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109-400: Sciaena undecimalis Bloch, 1792 The common snook ( Centropomus undecimalis ) is a species of marine fish in the family Centropomidae of the order Perciformes . The common snook is also known as the sergeant fish or robalo . It was originally assigned to the sciaenid genus Sciaena ; Sciaena undecimradiatus and Centropomus undecimradiatus are obsolete synonyms for

218-487: A cold snap . Snook are very susceptible to cold temperatures, with the effects ranging from the complete halt of all feeding at a water temperature of 14.2 °C (57.6 °F), to the loss of equilibrium at 12.7 °C (54.9 °F), to death at a temperature of 12.5 °C (54.5 °F). A cold snap in January 2010 resulted in a 41.88% decline in nominal abundance of the common snook population in southwest Florida from

327-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles

436-542: A two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,

545-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed

654-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this

763-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing

872-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by

981-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider

1090-573: A harsh environment for organisms. Sediment often settles in intertidal mudflats which are extremely difficult to colonize. No points of attachment exist for algae , so vegetation based habitat is not established. Sediment can also clog feeding and respiratory structures of species, and special adaptations exist within mudflat species to cope with this problem. Lastly, dissolved oxygen variation can cause problems for life forms. Nutrient-rich sediment from human-made sources can promote primary production life cycles, perhaps leading to eventual decay removing

1199-469: A high level of habitat plasticity . Common snook are opportunistic predators whose feeding habits indicate a positive relationship between their size and the size of their prey, meaning that as the snook grows, it feeds on larger and larger prey. Common snook occasionally engage in cannibalism . This usually occurs during the winter when adults and juveniles are in close proximity to one another within their estuarine habitats. This form of cannibalism where

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1308-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On

1417-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides

1526-529: A number of coastal water bodies such as coastal lagoons and brackish seas. A more comprehensive definition of an estuary is "a semi-enclosed body of water connected to the sea as far as the tidal limit or the salt intrusion limit and receiving freshwater runoff; however the freshwater inflow may not be perennial, the connection to the sea may be closed for part of the year and tidal influence may be negligible". This broad definition also includes fjords , lagoons , river mouths , and tidal creeks . An estuary

1635-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as

1744-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that

1853-634: A severe cold kill affected snook population number." All snook were "catch-and-release only" in the Gulf of Mexico until August 31, 2013. At that time, the FWC could choose to open or close snook harvest for another season. The commercial harvest or sale of snook is prohibited by the same regulations. At the June 2013 FWC meeting, commissioners voted to let the recreational harvest of snook reopen in Gulf of Mexico waters from September 1 that year. The next stock assessment for snook

1962-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,

2071-406: A significant impact on their behavior. For example, common snook are able to determine when to start and stop spawning based on the temperature and salinity of the water they inhabit, the amount of rainfall in the area, and whether or not the moon is full. In some cases, disturbances in their environment can have very negative effects on the snook population. One example is the devastating results of

2180-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of

2289-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in

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2398-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as

2507-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there

2616-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define

2725-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including

2834-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines

2943-408: A type of ecosystem in some estuaries that have been negatively impacted by eutrophication. Cordgrass vegetation dominates the salt marsh landscape. Excess nutrients allow the plants to grow at greater rates in above ground biomass, however less energy is allocated to the roots since nutrients is abundant. This leads to a lower biomass in the vegetation below ground which destabilizes the banks of

3052-469: A unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given

3161-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species

3270-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,

3379-550: A well-mixed water column and the disappearance of the vertical salinity gradient . The freshwater-seawater boundary is eliminated due to the intense turbulent mixing and eddy effects . The lower reaches of Delaware Bay and the Raritan River in New Jersey are examples of vertically homogeneous estuaries. Inverse estuaries occur in dry climates where evaporation greatly exceeds the inflow of freshwater. A salinity maximum zone

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3488-491: A while. Occurring in shallow coastal waters (up to 20 m (66 ft) in depth), estuaries and lagoons , the fish often enter fresh water . They are carnivorous , with a diet dominated by smaller fishes, and crustaceans such as shrimp , and occasionally crabs . Due to their limited cold tolerance, snook have been physiologically restricted from inhabiting temperate climates. In Florida, northern limits in their geographic range have historically been Tarpon Springs along

3597-436: A wholly marine embayment to any of the other estuary types. The most important variable characteristics of estuary water are the concentration of dissolved oxygen, salinity and sediment load. There is extreme spatial variability in salinity, with a range of near-zero at the tidal limit of tributary rivers to 3.4% at the estuary mouth. At any one point, the salinity will vary considerably over time and seasons, making it

3706-470: A wide effect on the surrounding water bodies.  In turn, this can decrease fishing industry sales in one area and across the country. Production in 2016 from recreational and commercial fishing contributes billions of dollars to the United States' gross domestic product (GDP). A decrease in production within this industry can affect any of the 1.7 million people the fishing industry employs yearly across

3815-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that

3924-413: Is a protandric hermaphrodite fish species. Its spawning season spans from April to October. The peak spawning occurs during July and August. Spawning typically occurs in near-shore waters with high salinities . Following the spawning period, the juveniles then migrate to the brackish waters of the nearby estuarine environments. When these juveniles mature, they return to the higher-salinity waters of

4033-455: Is a dynamic ecosystem having a connection to the open sea through which the sea water enters with the rhythm of the tides . The effects of tides on estuaries can show nonlinear effects on the movement of water which can have important impacts on the ecosystem and waterflow. The seawater entering the estuary is diluted by the fresh water flowing from rivers and streams. The pattern of dilution varies between different estuaries and depends on

4142-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if

4251-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This

4360-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation

4469-509: Is derived from the Latin word aestuarium meaning tidal inlet of the sea, which in itself is derived from the term aestus , meaning tide. There have been many definitions proposed to describe an estuary. The most widely accepted definition is: "a semi-enclosed coastal body of water, which has a free connection with the open sea, and within which seawater is measurably diluted with freshwater derived from land drainage". However, this definition excludes

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4578-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when

4687-411: Is formed, and both riverine and oceanic water flow close to the surface towards this zone. This water is pushed downward and spreads along the bottom in both the seaward and landward direction. Examples of an inverse estuary are Spencer Gulf , South Australia, Saloum River and Casamance River , Senegal. Estuary type varies dramatically depending on freshwater input, and is capable of changing from

4796-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as

4905-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,

5014-560: Is less restricted, and there is a slow but steady exchange of water between the estuary and the ocean. Fjord-type estuaries can be found along the coasts of Alaska , the Puget Sound region of western Washington state , British Columbia , eastern Canada, Greenland , Iceland , New Zealand, and Norway. These estuaries are formed by subsidence or land cut off from the ocean by land movement associated with faulting , volcanoes , and landslides . Inundation from eustatic sea-level rise during

5123-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept

5232-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with

5341-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,

5450-408: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Estuary Most existing estuaries formed during

5559-639: Is the whitefish species from the European Alps . Eutrophication reduced the oxygen levels in their habitats so greatly that whitefish eggs could not survive, causing local extinctions. However, some animals, such as carnivorous fish, tend to do well in nutrient-enriched environments and can benefit from eutrophication. This can be seen in populations of bass or pikes. Eutrophication can affect many marine habitats which can lead to economic consequences. The commercial fishing industry relies upon estuaries for approximately 68 percent of their catch by value because of

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5668-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,

5777-719: The Holocene Epoch has also contributed to the formation of these estuaries. There are only a small number of tectonically produced estuaries; one example is the San Francisco Bay , which was formed by the crustal movements of the San Andreas Fault system causing the inundation of the lower reaches of the Sacramento and San Joaquin rivers . In this type of estuary, river output greatly exceeds marine input and tidal effects have minor importance. Freshwater floats on top of

5886-468: The Holocene epoch with the flooding of river-eroded or glacially scoured valleys when the sea level began to rise about 10,000–12,000 years ago. Estuaries are typically classified according to their geomorphological features or to water-circulation patterns. They can have many different names, such as bays , harbors , lagoons , inlets , or sounds , although some of these water bodies do not strictly meet

5995-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as

6104-595: The Mandovi estuary in Goa during the monsoon period. As tidal forcing increases, river output becomes less than the marine input. Here, current induced turbulence causes mixing of the whole water column such that salinity varies more longitudinally rather than vertically, leading to a moderately stratified condition. Examples include the Chesapeake Bay and Narragansett Bay . Tidal mixing forces exceed river output, resulting in

6213-628: The Mid-Atlantic coast, and Galveston Bay and Tampa Bay along the Gulf Coast . Bar-built estuaries are found in a place where the deposition of sediment has kept pace with rising sea levels so that the estuaries are shallow and separated from the sea by sand spits or barrier islands. They are relatively common in tropical and subtropical locations. These estuaries are semi-isolated from ocean waters by barrier beaches ( barrier islands and barrier spits ). Formation of barrier beaches partially encloses

6322-702: The Severn Estuary in the United Kingdom and the Ems Dollard along the Dutch-German border. The width-to-depth ratio of these estuaries is typically large, appearing wedge-shaped (in cross-section) in the inner part and broadening and deepening seaward. Water depths rarely exceed 30 m (100 ft). Examples of this type of estuary in the U.S. are the Hudson River , Chesapeake Bay , and Delaware Bay along

6431-478: The black-tailed godwit , rely on estuaries. Two of the main challenges of estuarine life are the variability in salinity and sedimentation . Many species of fish and invertebrates have various methods to control or conform to the shifts in salt concentrations and are termed osmoconformers and osmoregulators . Many animals also burrow to avoid predation and to live in a more stable sedimental environment. However, large numbers of bacteria are found within

6540-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is

6649-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen

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6758-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept

6867-542: The Gulf Coast and Cape Canaveral on the east coast. Over the past 15 years, snook have extended their range as far north as the Suwannee River Estuary (~29°N), where an exponential increase in their population has occurred. Warmer winter temperatures resulting from climate change has been suggested as a likely cause. There may be potential food-web effects, particularly for sea trout ( Cynoscion species), related to

6976-649: The Ice Age, snook moved northward along the Mexico shoreline. They followed the perimeter of the Gulf of Mexico, along the west and east coasts of Florida. Massive snook are found in Central America, although they seem to look a little different because of the weather and water quality, but they are the same. No restrictions exist in most of Central America on the size or quantity of snook one can keep, consequently many locals have been keeping and killing these large snook for quite

7085-399: The June 2012 Florida Fish and Wildlife Conservation Commission (FWC) meeting, Commissioners voted to keep the recreational harvest of snook in Gulf of Mexico waters closed through Aug. 31, 2013. This closure will offer the species additional protection after a 2010 cold kill detrimentally affected the population. Snook closed to harvest in Gulf of Mexico and Atlantic waters in January 2010 after

7194-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for

7303-400: The United States. Estuaries are incredibly dynamic systems, where temperature, salinity, turbidity, depth and flow all change daily in response to the tides. This dynamism makes estuaries highly productive habitats, but also make it difficult for many species to survive year-round. As a result, estuaries large and small experience strong seasonal variation in their fish communities. In winter,

7412-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as

7521-513: The above definition of an estuary and could be fully saline. Many estuaries suffer degeneration from a variety of factors including soil erosion , deforestation , overgrazing , overfishing and the filling of wetlands. Eutrophication may lead to excessive nutrients from sewage and animal wastes; pollutants including heavy metals , polychlorinated biphenyls , radionuclides and hydrocarbons from sewage inputs; and diking or damming for flood control or water diversion. The word "estuary"

7630-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to

7739-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species

7848-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,

7957-452: The bottom where they are harmless. Historically the oysters filtered the estuary's entire water volume of excess nutrients every three or four days. Today that process takes almost a year, and sediment, nutrients, and algae can cause problems in local waters. Some major rivers that run through deserts historically had vast, expansive estuaries that have been reduced to a fraction of their former size, because of dams and diversions. One example

8066-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be

8175-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered

8284-409: The dissolved oxygen from the water; thus hypoxic or anoxic zones can develop. Nitrogen is often the lead cause of eutrophication in estuaries in temperate zones. During a eutrophication event, biogeochemical feedback decreases the amount of available silica . These feedbacks also increase the supply of nitrogen and phosphorus, creating conditions where harmful algal blooms can persist. Given

8393-401: The effects of modifying the estuarine circulation. Fjord -type estuaries are formed in deeply eroded valleys formed by glaciers . These U-shaped estuaries typically have steep sides, rock bottoms, and underwater sills contoured by glacial movement. The estuary is shallowest at its mouth, where terminal glacial moraines or rock bars form sills that restrict water flow. In the upper reaches of

8502-595: The estuary impacted by human activities, and over time may shift the basic composition of the ecosystem, and the reversible or irreversible changes in the abiotic and biotic parts of the systems from the bottom up. For example, Chinese and Russian industrial pollution, such as phenols and heavy metals, has devastated fish stocks in the Amur River and damaged its estuary soil. Estuaries tend to be naturally eutrophic because land runoff discharges nutrients into estuaries. With human activities, land run-off also now includes

8611-424: The estuary, the depth can exceed 300 m (1,000 ft). The width-to-depth ratio is generally small. In estuaries with very shallow sills, tidal oscillations only affect the water down to the depth of the sill, and the waters deeper than that may remain stagnant for a very long time, so there is only an occasional exchange of the deep water of the estuary with the ocean. If the sill depth is deep, water circulation

8720-771: The estuary, with only narrow inlets allowing contact with the ocean waters. Bar-built estuaries typically develop on gently sloping plains located along tectonically stable edges of continents and marginal sea coasts. They are extensive along the Atlantic and Gulf coasts of the U.S. in areas with active coastal deposition of sediments and where tidal ranges are less than 4 m (13 ft). The barrier beaches that enclose bar-built estuaries have been developed in several ways: Fjords were formed where Pleistocene glaciers deepened and widened existing river valleys so that they become U-shaped in cross-sections. At their mouths there are typically rocks, bars or sills of glacial deposits , which have

8829-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising

8938-609: The fish community is dominated by hardy marine residents, and in summer a variety of marine and anadromous fishes move into and out of estuaries, capitalizing on their high productivity. Estuaries provide a critical habitat to a variety of species that rely on estuaries for life-cycle completion. Pacific Herring ( Clupea pallasii ) are known to lay their eggs in estuaries and bays, surfperch give birth in estuaries, juvenile flatfish and rockfish migrate to estuaries to rear, and anadromous salmonids and lampreys use estuaries as migration corridors. Also, migratory bird populations, such as

9047-555: The fish. Snook are prized as game fish , being known for their great fighting capabilities. The IGFA all-tackle world record for common snook stands at 53 lb 10 oz (24.3 kg) caught by Gilbert Ponzi near Parismina Ranch, Costa Rica. Previous world records were caught in Fort Myers, Florida , and Gatun Spillway Canal Zone, Panama. Anglers regularly use a variety of lures to catch snook. Common lures include jerkbaits, bucktail jigs, topwater lures, spoons, and live bait. "At

9156-400: The great biodiversity of this ecosystem. During an algal bloom , fishermen have noticed a significant increase in the quantity of fish. A sudden increase in primary productivity causes spikes in fish populations which leads to more oxygen being utilized. It is the continued deoxygenation of the water that then causes a decline in fish populations. These effects can begin in estuaries and have

9265-420: The impacts do not end there. Plant death alters the entire food web structure which can result in the death of animals within the afflicted biome . Estuaries are hotspots for biodiversity , containing a majority of commercial fish catch, making the impacts of eutrophication that much greater within estuaries. Some specific estuarine animals feel the effects of eutrophication more strongly than others. One example

9374-401: The juveniles are fed on by the adults is referred to as intercohort cannibalism. The adult common snook that cannibalize juveniles most likely target them because the juveniles may be the largest of the available prey, so are nutritionally efficient to prey upon. Common snook, like many species of fish, are very in tune with their environments. Even a slight change in their surroundings can have

9483-525: The many chemicals used as fertilizers in agriculture as well as waste from livestock and humans. Excess oxygen-depleting chemicals in the water can lead to hypoxia and the creation of dead zones . This can result in reductions in water quality, fish, and other animal populations. Overfishing also occurs. Chesapeake Bay once had a flourishing oyster population that has been almost wiped out by overfishing. Oysters filter these pollutants, and either eat them or shape them into small packets that are deposited on

9592-422: The marine environment, such as plastics , pesticides , furans , dioxins , phenols and heavy metals . Such toxins can accumulate in the tissues of many species of aquatic life in a process called bioaccumulation . They also accumulate in benthic environments, such as estuaries and bay muds : a geological record of human activities of the last century. The elemental composition of biofilm reflect areas of

9701-575: The marsh causing increased rates of erosion . A similar phenomenon occurs in mangrove swamps , which are another potential ecosystem in estuaries. An increase in nitrogen causes an increase in shoot growth and a decrease in root growth. Weaker root systems cause a mangrove tree to be less resilient in seasons of drought, which can lead to the death of the mangrove. This shift in above ground and below ground biomass caused by eutrophication could hindered plant success in these ecosystems. Across all biomes, eutrophication often results in plant death but

9810-440: The now off-balance nitrogen cycle , estuaries can be driven to phosphorus limitation instead of nitrogen limitation. Estuaries can be severely impacted by an unbalanced phosphorus cycle, as phosphorus interacts with nitrogen and silica availability. With an abundance of nutrients in the ecosystem, plants and algae overgrow and eventually decompose, which produce a significant amount of carbon dioxide. While releasing CO 2 into

9919-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting

10028-542: The open ocean to join the breeding population. The common snook is an estuarine and freshwater-dependent fish species. Within estuaries, juvenile common snook are most often found inhabiting areas such as coastal wetland ponds, island networks, and creeks. Despite being a euryhaline species of fish, the common snook shows a tendency to gravitate towards lower-salinity conditions in the early stages of its life. By being able to adapt and thrive in both high- and low-salinity conditions through osmoregulation , common snook display

10137-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using

10246-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species

10355-626: The previous year and a 96–97% decrease in apparent survival estimates. C. undecimalis is widespread throughout the tropical waters of the western Atlantic Ocean , from the coast of the North Carolina to Brazil including the Gulf of Mexico and the Caribbean Sea . Snook likely originated in Central America, and changes in the Earth's climate brought the snook to Florida. During a great warming trend after

10464-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean

10573-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be

10682-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that

10791-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered

10900-427: The seawater in a layer that gradually thins as it moves seaward. The denser seawater moves landward along the bottom of the estuary, forming a wedge-shaped layer that is thinner as it approaches land. As a velocity difference develops between the two layers, shear forces generate internal waves at the interface, mixing the seawater upward with the freshwater. An examples of a salt wedge estuary is Mississippi River and

11009-411: The sediment which has a very high oxygen demand. This reduces the levels of oxygen within the sediment often resulting in partially anoxic conditions, which can be further exacerbated by limited water flow. Phytoplankton are key primary producers in estuaries. They move with the water bodies and can be flushed in and out with the tides . Their productivity is largely dependent upon the turbidity of

11118-551: The snook colonization. Three United States Navy submarines have been named for this species, USS Robalo (SS-273) and USS Snook (SS-279) in the Second World War and USS Snook (SSN-592) in the 1950s. Considered an excellent food fish, the common snook is fished commercially and foreign-caught fish are sold in the US. When cooking snook, the skin must be removed, because it imparts an unpleasant taste, described as soapy, to

11227-502: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike

11336-558: The species. One of the largest snooks , C. undecimalis grows to a maximum overall length of 140 cm (4.6 ft). The common length is 50 cm (1.6 ft). The IGFA world record is 24.32 kg (53 lb 10 oz) caught in Parismina Ranch, Costa Rica, by Rafael Montalvo. Of typical centropomid form, it possesses drab coloration except for a distinctive black lateral line . It can also have bright yellow pelvic and caudal fins , especially during spawn. The common snook

11445-456: The volume of freshwater, the tidal range, and the extent of evaporation of the water in the estuary. Drowned river valleys are also known as coastal plain estuaries. In places where the sea level is rising relative to the land, sea water progressively penetrates into river valleys and the topography of the estuary remains similar to that of a river valley. This is the most common type of estuary in temperate climates. Well-studied estuaries include

11554-453: The water and atmosphere, these organisms are also intaking all or nearly all of the available oxygen creating a hypoxic environment and unbalanced oxygen cycle . The excess carbon in the form of CO 2 can lead to low pH levels and ocean acidification , which is more harmful for vulnerable coastal regions like estuaries. Eutrophication has been seen to negatively impact many plant communities in estuarine ecosystems . Salt marshes are

11663-835: The water. The main phytoplankton present are diatoms and dinoflagellates which are abundant in the sediment. A primary source of food for many organisms on estuaries, including bacteria , is detritus from the settlement of the sedimentation. Of the thirty-two largest cities in the world in the early 1990s, twenty-two were located on estuaries. As ecosystems, estuaries are under threat from human activities such as pollution and overfishing . They are also threatened by sewage, coastal settlement, land clearance and much more. Estuaries are affected by events far upstream, and concentrate materials such as pollutants and sediments. Land run-off and industrial, agricultural, and domestic waste enter rivers and are discharged into estuaries. Contaminants can be introduced which do not disintegrate rapidly in

11772-537: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of

11881-441: Was scheduled for 2015, but had not yet occurred as of June 2016, effectively leaving the fish under a protected status. Species A species ( pl. : species) is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as

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