, after his father's replacement, he seems to have been transferred to an unspecified Rhine province, and Pliny implies that he engaged in active combat duty during both commissions. In about 86, Trajan's cousin Aelius Afer died, leaving his young children Hadrian and Paulina orphans. Trajan and his colleague Publius Acilius Attianus became co-guardians of
19788-434: The East, that meant the families of Greek notables. The Greeks, though, had their own memories of independence – and a commonly acknowledged sense of cultural superiority – and, instead of seeing themselves as Roman, disdained Roman rule. What the Greek oligarchies wanted from Rome was, above all, to be left in peace, to be allowed to exert their right to self-government (i.e., to be excluded from
19992-563: The Great , suffect consul in 116. Trajan created at least fourteen new senators from the Greek-speaking half of the empire, an unprecedented recruitment number that opens to question the issue of the "traditionally Roman" character of his reign, as well as the "Hellenism" of his successor Hadrian. But then Trajan's new Eastern senators were mostly very powerful and very wealthy men with more than local influence and much interconnected by marriage, so that many of them were not altogether "new" to
20196-552: The Iron Gate's gorge. A canal was built between the Danube's Kasajna tributary and Ducis Pratum, circumventing rapids and cataracts. Trajan's Forum Traiani was Rome's largest forum. It was built to commemorate his victories in Dacia , and was largely financed from that campaign's loot. To accommodate it, parts of the Capitoline and Quirinal Hills had to be removed, the latter enlarging
20400-559: The Royal House of Commagene , left behind him a funeral monument on the Mouseion Hill that was later disparagingly described by Pausanias as "a monument built to a Syrian man". As a senatorial Emperor, Trajan was inclined to choose his local base of political support from among the members of the ruling urban oligarchies. In the West, that meant local senatorial families like his own. In
20604-471: The Sarmatians living at the borders. Therefore, the indefensible character of the province did not appear to be a problem for Trajan, as the province was conceived more as a sally-base for further attacks. Even in the absence of further Roman expansion, the value of the province depended on Roman overall strength: while Rome was strong, the Dacian salient was an instrument of military and diplomatic control over
20808-548: The Senate. On the local level, among the lower section of the Eastern propertied, the alienation of most Greek notables and intellectuals towards Roman rule, and the fact that the Romans were seen by most such Greek notables as aliens, persisted well after Trajan's reign. One of Trajan's senatorial creations from the East, the Athenian Gaius Julius Antiochus Epiphanes Philopappos , a member of
21012-695: The Temple of Khnum at Esna . He built palatial villas outside Rome at Arcinazzo , at Centumcellae and at Talamone . He also built a bath complex as well as the Aqua Traiana . Trajan invested heavily in the provision of popular amusements. He carried out a "massive reconstruction" of the Circus Maximus , which was already the empire's biggest and best appointed circuit for the immensely popular sport of chariot racing . The Circus also hosted religious theatrical spectacles and games , and public processions on
21216-946: The U.S. and Canada by Species Survival Plans; in Australasia, by the Australasian Species Management Program; in Europe, by the European Endangered Species Program; and in Japan, South Asia, and South East Asia, by the Japanese Association of Zoos and Aquariums, the South Asian Zoo Association for Regional Cooperation , and the South East Asian Zoo Association. Besides conservation of captive species, large zoos may form
21420-541: The United States, Europe, and Asia are frequently embedded in zoos and zoological parks. The position of most modern zoos in Australasia , Asia , Europe , and North America , particularly those with scientific societies, is that they display wild animals primarily for the conservation of endangered species , as well as for research purposes and education, and secondarily for the entertainment of visitors. The Zoological Society of London states in its charter that its aim
21624-542: The VII Gemina legion to Legio in Hispania Tarraconensis. In 91 he held a consulate with Acilius Glabrio , a rarity in that neither consul was a member of the ruling dynasty. He held an unspecified consular commission as governor of either Pannonia or Germania Superior , or possibly both. Pliny – who seems to deliberately avoid offering details that would stress personal attachment between Trajan and
21828-528: The aid of the troops in his rearguard. The Dacians and their allies were repulsed after two battles in Moesia, at Nicopolis ad Istrum and Adamclisi . Trajan's army then advanced further into Dacian territory, and, a year later, forced Decebalus to submit. He had to renounce claim to some regions of his kingdom, return runaways from Rome then under his protection (most of them technical experts), and surrender all his war machines. Trajan returned to Rome in triumph and
22032-449: The animals and plants, considering the adaptations of different organisms and the function of their parts. Four hundred years later, Roman physician Galen dissected animals to study their anatomy and the function of the different parts, because the dissection of human cadavers was prohibited at the time. This resulted in some of his conclusions being false, but for many centuries it was considered heretical to challenge any of his views, so
22236-486: The animals might snatch. Some zoos keep animals in larger, outdoor enclosures, confining them with moats and fences, rather than in cages. Safari parks , also known as zoo parks and lion farms, allow visitors to drive through them and come in close proximity to the animals. Sometimes, visitors are able to feed animals through the car windows. The first safari park was Whipsnade Park in Bedfordshire, England, opened by
22440-648: The animals were moved to the Bulwark, renamed the Lion Tower, near the main western entrance of the Tower. It was opened to the public during the reign of Elizabeth I in the 16th century. During the 18th century, the price of admission was three half-pence, or the supply of a cat or dog for feeding to the lions. The animals were moved to the London Zoo when it opened. Aztec emperor Moctezuma had in his capital city of Tenochtitlan
22644-432: The animals' needs, but constraints such as size and expense can complicate this. The type of enclosure and the husbandry are of great importance in determining the welfare of animals. Substandard enclosures can lead to decreased lifespans, caused by factors as human diseases, unsafe materials in the cages and possible escape attempts (Bendow 382). However, when zoos take time to think about the animal's welfare, zoos can become
22848-434: The appointing of imperial correctores to audit the civic finances of the technically free Greek cities . The main goal was to curb the overenthusiastic spending on public works that served to channel ancient rivalries between neighbouring cities. As Pliny wrote to Trajan, this had as its most visible consequence a trail of unfinished or ill-kept public utilities. Competition among Greek cities and their ruling oligarchies
23052-511: The besieged capital with him. Decebalus fled but, when later cornered by Roman cavalry, committed suicide. His severed head, brought to Trajan by the cavalryman Tiberius Claudius Maximus , was later exhibited in Rome on the steps leading up to the Capitol and thrown on the Gemonian stairs . The famous Dacian treasures were not found in the captured capital and their whereabouts were only revealed when
23256-545: The best way to achieve this was to lower the minimum age for holding a seat on the council, making it possible for more sons of the established oligarchical families to join and thus contribute to civic spending; this was seen as preferable to enrolling non-noble wealthy upstarts. Such an increase in the number of council members was granted to Dio's city of Prusa, to the dismay of existing councilmen who felt their status lowered. A similar situation existed in Claudiopolis , where
23460-461: The case of the Galatian notable and "leading member of the Greek community" (according to one inscription) Gaius Julius Severus, who was a descendant of several Hellenistic dynasts and client kings. Severus was the grandfather of the prominent general Gaius Julius Quadratus Bassus , consul in 105. Other prominent Eastern senators included Gaius Julius Alexander Berenicianus , a descendant of Herod
23664-427: The cells have nuclei or not, as well as differences in the chemical composition of the cell exteriors. Further, each kingdom is broken down recursively until each species is separately classified. The order is: Domain ; kingdom ; phylum ; class ; order ; family ; genus ; species . The scientific name of an organism is generated from its genus and species. For example, humans are listed as Homo sapiens . Homo
23868-434: The clades can be shown diagrammatically in a cladogram . Although someone who made a scientific study of animals would historically have described themselves as a zoologist, the term has come to refer to those who deal with individual animals, with others describing themselves more specifically as physiologists, ethologists, evolutionary biologists, ecologists, pharmacologists, endocrinologists or parasitologists. Although
24072-406: The claims of Dio and other Greek notables to political influence based on what they saw as their "special connection" to their Roman overlords. Pliny tells of Dio of Prusa placing a statue of Trajan in a building complex where Dio's wife and son were buried – therefore incurring a charge of treason for placing the emperor's statue near a grave. Trajan, however, dropped the charge. Nevertheless, while
24276-632: The classical zoo, such as stage shows, roller coasters, and mythical creatures. Some examples are Busch Gardens Tampa Bay in Tampa, Florida , both Disney's Animal Kingdom and Gatorland in Orlando, Florida , Flamingo Land in North Yorkshire, England , and Six Flags Discovery Kingdom in Vallejo, California . By 2000 most animals being displayed in zoos were the offspring of other zoo animals. This trend, however
24480-564: The construction or reconstruction of Old Cairo 's Roman fortress (also known as "Babylon Fort") to Trajan, and the building of a canal between the River Nile and the Red Sea . In Egypt, Trajan was "quite active" in constructing and embellishing buildings. He is portrayed, together with Domitian , on the propylon of the Temple of Hathor at Dendera . His cartouche also appears in the column shafts of
24684-707: The construction, renovation, and expansion of zoos when the Great Depression severely reduced local budgets. It was "a new deal for animals." The Atlanta Zoo , founded in 1886, suffered neglect. By 1984 it was ranked among the ten worst zoos in the United States. Systematic reform by 2000 put it on the list of the ten best. By 2020, the United States featured 230 accredited zoos and aquariums across 45 states, accommodating 800,000 animals, and 6,000 species out of which about 1,000 are endangered. The zoos provide 208,000 jobs, and with an annual budget of $ 230 million for wildlife conservation . They attract over 200 million visits
24888-735: The continent of Africa . Housing over 10 thousand animals, the facility's breadth extends from housing insects such cockroaches to large mammals like elephants on an area of over 33 hectares . In the United States, the Philadelphia Zoo , opened on July 1, 1874, earning its motto "America's First Zoo." The Lincoln Park Zoological Gardens in Chicago and the Cincinnati Zoo opened in 1875. In the 1930s, federal relief programs provided financial aid to most local zoos. The Works Progress Administration and similar New Deal government agencies helped greatly in
25092-520: The culling proceeded. Although zoos in some countries have been open about culling, the controversy of the subject and pressure from the public has resulted in others being closed. This stands in contrast to most zoos publicly announcing animal births. Furthermore, while many zoos are willing to cull smaller and/or low-profile animals, fewer are willing to do it with larger high-profile species. Many animals breed readily in captivity. Zoos frequently are forced to intentionally limit captive breeding because of
25296-514: The determination of the double helical structure of the DNA molecule by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The study of systematics was transformed as DNA sequencing elucidated the degrees of affinity between different organisms. Zoology is the branch of science dealing with animals . A species can be defined as
25500-643: The discovery of the structure of DNA by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The history of zoology traces the study of the animal kingdom from ancient to modern times. Prehistoric people needed to study the animals and plants in their environment to exploit them and survive. Cave paintings, engravings and sculptures in France dating back 15,000 years show bison, horses, and deer in carefully rendered detail. Similar images from other parts of
25704-456: The diversity of life and the fossil record , as well as studying the development and behavior of organisms. Antonie van Leeuwenhoek did pioneering work in microscopy and revealed the previously unknown world of microorganisms , laying the groundwork for cell theory . van Leeuwenhoek's observations were endorsed by Robert Hooke ; all living organisms were composed of one or more cells and could not generate spontaneously. Cell theory provided
25908-567: The emperor and the Senate, especially after the supposed bloodiness that had marked Domitian's reign and his dealings with the Curia . By feigning reluctance to hold power, Trajan was able to start building a consensus around him in the Senate. His belated ceremonial entry into Rome in 99 was notably understated, something on which Pliny the Younger elaborated. By not openly supporting Domitian's preference for equestrian officers, Trajan appeared to conform to
26112-554: The enclosures and come into close proximity with the animals. The Zoological Society of London was founded in 1826 by Stamford Raffles and established the London Zoo in Regent's Park two years later in 1828. At its founding, it was the world's first scientific zoo. Originally intended to be used as a collection for scientific study , it was opened to the public in 1847. The Zoo was located in Regent's Park —then undergoing development at
26316-463: The existing quasi-urban Dacian settlements disappeared after the Roman conquest. A number of unorganized urban settlements ( vici ) developed around military encampments in Dacia proper – the most important being Apulum – but were only acknowledged as cities proper well after Trajan's reign. The main regional effort of urbanization was concentrated by Trajan at the rearguard, in Moesia, where he created
26520-575: The five successive good emperors "from Nerva to Marcus " – a trope out of which the 18th-century historian Edward Gibbon popularized the notion of the Five Good Emperors , of whom Trajan was the second. An account of the Dacian Wars , the Commentarii de bellis Dacicis , written by Trajan himself or a ghostwriter and modelled after Caesar 's Commentarii de Bello Gallico ,
26724-458: The food is supplied by the zoo, either from vending machines or a kiosk nearby. An animal theme park is a combination of an amusement park and a zoo, mainly for entertaining and commercial purposes. Marine mammal parks such as Sea World and Marineland are more elaborate dolphinariums keeping whales , and containing additional entertainment attractions. Another kind of animal theme park contains more entertainment and amusement elements than
26928-404: The forms of macroscopic structures such as organs and organ systems. It focuses on how organs and organ systems work together in the bodies of humans and other animals, in addition to how they work independently. Anatomy and cell biology are two studies that are closely related, and can be categorized under "structural" studies. Comparative anatomy is the study of similarities and differences in
27132-431: The foundations for biogeography , ecology and ethology . Naturalists began to reject essentialism and consider the importance of extinction and the mutability of species . These developments, as well as the results from embryology and paleontology , were synthesized in the 1859 publication of Charles Darwin 's theory of evolution by natural selection ; in this Darwin placed the theory of organic evolution on
27336-636: The fourth century. It accommodated Trajan's Market, and an adjacent brick market. Trajan was also a prolific builder of triumphal arches, many of which survive. He built roads, such as the Via Traiana , an extension of the Via Appia from Beneventum to Brundisium and the Via Traiana Nova , a mostly military road between Damascus and Aila , which Rome employed in its annexation of Nabataea and founding of Arabia Province . Some historians attribute
27540-535: The free cities", as it was felt that the old method of ad hoc intervention by the Emperor and/or the proconsuls had not been enough to curb the pretensions of the Greek notables. It is noteworthy that an embassy from Dio's city of Prusa was not favourably received by Trajan, and that this had to do with Dio's chief objective, which was to elevate Prusa to the status of a free city, an "independent" city-state exempt from paying taxes to Rome. Eventually, Dio gained for Prusa
27744-405: The gifted architect Apollodorus of Damascus , including a massive bridge over the Danube , which the Roman army and its reinforcements could use regardless of weather; the Danube sometimes froze over in winter, but seldom enough to bear the passage of a party of soldiers. Trajan's works at the Iron Gates region of the Danube created or enlarged the boardwalk road cut into the cliff-face along
27948-504: The hands of the architect John Nash . What set the London zoo apart from its predecessors was its focus on society at large. The zoo was established in the middle of a city for the public, and its layout was designed to cater for the large London population. The London zoo was widely copied as the archetype of the public city zoo. In 1853, the Zoo opened the world's first public aquarium . Dublin Zoo
28152-409: The high mortality rate was the reason for the "massive scale of importations." One 2-year study indicated that of 19,361 mammals that left accredited zoos in the U.S. between 1992 and 1998, 7,420 (38%) went to dealers, auctions, hunting ranches, unaccredited zoos and individuals, and game farms. The welfare of zoo animals varies widely. Many zoos work to improve their animal enclosures and make it fit
28356-422: The idea (developed by Pliny) that an emperor derived his legitimacy from his adherence to traditional hierarchies and senatorial morals. Therefore, he could point to the allegedly republican character of his rule. In a speech at the inauguration of his third consulship, on 1 January 100, Trajan exhorted the senate to share the care-taking of the empire with him – an event later celebrated on
28560-423: The importance of animal conservation, often through letting visitors witness the animals firsthand. Some critics, and the majority of animal rights activists, say that zoos, no matter their intentions, or how noble these intentions, are immoral and serve as nothing but to fulfill human leisure at the expense of the animals (an opinion that has spread over the years). However, zoo advocates argue that their efforts make
28764-449: The incorporation of Armenia , Mesopotamia , and Assyria as Roman provinces. In August AD 117, while sailing back to Rome, Trajan fell ill and died of a stroke in the city of Selinus . He was deified by the senate and his successor Hadrian (Trajan's cousin). According to historical tradition, Trajan's ashes were entombed in a small room beneath Trajan's Column . As an emperor, Trajan's reputation has endured – he
28968-476: The interactions of molecules in their own right such as in cell biology and developmental biology , or indirectly, where molecular techniques are used to infer historical attributes of populations or species , as in fields in evolutionary biology such as population genetics and phylogenetics . There is also a long tradition of studying biomolecules "from the ground up", or molecularly, in biophysics . Animals generally reproduce by sexual reproduction ,
29172-447: The interests of justice, and to reflect "the spirit of the age". Non-citizens who admitted to being Christians and refused to recant were to be executed "for obstinacy". Citizens were sent to Rome for trial. Further tests faced by Christians in Pontus are alluded to in correspondence between Pliny the Younger, governor of the Roman province of Bithynia and Pontus, and Emperor Trajan. Writing from Pontus in about AD 112, Pliny reported that
29376-404: The largest group of organisms in which any two individuals of the appropriate sex can produce fertile offspring; about 1.5 million species of animal have been described and it has been estimated that as many as 8 million animal species may exist. An early necessity was to identify the organisms and group them according to their characteristics, differences and relationships, and this is the field of
29580-399: The lawsuit, ALDF has obtained records from investigations conducted by the USDA's Animal and Plant Health Inspection Services; these records show that the zoo is also violating the Animal Welfare Act. A petting zoo, also called petting farms or children's zoos, features a combination of domestic animals and wild species that are docile enough to touch and feed. To ensure the animals' health,
29784-408: The more popular Trajan, who had distinguished himself in military campaigns against Germanic tribes. As emperor of Rome, Trajan oversaw the construction of building projects such as the forum named after him , the introduction of social welfare policies such as the alimenta , and new military conquests. He annexed Nabataea and Dacia , and his war against the Parthian Empire ended with
29988-831: The natural history of the animals they saw around them, and used this knowledge to domesticate certain species, the formal study of zoology can be said to have originated with Aristotle . He viewed animals as living organisms, studied their structure and development, and considered their adaptations to their surroundings and the function of their parts. Modern zoology has its origins during the Renaissance and early modern period, with Carl Linnaeus , Antonie van Leeuwenhoek , Robert Hooke , Charles Darwin , Gregor Mendel and many others. The study of animals has largely moved on to deal with form and function, adaptations, relationships between groups, behaviour and ecology. Zoology has increasingly been subdivided into disciplines such as classification , physiology , biochemistry and evolution . With
30192-419: The new cities of Nicopolis ad Istrum and Marcianopolis . A vicus was also created around the Tropaeum Traianum. The garrison city of Oescus received the status of Roman colony after its legionary garrison was redeployed. The fact that these former Danubian outposts had ceased to be frontier bases and were now in the deep rear acted as an inducement to their urbanization and development. Not all of Dacia
30396-413: The office of corrector was intended as a tool to curb any hint of independent political activity among local notables in the Greek cities, the correctores themselves were all men of the highest social standing entrusted with an exceptional commission. The post seems to have been conceived partly as a reward for senators who had chosen to make a career solely on the emperor's behalf. Therefore, in reality
30600-449: The other hand, commercial agricultural exploitation on the villa model, based on the centralized management of a huge landed estate by a single owner ( fundus ) was poorly developed. Therefore, use of slave labor in the province itself seems to have been relatively undeveloped, and epigraphic evidence points to work in the gold mines being conducted by means of labor contracts ( locatio conductio rei ) and seasonal wage-earning. The victory
30804-524: The political history of Trajan's rule. Besides this, Pliny the Younger 's Panegyricus and Dio Chrysostom 's orations are the best surviving contemporary sources. Both are adulatory perorations , typical of the High Imperial period, that describe an idealized monarch and an equally idealized view of Trajan's rule, and concern themselves more with ideology than with fact. The 10th volume of Pliny's letters contains his correspondence with Trajan, which deals with various aspects of imperial Roman government. It
31008-412: The populace; the more "serious matter" of the corn dole aimed to satisfy individuals. During the period of peace that followed the Dacian war, Trajan exchanged letters with Pliny the Younger on how best to deal with the Christians of Pontus . Trajan told Pliny to continue prosecutions of Christians if they merited that, but not to accept anonymous or malicious denunciations. He considered this to be in
31212-426: The post was conceived as a means for "taming" both Greek notables and Roman senators. It must be added that, although Trajan was wary of the civic oligarchies in the Greek cities, he also admitted into the senate a number of prominent Eastern notables already slated for promotion during Domitian's reign by reserving for them one of the twenty posts open each year for minor magistrates (the vigintiviri ). Such must be
31416-443: The practice continuing in Belgium at least to as late as 1958 in a "Congolese village" display at Expo '58 in Brussels. These displays, while sometimes called "human zoos", usually did not take place in zoos or use cages. Zoo animals live in enclosures that often attempt to replicate their natural habitats or behavioral patterns, for the benefit of both the animals and visitors. Nocturnal animals are often housed in buildings with
31620-494: The practice of selling animals from certified zoos has declined. A large number of animals are culled each year in zoos, but this is controversial. A highly publicized culling as part of population management was that of a healthy giraffe at Copenhagen Zoo in 2014. The zoo argued that his genes already were well-represented in captivity, making the giraffe unsuitable for future breeding. There were offers to adopt him and an online petition to save him had many thousand signatories, but
31824-517: The processes by which animals and plants reproduce and grow. The discipline includes the study of embryonic development , cellular differentiation , regeneration , asexual and sexual reproduction , metamorphosis , and the growth and differentiation of stem cells in the adult organism. Development of both animals and plants is further considered in the articles on evolution , population genetics , heredity , genetic variability , Mendelian inheritance , and reproduction . Evolutionary biology
32028-457: The provincial government, as was Italy) and to concentrate on their local interests. This was something the Romans were not disposed to do as from their perspective the Greek notables were shunning their responsibilities in regard to the management of Imperial affairs – primarily in failing to keep the common people under control, thus creating the need for the Roman governor to intervene. An excellent example of this Greek alienation
32232-493: The questions regarding the mechanisms of genetic inheritance and the structure of the gene . In 1953, James Watson and Francis Crick described the structure of DNA and the interactions within the molecule, and this publication jump-started research into molecular biology and increased interest in the subject. While researchers practice techniques specific to molecular biology, it is common to combine these with methods from genetics and biochemistry . Much of molecular biology
32436-655: The races, alongside his family and images of the gods, At some time during 108 or 109, Trajan held 123 days of games to celebrate his Dacian victory. They involved "fully 10,000" gladiators and the slaughter of thousands, "possibly tens of thousands," of animals, both wild and domestic. Trajan's careful management of public spectacles led the orator Fronto to congratulate him for paying equal attention to public entertainments and more serious issues, acknowledging that "neglect of serious matters can cause greater damage, but neglect of amusements greater discontent". State-funded public entertainments helped to maintain contentment among
32640-494: The results of his own investigations. His general approach was surprisingly modern, and he wrote, "For it is [the task] of natural science not simply to accept what we are told but to inquire into the causes of natural things." An early pioneer was Conrad Gessner , whose monumental 4,500-page encyclopedia of animals, Historia animalium , was published in four volumes between 1551 and 1558. In Europe, Galen's work on anatomy remained largely unsurpassed and unchallenged up until
32844-468: The right to become the head of the assize-district, conventus (meaning that Prusans did not have to travel to be judged by the Roman governor), but eleutheria (freedom, in the sense of full political autonomy) was denied. Eventually, it fell to Pliny, as imperial governor of Bithynia in AD 110, to deal with the consequences of the financial mess wrought by Dio and his fellow civic officials. "It's well established that [the cities' finances] are in
33048-503: The rise of the germ theory of disease , though the mechanism of inheritance remained a mystery. In the early 20th century, the rediscovery of Mendel's work led to the rapid development of genetics , and by the 1930s the combination of population genetics and natural selection in the modern synthesis created evolutionary biology . Research in cell biology is interconnected to other fields such as genetics, biochemistry , medical microbiology , immunology , and cytochemistry . With
33252-557: The same full name, Sarmizegetusa. This capital city was conceived as a purely civilian administrative centre and was provided the usual Romanized administrative apparatus ( decurions , aediles , etc.). Urban life in Roman Dacia seems to have been restricted to Roman colonists, mostly military veterans; there is no extant evidence for the existence in the province of peregrine cities. Native Dacians continued to live in scattered rural settlements, according to their own ways. In another arrangement with no parallels in any other Roman province,
33456-406: The second of the Five Good Emperors of the Nerva–Antonine dynasty . He was a philanthropic ruler and a successful soldier-emperor who presided over one of the greatest military expansions in Roman history , during which, by the time of his death, the Roman Empire reached its maximum territorial extent. He was given the title of Optimus ('the best') by the Roman Senate . Trajan was born in
33660-455: The senatorial provinces of Achaea and Bithynia into imperial ones in order to deal with the inordinate spending on public works by local magnates and the general mismanagement of provincial affairs by various proconsuls appointed by the Senate. In the formula developed by Pliny, however, Trajan was a "good" emperor in that, by himself, he approved or blamed the same things that the Senate would have approved or blamed. If in reality Trajan
33864-420: The servile. Some authors have even proposed that much of the text was written and/or edited by Trajan's Imperial secretary, his ab epistulis . Given the scarcity of literary sources, discussion of Trajan and his rule in modern historiography cannot avoid speculation. Non-literary sources such as archaeology, epigraphy , and numismatics are also useful for reconstructing his reign. Marcus Ulpius Traianus
34068-490: The spectacle. Charlemagne had an elephant named Abul-Abbas that was given to him by the Abbasid caliph. King Henry I of England kept a collection of animals at his palace in Woodstock which reportedly included lions, leopards, and camels. The most prominent collection in medieval England was in the Tower of London , created as early as 1204 by King John I . Henry III received a wedding gift in 1235 of three leopards from Frederick II, Holy Roman Emperor , and in 1264,
34272-499: The structures and mechanisms specific to those groups. The rest of the animal kingdom is dealt with by invertebrate zoology , a vast and very diverse group of animals that includes sponges , echinoderms , tunicates , worms , molluscs , arthropods and many other phyla , but single-celled organisms or protists are not usually included. Cell biology studies the structural and physiological properties of cells , including their behavior , interactions, and environment . This
34476-529: The study of anatomy stultified. During the post-classical era , Middle Eastern science and medicine was the most advanced in the world, integrating concepts from Ancient Greece, Rome, Mesopotamia and Persia as well as the ancient Indian tradition of Ayurveda , while making numerous advances and innovations. In the 13th century, Albertus Magnus produced commentaries and paraphrases of all Aristotle's works; his books on topics like botany , zoology, and minerals included information from ancient sources, but also
34680-516: The study of animal life is ancient, its scientific incarnation is relatively modern. This mirrors the transition from natural history to biology at the start of the 19th century. Since Hunter and Cuvier , comparative anatomical study has been associated with morphography , shaping the modern areas of zoological investigation: anatomy , physiology , histology , embryology , teratology and ethology . Modern zoology first arose in German and British universities. In Britain, Thomas Henry Huxley
34884-433: The time. Darwin gave a new direction to morphology and physiology , by uniting them in a common biological theory: the theory of organic evolution. The result was a reconstruction of the classification of animals upon a genealogical basis, fresh investigation of the development of animals, and early attempts to determine their genetic relationships. The end of the 19th century saw the fall of spontaneous generation and
35088-403: The tremendous expansion in the urbanization of London, led to a heightened demand for a greater variety of public forms of entertainment to be made available. The need for public entertainment, as well as the requirements of scholarly research, came together in the founding of the first modern zoos. Whipsnade Park Zoo in Bedfordshire , England, opened in 1931. It allowed visitors to drive through
35292-455: The two children. Trajan, in his late thirties, was created ordinary consul for the year 91. This early appointment may reflect the prominence of his father's career, as his father had been instrumental to the ascent of the ruling Flavian dynasty , held consular rank himself and had just been made a patrician . Around this time Trajan brought the architect and engineer Apollodorus of Damascus with him to Rome , and married Pompeia Plotina ,
35496-413: The very last!" These same Roman authorities had also an interest in assuring the cities' solvency and therefore ready collection of Imperial taxes. Last but not least, inordinate spending on civic buildings was not only a means to achieve local superiority, but also a means for the local Greek elites to maintain a separate cultural identity – something expressed in the contemporary rise of
35700-428: The wild The breeding of endangered species is coordinated by cooperative breeding programmes containing international studbooks and coordinators, who evaluate the roles of individual animals and institutions from a global or regional perspective, and there are regional programmes all over the world for the conservation of endangered species . In Africa, conservation is handled by the African Preservation Program (APP); in
35904-403: The world and many species such as elephants , big cats, penguins , tropical birds, primates, rhinos , exotic reptiles, and many others are in danger of dying out. Many of today's zoos hope to stop or slow the decline of many endangered species and see their primary purpose as breeding endangered species in captivity and reintroducing them into the wild. Modern zoos also aim to help teach visitors
36108-516: The world fight to protect species from going extinct , but many conservation programs are underfunded and under-represented. Conservation programs can struggle to fight bigger issues like habitat loss and illness. It often takes significant funding and long time periods to rebuild degraded habitats, both of which are scarce in conservation efforts. The current state of conservation programs cannot rely solely in situ (on-site conservation) plans alone, ex situ (off-site conservation) may therefore provide
36312-437: The world illustrated mostly the animals hunted for food and the savage animals. The Neolithic Revolution , which is characterized by the domestication of animals , continued throughout Antiquity. Ancient knowledge of wildlife is illustrated by the realistic depictions of wild and domestic animals in the Near East, Mesopotamia, and Egypt, including husbandry practices and techniques, hunting and fishing. The invention of writing
36516-461: The year in order to regroup and reinforce his army. Nevertheless, the battle was considered a Roman victory and Trajan strived to ultimately consolidate his position, including other major engagements, as well as the capture of Decebalus' sister as depicted on Trajan's Column. The following winter, Decebalus took the initiative by launching a counter-attack across the Danube further downstream, supported by Sarmatian cavalry, forcing Trajan to come to
36720-399: The zoo inside the Jardin des Plantes in Paris was founded by Jacques-Henri Bernardin , with animals from the royal menagerie at Versailles, primarily for scientific research and education. The planning about a space for the conservation and observation of animals was expressed in connection with the political construction of republican citizenship. The Kazan Zoo , the first zoo in Russia
36924-448: The zoological garden is the menagerie , which has a long history from the ancient world to modern times. The oldest known zoological collection was revealed during excavations at Hierakonpolis , Egypt in 2009, of a c. 3500 BCE menagerie. The exotic animals included hippopotami , hartebeest , elephants , baboons and wildcats . King Ashur-bel-kala of the Middle Assyrian Empire created zoological and botanical gardens in
37128-432: The zoos they can live out the rest of their lives healthy and happy (McGaffin). In recent years, some zoos have chosen to move out some larger animals because they do not have the space available to provide an adequate enclosure for them (Lemonic, McDowell, and Bjerklie 50). An issue with animal welfare in zoos is that best animal husbandry practices are often not completely known, especially for species that are only kept in
37332-552: Was Lucius Licinius Sura , a Roman senator born in Spain and the governor of Germania Inferior , who was Trajan's personal friend and became an official adviser of the Emperor. Sura was highly influential, and was appointed consul for a third term in 107. Some senators may have resented Sura's activities as a kingmaker and éminence grise , among them the historian Tacitus, who acknowledged Sura's military and oratorical talents, but compared his rapacity and devious ways to those of Vespasian 's éminence grise Licinius Mucianus . Sura
37536-420: Was Ulpia Marciana , and his niece was Salonia Matidia . Very little is known about Trajan's early formative years, but it is thought likely that he spent his first months or years in Italica before moving to Rome and then, perhaps at around eight or nine years of age, he almost certainly would have returned temporarily to Italica with his father during Trajanus's governorship of Baetica (ca. 64–65). The lack of
37740-483: Was a conservative one, argued as well by the orations of Dio Chrysostom—in particular his four Orations on Kingship , composed early during Trajan's reign. Dio, as a Greek notable and intellectual with friends in high places, and possibly an official friend to the emperor ( amicus caesaris ), saw Trajan as a defender of the status quo . In his third kingship oration, Dio describes an ideal king ruling by means of "friendship" – that is, through patronage and
37944-502: Was a prominent figure. His ideas were centered on the morphology of animals. Many consider him the greatest comparative anatomist of the latter half of the 19th century. Similar to Hunter , his courses were composed of lectures and laboratory practical classes in contrast to the previous format of lectures only. Scientific classification in zoology , is a method by which zoologists group and categorize organisms by biological type , such as genus or species . Biological classification
38148-406: Was an ex post facto fiction developed by authors writing under Trajan, including Tacitus and Pliny . According to the Historia Augusta , the future Emperor Hadrian brought word to Trajan of his adoption. Trajan retained Hadrian on the Rhine frontier as a military tribune , and Hadrian thus became privy to the circle of friends and relations with whom Trajan surrounded himself. Among them
38352-443: Was an autocrat, his deferential behavior towards his peers qualified him to be viewed as a virtuous monarch. The idea is that Trajan wielded autocratic power through moderatio instead of contumacia – moderation instead of insolence. In short, according to the ethics for autocracy developed by most political writers of the Imperial Roman Age, Trajan was a good ruler in that he ruled less by fear, and more by acting as
38556-492: Was and still is somewhat species-specific. When animals are transferred between zoos, they usually spend time in quarantine, and are given time to acclimatize to their new enclosures which are often designed to mimic their natural environment. For example, some species of penguins may require refrigerated enclosures. Guidelines on necessary care for such animals is published in the International Zoo Yearbook . Animal exchanges between facilities are usually made voluntarily, based on
38760-419: Was apparently executed forthwith ("put out of the way"), and his now-vacant post taken by Attius Suburanus . Trajan's accession, therefore, could qualify more as a successful coup than an orderly succession. On his entry to Rome, Trajan granted the plebs a direct gift of money. The traditional donative to the troops, however, was reduced by half. There remained the issue of the strained relations between
38964-433: Was between 150,000 and 175,000, while Decebalus could dispose of up to 200,000. Other estimates for the Roman forces involved in Trajan's second Dacian War cite around 86,000 for active campaigning with large reserves retained in the proximal provinces, and potentially much lower numbers around 50,000 for Decebalus' depleted forces and absent allies. In a fierce campaign that seems to have consisted mostly of static warfare,
39168-410: Was born on 18 September AD 53 in the Roman province of Hispania Baetica (in what is now Andalusia in modern Spain ), in the municipium of Italica (now in the municipal area of Santiponce , in the outskirts of Seville ), a Roman colony established in 206 BC by Scipio Africanus . At the time of Trajan's birth it was a small town, without baths, theatre and amphitheatre, and with
39372-441: Was entirely due to Trajan's outstanding military merits. There are hints, however, in contemporary literary sources that Trajan's adoption was imposed on Nerva. Pliny implied as much when he wrote that, although an emperor could not be coerced into doing something, if this was the way in which Trajan was raised to power, then it was worth it. Alice König argues that the notion of a natural continuity between Nerva's and Trajan's reigns
39576-473: Was founded in 1806 by the Professor of Kazan State University Karl Fuchs . Until the early 19th century, the function of the zoo was often to symbolize royal power, like King Louis XIV 's menagerie at Versailles . Major cities in Europe set up zoos in the 19th century, usually using London and Paris as models. The transition was made from princely menageries designed to entertain high society with strange novelties into public zoological gardens. The new goal
39780-532: Was granted the title Dacicus . The peace of 102 had returned Decebalus to the condition of more or less harmless client king; however, he soon began to rearm, to again harbour Roman runaways, and to pressure his Western neighbours, the Iazyges Sarmatians, into allying themselves with him. Through his efforts to develop an anti-Roman bloc, Decebalus prevented Trajan from treating Dacia as a protectorate instead of an outright conquest. In 104, Decebalus devised an attempt on Trajan's life by means of some Roman deserters,
39984-406: Was mainly for marks of pre-eminence, especially for titles bestowed by the Roman emperor. Such titles were ordered in a ranking system that determined how the cities were to be outwardly treated by Rome. The usual form that such rivalries took was that of grandiose building plans, giving the cities the opportunity to vie with each other over "extravagant, needless ... structures that would make
40188-451: Was not until some 20 years later that the shortened form became popular in the rhyming song " Walking in the Zoo " by music-hall artist Alfred Vance . The term "zoological park" was used for more expansive facilities in Halifax, Nova Scotia , Washington, D.C. , and the Bronx in New York, which opened in 1846, 1891 and 1899 respectively. Relatively new terms for zoos in the late 20th century are " conservation park" or "bio park". Adopting
40392-541: Was opened at the London Zoo in 1853. This was followed by the opening of public aquaria in continental Europe (e.g. Paris in 1859, Hamburg in 1864, Berlin in 1869, and Brighton in 1872) and the United States (e.g. Boston in 1859, Washington in 1873, San Francisco Woodward's Garden in 1873, and the New York Aquarium at Battery Park in 1896). Roadside zoos are found throughout North America , particularly in remote locations. They are often small, for-profit zoos, often intended to attract visitors to some other facility, such as
40596-522: Was opened in 1831 by members of the medical profession interested in studying animals while they were alive and more particularly getting hold of them when they were dead. Downs' Zoological Gardens created by Andrew Downs and opened to the Nova Scotia public in 1847. It was originally intended to be used as a collection for scientific study. By the early 1860s, the zoo grounds covered 40 hectares with many fine flowers and ornamental trees, picnic areas, statues, walking paths, The Glass House (which contained
40800-448: Was permanently occupied. After the post-Trajanic evacuation of lands across the lower Danube, land extending from the Danube to the inner arch of the Carpathian Mountains , including Transylvania , the Metaliferi Mountains and Oltenia was absorbed into the Roman province, which eventually took the form of an "excrescence" with ill-defined limits, stretching from the Danube northwards to the Carpathians . This may have been intended as
41004-442: Was posted to Brigetio , in Pannonia . By 105, the concentration of Roman troops assembled in the middle and lower Danube amounted to fourteen legions (up from nine in 101) – about half of the entire Roman army. Even after the Dacian wars, the Danube frontier would permanently replace the Rhine as the main military axis of the Roman Empire. Including auxiliaries , the number of Roman troops engaged on both campaigns
41208-414: Was settled by and named after Italic veterans who fought in Spain under Scipio, and new settlers arrived there from Italy in the following centuries. Among the Italic settlers were the Ulpii and the Traii , who were either part of the original colonists or arrived as late as the end of the 1st century BC. Their original home, according to the description of Trajan as "Ulpius Traianus ex urbe Tudertina" in
41412-425: Was the personal role played by Dio's relationship with Trajan. Dio is described by Philostratus as Trajan's close friend, and Trajan as supposedly engaging publicly in conversations with Dio. Nevertheless, as a Greek local magnate with a taste for costly building projects and pretensions of being an important political agent for Rome, Dio of Prusa was actually a target for one of Trajan's authoritarian innovations:
41616-420: Was to educate the entire population with information along modern scientific lines. Zoos were supported by local commercial or scientific societies. The modern zoo that emerged in the 19th century in the United Kingdom , was focused on providing scientific study and later educational exhibits to the public for entertainment and inspiration. A growing fascination for natural history and zoology , coupled with
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