182-577: Rhinocarcinosoma is a genus of eurypterid , an extinct group of aquatic arthropods . Fossils of Rhinocarcinosoma have been discovered in deposits ranging of Late Silurian age in the United States , Canada and Vietnam . The genus contains three species, the American R. cicerops and R. vaningeni and the Vietnamese R. dosonensis . The generic name is derived from the related genus Carcinosoma , and
364-489: A Rhinocarcinosoma specimen, a carapace, from the McKenzie Formation just east of Lock Haven, Pennsylvania . Rhinocarcinosoma is classified as part of the family Carcinosomatidae , a family within the superfamily Carcinosomatoidea , alongside the genera Carcinosoma , Eocarcinosoma , Eusarcana and possibly Holmipterus . The first cladogram below is adapted from a larger cladogram (simplified to only display
546-644: A chateau on Castle Island , within present-day Albany; it was abandoned the following year due to flooding. In 1614, the Dutch, under the command of Hendrick Corstiaensen, rebuilt the French chateau, which they called Fort Nassau . Fort Nassau was the first Dutch settlement in North America, and was located along the Hudson River, also within present-day Albany. The small fort served as a trading post and warehouse. Located on
728-564: A cosmopolitan distribution . Though the eurypterids continued to be abundant and diversify during the Early Devonian (for instance leading to the evolution of the pterygotid Jaekelopterus , the largest of all arthropods), the group was one of many heavily affected by the Late Devonian extinction . The extinction event, only known to affect marine life (particularly trilobites, brachiopods and reef -building organisms) effectively crippled
910-509: A cuticle composed of proteins and chitin . As in other chelicerates , the body was divided into two tagmata (sections); the frontal prosoma (head) and posterior opisthosoma (abdomen). The prosoma was covered by a carapace (sometimes called the "prosomal shield") on which both compound eyes and the ocelli (simple eye-like sensory organs) were located. The prosoma also bore six pairs of appendages which are usually referred to as appendage pairs I to VI. The first pair of appendages,
1092-608: A humid subtropical climate . Weather in New York is heavily influenced by two continental air masses: a warm, humid one from the southwest and a cold, dry one from the northwest. Downstate New York (comprising New York City, Long Island, and lower portions of the Hudson Valley) have rather hot summers with some periods of high humidity and cold, damp winters which are relatively mild compared to temperatures in Upstate New York, due to
1274-408: A lung , plastron or a pseudotrachea . Plastrons are organs that some arthropods evolved secondarily to breathe air underwater. This is considered an unlikely explanation since eurypterids had evolved in water from the start and they would not have organs evolved from air-breathing organs present. In addition, plastrons are generally exposed on outer parts of the body while the eurypterid gill tract
1456-497: A varied geography . The southeastern part of the state, known as Downstate , encompasses New York City , the United States's largest city; Long Island , the nation's most populous island; and the suburbs and wealthy enclaves of the lower Hudson Valley . These areas are the center of the expansive New York metropolitan area and account for approximately two-thirds of the state's population. The larger Upstate area spreads from
1638-925: A bottom-dwelling genus comes in the form of its swimming paddles being reduced in size compared to those of its relatives, such as Carcinosoma . It is likely that Rhinocarcinosoma was either a top predator, actively digging for prey or burrowing and lying in wait, or a scavenger, digging for scraps, in its environment. Given its size, it is possible that Rhinocarcinosoma fed on worms , other arthropods, lingulids and small fish. Rhinocarcinosoma would most likely have fed through using its spined forward-facing appendages to push food into its mouth. The deposits at Otisville where R. cicerops have been discovered have also yielded other eurypterids, including Hughmilleria shawangunk , Nanahughmilleria clarkei , Ruedemannipterus stylonuroides , Erettopterus globiceps , Hardieopterus myops , Kiaeropterus otisius and Ctenopterus cestrotus . Also present were early jawed fish of
1820-519: A career most natives had turned to in hopes of trading with whites first. This completely changed the ethnography of the region, and most large game was hunted out before whites ever fully explored the land. Still, afterward, the Iroquois Confederacy offered shelter to refugees of the Mascouten , Erie, Chonnonton , Tutelo , Saponi , and Tuscarora nations. The Tuscarora became the sixth nation of
2002-454: A dual respiratory system was present, which would have allowed for short periods of time in terrestrial environments. The name Eurypterida comes from the Ancient Greek words εὐρύς ( eurús ), meaning 'broad' or 'wide', and πτερόν ( pterón ), meaning 'wing', referring to the pair of wide swimming appendages present in many members of the group. The eurypterid order includes
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#17331044951762184-726: A gait like that of most modern insects. The weight of its long abdomen would have been balanced by two heavy and specialized frontal appendages, and the center of gravity might have been adjustable by raising and positioning the tail. Preserved fossilized eurypterid trackways tend to be large and heteropodous and often have an associated telson drag mark along the mid-line (as with the Scottish Hibbertopterus track). Such trackways have been discovered on every continent except for South America. In some places where eurypterid fossil remains are otherwise rare, such as in South Africa and
2366-682: A major threat to the Abenaki and Mohicans, while the Susquehannock briefly conquered the Lenape in the 1600s. The most devastating event of the century, however, was the Beaver Wars . From approximately 1640–1680, the Iroquois peoples waged campaigns which extended from modern-day Michigan to Virginia against Algonquian and Siouan tribes, as well as each other. The aim was to control more land for animal trapping ,
2548-416: A manner similar to modern horseshoe crabs, by grabbing and shredding food with their appendages before pushing it into their mouth using their chelicerae. Fossils preserving digestive tracts have been reported from fossils of various eurypterids, among them Carcinosoma , Acutiramus and Eurypterus . Though a potential anal opening has been reported from the telson of a specimen of Buffalopterus , it
2730-425: A meter (1.64 ft) even if the extended chelicerae are not included. Two other eurypterids have also been estimated to have reached lengths of 2.5 metres; Erettopterus grandis (closely related to Jaekelopterus ) and Hibbertopterus wittebergensis , but E. grandis is very fragmentary and the H. wittenbergensis size estimate is based on trackway evidence, not fossil remains. The family of Jaekelopterus ,
2912-471: A rowing type of propulsion similar to that of crabs and water beetles . Larger individuals may have been capable of underwater flying (or subaqueous flight ) in which the motion and shape of the paddles are enough to generate lift , similar to the swimming of sea turtles and sea lions . This type of movement has a relatively slower acceleration rate than the rowing type, especially since adults have proportionally smaller paddles than juveniles. However, since
3094-583: A small group known as the Ouabano migrated southwest into West Virginia at an earlier time. They may have merged with the Shawnee . The Mohawk and Susquehannock were the most militaristic . Trying to corner trade with the Europeans, they targeted other tribes. The Mohawk were also known for refusing white settlement on their land and discriminating against any of their people who converted to Christianity . They posed
3276-641: A temperature of −13 °F (−25 °C) or lower can be expected in the northern highlands (Northern Plateau) and 5 °F (−15 °C) or colder in the southwestern and east-central highlands of the Southern Tier. New York had a record-high temperature of 108 °F (42.2 °C) on July 22, 1926, in the Albany area. Its record-lowest temperature during the winter was −52 °F (−46.7 °C) in 1979. Governors Island, Manhattan , in New York Harbor ,
3458-644: A total area of 54,555 square miles (141,297 km ) and ranks as the 27th-largest state by size . The highest elevation in New York is Mount Marcy in the Adirondack High Peaks in Northern New York , at 5,344 feet (1,629 meters) above sea level ; while the state's lowest point is at sea level, on the Atlantic Ocean in Downstate New York . In contrast with New York City's urban landscape,
3640-592: A water border with Rhode Island , while Connecticut has land and sea borders. Except for areas near the New York Harbor and the Upper Delaware River , New York has a mostly land border with two Mid-Atlantic states , New Jersey and Pennsylvania . New York is the only state that borders both the Great Lakes and the Atlantic Ocean . The Hudson River begins near Lake Tear of the Clouds and flows south through
3822-547: Is a genital appendage. This appendage, an elongated rod with an internal duct, is found in two distinct morphs, generally referred to as "type A" and "type B". These genital appendages are often preserved prominently in fossils and have been the subject of various interpretations of eurypterid reproduction and sexual dimorphism. Type A appendages are generally longer than those of type B. In some genera they are divided into different numbers of sections, such as in Eurypterus where
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#17331044951764004-495: Is a lightweight build. Factors such as locomotion, energy costs in molting and respiration, as well as the actual physical properties of the exoskeleton , limit the size that arthropods can reach. A lightweight construction significantly decreases the influence of these factors. Pterygotids were particularly lightweight, with most fossilized large body segments preserving as thin and unmineralized. Lightweight adaptations are present in other giant paleozoic arthropods as well, such as
4186-553: Is a very rare genus of eurypterids, specimens have also been found elsewhere, though they have not been assigned to any particular species. In 1985, Brian Jones and Erik N. Kjellesvig-Waering referred a poorly preserved part of a prosoma recovered in the Leopold Formation on Somerset Island in Canada to Rhinocarcinosoma sp. indet., an assignment which has been maintained as correct in later research. In 1992, Samuel J. Ciurca reported
4368-523: Is also possible and the structure may represent the unfused tips of the appendages. Located between the dorsal and ventral surfaces of the Blattfüsse associated with the type A appendages is a set of organs traditionally described as either "tubular organs" or "horn organs". These organs are most often interpreted as spermathecae (organs for storing sperm ), though this function is yet to be proven conclusively. In arthropods, spermathecae are used to store
4550-516: Is covered by ice. The summer climate is cool in the Adirondacks, Catskills, and at higher elevations of the Southern Tier. Buffalo and its metropolitan area are described as climate change havens for their weather pattern in Western New York. Summer daytime temperatures range from the high 70s to low 80s °F (25 to 28 °C), over most of the state. In the majority of winter seasons,
4732-483: Is located behind the Blattfüssen . Instead, among arthropod respiratory organs, the eurypterid gill tracts most closely resemble the pseudotracheae found in modern isopods . These organs, called pseudotracheae, because of some resemblance to the tracheae (windpipes) of air-breathing organisms, are lung-like and present within the pleopods (back legs) of isopods. The structure of the pseudotracheae has been compared to
4914-487: Is made up of the first six exoskeleton segments fused together into a larger structure. The seventh segment (thus the first opisthosomal segment) is referred to as the metastoma and the eighth segment (distinctly plate-like) is called the operculum and contains the genital aperature. The underside of this segment is occupied by the genital operculum, a structure originally evolved from ancestral seventh and eighth pair of appendages. In its center, as in modern horseshoe crabs,
5096-517: Is more likely that the anus was opened through the thin cuticle between the last segment before the telson and the telson itself, as in modern horseshoe crabs. Eurypterid coprolites discovered in deposits of Ordovician age in Ohio containing fragments of a trilobite and eurypterid Megalograptus ohioensis in association with full specimens of the same eurypterid species have been suggested to represent evidence of cannibalism . Similar coprolites referred to
5278-537: Is much more of a marine influence in many of the sections yielding Adelophthalmus than has previously been acknowledged." Similarly, a study of the eurypterid Hibbertopterus from the Carboniferous of New Mexico concluded that the habitat of some eurypterids "may need to be re-evaluated". The sole surviving eurypterine family, Adelophthalmidae, was represented by only a single genus, Adelophthalmus . The hibbertopterids, mycteroptids and Adelophthalmus survived into
5460-419: Is only confidently known from R. vaningeni , given that the relevant portion of the carapace has not been preserved in adult fossils of R. dosonensis and R. cicerops (which lacks known adult fossils altogether). All species of Rhinocarcinosoma also share certain other features of the carapace, including the carapace being more or less subtriangular in shape, with a width to length ratio of about 3:2, and
5642-528: Is planned to host a US$ 1 billion research and education center poised to make New York the global leader in addressing the climate crisis . Due to New York's relatively large land area and unique geography compared to other eastern states, there are several distinct ecoregions present in the state, many of them reduced heavily due to urbanization and other human activities: Southern Great Lakes forests in Western New York , New England–Acadian forests on
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5824-625: Is possible that many eurypterid species thought to be distinct actually represent juvenile specimens of other species, with paleontologists rarely considering the influence of ontogeny when describing new species. Studies on a well-preserved fossil assemblage of eurypterids from the Pragian -aged Beartooth Butte Formation in Cottonwood Canyon , Wyoming , composed of multiple specimens of various developmental stages of eurypterids Jaekelopterus and Strobilopterus , revealed that eurypterid ontogeny
6006-620: Is possible that the adults were even more similar to R. vaningeni . Although formal synonymisation has never been conducted, it is possible that R. cicerops were actually juvenile R. vaningeni . Rhinocarcinosoma fossils were first found in the Illion Shale of New York State in the United States , and they were first described and discussed in detail in John Mason Clarke 's and Rudolf Ruedemann 's 1912 The Eurypterida of New York . The earliest described species of Rhinocarcinosoma
6188-637: Is shared between New York and Ontario as it flows on the Niagara River from Lake Erie to Lake Ontario. The Delaware River Basin Compact, signed in 1961 by New York, New Jersey, Pennsylvania , Delaware , and the federal government, regulates the utilization of water of the Delaware system. Under the Köppen climate classification , most of New York has a humid continental climate , though New York City and Long Island have
6370-439: Is the first record of land locomotion by a eurypterid. The trackway provides evidence that some eurypterids could survive in terrestrial environments, at least for short periods of time, and reveals information about the stylonurine gait. In Hibbertopterus , as in most eurypterids, the pairs of appendages are different in size (referred to as a heteropodous limb condition). These differently sized pairs would have moved in phase, and
6552-795: Is the metastoma becoming proportionally less wide. This ontogenetic change has been observed in members of several superfamilies, such as the Eurypteroidea, the Pterygotioidea and the Moselopteroidea . No fossil gut contents from eurypterids are known, so direct evidence of their diet is lacking. The eurypterid biology is particularly suggestive of a carnivorous lifestyle. Not only were many large (in general, most predators tend to be larger than their prey), but they had stereoscopic vision (the ability to perceive depth). The legs of many eurypterids were covered in thin spines, used both for locomotion and
6734-514: Is true for New York City: four of its five boroughs are situated on three islands at the mouth of the Hudson River : Manhattan Island ; Staten Island ; and Long Island , which contains Brooklyn and Queens at its western end. The state's borders include a water boundary in (clockwise from the west) two Great Lakes ( Lake Erie and Lake Ontario , which are connected by the Niagara River );
6916-418: Is unlikely the "gill tract" contained functional gills when comparing the organ to gills in other invertebrates and even fish. Previous interpretations often identified the eurypterid "gills" as homologous with those of other groups (hence the terminology), with gas exchange occurring within the spongy tract and a pattern of branchio-cardiac and dendritic veins (as in related groups) carrying oxygenated blood into
7098-519: The American Civil War , in terms of finances, volunteer soldiers, and supplies. The state provided more than 370,000 soldiers to the Union armies. Over 53,000 New Yorkers died in service, roughly one of every seven who served. However, Irish draft riots in 1862 were a significant embarrassment. Since the early 19th century, New York City has been the largest port of entry for legal immigration into
7280-637: The Capital Region entered Phase 1 of reopening. On May 26, the Hudson Valley began Phase 1, and New York City partially reopened on June 8. During July 2020, a federal judge ruled Governor Andrew Cuomo and Mayor Bill de Blasio exceeded authority by limiting religious gatherings to 25% when others operated at 50% capacity. On Thanksgiving Eve, the U.S. Supreme Court blocked additional religious restrictions imposed by Cuomo for areas with high infection rates. The state of New York covers
7462-597: The Continental Army at the Battle of Saratoga in 1777, a success that influenced France to ally with the revolutionaries; the state constitution was enacted in 1777. New York became the 11th state to ratify the United States Constitution , on July 26, 1788. In an attempt to retain their sovereignty and remain an independent nation positioned between the new United States and British North America , four of
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7644-459: The Declaration of Independence on July 9, 1776. The New York State Constitution was framed by a convention which assembled at White Plains on July 10, 1776, and after repeated adjournments and changes of location, finished its work at Kingston on Sunday evening, April 20, 1777, when the new constitution drafted by John Jay was adopted with but one dissenting vote. It was not submitted to
7826-661: The Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian , from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though
8008-581: The Great Lakes to Lake Champlain and includes the Adirondack Mountains and the Catskill Mountains (part of the wider Appalachian Mountains ). The east–west Mohawk River Valley bisects the more mountainous regions of Upstate and flows into the north–south Hudson River valley near the state capital of Albany . Western New York , home to the cities of Buffalo and Rochester , is part of
8190-681: The Great Lakes , with the port of New York City. Improving transportation, it enabled additional population migration to territories west of New York. After 1850, railroads largely replaced the canal. The connectivity offered by the canal, and subsequently the railroads, led to an economic boom across the entire state through the 1950s. Major corporations that got their start in New York during this time include American Express , AT&T , Bristol Myers Squibb , Carrier , Chase , General Electric , Goldman Sachs , IBM , Kodak , Macy's , NBC , Pfizer , Random House , RCA , Tiffany & Co. , Wells Fargo , Western Union , and Xerox . New York City
8372-609: The Great Lakes region and borders Lake Ontario and Lake Erie . Central New York is anchored by the city of Syracuse ; between the central and western parts of the state, New York is dominated by the Finger Lakes , a popular tourist destination. To the south, along the state border with Pennsylvania , the Southern Tier sits atop the Allegheny Plateau , representing some of the northernmost reaches of Appalachia . New York
8554-741: The Iroquois Nations fought on the side of the British; only the Oneida and their dependents, the Tuscarora, allied themselves with the Americans. In retaliation for attacks on the frontier led by Joseph Brant and Loyalist Mohawk forces , the Sullivan Expedition of 1779 destroyed nearly 50 Iroquois villages, adjacent croplands and winter stores, forcing many refugees to British-held Niagara. As allies of
8736-719: The Lake Champlain Valley . The Great Appalachian Valley dominates eastern New York and contains Lake Champlain Valley as its northern half and the Hudson Valley as its southern half within the state. The Tug Hill region arises as a cuesta east of Lake Ontario . The state of New York contains a part of the Marcellus shale , which extends into Ohio and Pennsylvania. Upstate and Downstate are often used informally to distinguish New York City or its greater metropolitan area from
8918-528: The New England border, Northeastern coastal forests in the lower Hudson Valley and western Long Island , Atlantic coastal pine barrens in southern Long Island, Northeastern interior dry–mesic oak forest in the eastern Southern Tier and upper Hudson Valley, Appalachian–Blue Ridge forests in the Hudson Valley), Central Appalachian dry oak–pine forest around the Hudson Valley, Eastern Great Lakes and Hudson Lowlands , Eastern forest–boreal transition in
9100-639: The New York State Legislature created the Board of Commissioners of Immigration to regulate immigration. The first permanent immigration depot in New York was established in 1855 at Castle Garden , a converted War of 1812 era fort located within what is now Battery Park , at the tip of Lower Manhattan. The first immigrants to arrive at the new depot were aboard three ships that had just been released from quarantine . Castle Garden served as New York's immigrant depot until it closed on April 18, 1890, when
9282-613: The Philadelphia Convention , which drafted the United States Constitution, in which he also took part. The new government was to be a strong federal national government to replace the relatively weaker confederation of individual states. Following heated debate, which included the publication of The Federalist Papers as a series of installments in New York City newspapers, New York was the 11th state to ratify
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#17331044951769464-521: The Stylonuroidea , Kokomopteroidea and Mycteropoidea as well as eurypterine groups such as the Pterygotioidea, Eurypteroidea and Waeringopteroidea . The most successful eurypterid by far was the Middle to Late Silurian Eurypterus , a generalist , equally likely to have engaged in predation or scavenging . Thought to have hunted mainly small and soft-bodied invertebrates, such as worms , species of
9646-778: The Susquehannock and the Erie . Many of the Wampanoag and Mohican peoples were caught up in King Philip's War , a joint effort of many New England tribes to push Europeans off their land. After the death of their leader, Chief Philip Metacomet , most of those peoples fled inland, splitting into the Abenaki and the Schaghticoke . Many of the Mohicans remained in the region until the 1800s, however,
9828-476: The United States Constitution , on July 26, 1788. New York City remained the national capital under the new constitution until 1790 when it was moved to Philadelphia until 1800, when it was relocated to its current location in Washington, D.C. and was the site of the inauguration of President George Washington, In the first session of the Supreme Court of the United States , the United States Bill of Rights were drafted. Transportation in Western New York
10010-471: The World Trade Center site . On October 29 and 30, 2012, Hurricane Sandy caused extensive destruction of the state's shorelines, ravaging portions of New York City, Long Island , and southern Westchester with record-high storm surge , with severe flooding and high winds causing power outages for hundreds of thousands of New Yorkers, and leading to gasoline shortages and disruption of mass transit systems. The storm and its profound effects have prompted
10192-477: The coxae (limb segments) used for feeding. These appendages were generally walking legs that were cylindrical in shape and were covered in spines in some species. In most lineages, the limbs tended to get larger the farther back they were. In the Eurypterina suborder , the larger of the two eurypterid suborders, the sixth pair of appendages was also modified into a swimming paddle to aid in traversing aquatic environments. The opisthosoma comprised 12 segments and
10374-442: The rhizodonts , were the new apex predators in marine environments. However, various recent findings raise doubts about this, and suggest that these eurypterids were euryhaline forms that lived in marginal marine environments, such as estuaries, deltas, lagoons, and coastal ponds. One argument is paleobiogeographical; pterygotoid distribution seems to require oceanic dispersal. A recent review of Adelophthalmoidea admitted that "There
10556-517: The ring-necked pheasant , northern bobwhite , ruffed grouse , spruce grouse , Canada jay , wild turkey , blue jay , eastern bluebird (the state bird ), American robin , and black-capped chickadee . Birds of prey that are present in the state are great horned owls , bald eagles , red-tailed hawks , American kestrels , and northern harriers . Waterfowl like mallards , wood ducks , canvasbacks , American black ducks , trumpeter swans , Canada geese , and blue-winged teals can be found in
10738-405: The spermatophore received from males. This would imply that the type A appendage is the female morph and the type B appendage is the male. Further evidence for the type A appendages representing the female morph of genital appendages comes in their more complex construction (a general trend for female arthropod genitalia). It is possible that the greater length of the type A appendage means that it
10920-487: The telson , the posteriormost division of the body, which in most species took the form of a blade-like shape. In some lineages, notably the Pterygotioidea , the Hibbertopteridae and the Mycteroptidae , the telson was flattened and may have been used as a rudder while swimming. Some genera within the superfamily Carcinosomatoidea , notably Eusarcana , had a telson similar to that of modern scorpions and may have been capable of using it to inject venom . The coxae of
11102-539: The white-footed mouse , North American least shrew , little brown bat , muskrat , eastern gray squirrel , eastern cottontail , American ermine , groundhog , striped skunk , fisher , North American river otter , raccoon , bobcat , eastern coyote , red fox , gray fox white-tailed deer , moose , and American black bear ; extirpated mammals include Canada lynx , American bison , wolverine , Allegheny woodrat , caribou , eastern elk , eastern cougar , and eastern wolf . Some species of birds in New York are
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#173310449517611284-548: The Adirondacks, Eastern Great Lakes lowland forests around the Adirondacks, and Allegheny Highlands forests , most of which are in the western Southern Tier. Some species that can be found in this state are American ginseng , starry stonewort , waterthyme , water chestnut, eastern poison ivy , poison sumac , giant hogweed , cow parsnip and common nettle . There are more than 70 mammal species, more than 20 bird species, some species of amphibians, and several reptile species. Species of mammals that are found in New York are
11466-438: The British, the Iroquois were forced out of New York, although they had not been part of treaty negotiations. They resettled in Canada after the war and were given land grants by the Crown. In the treaty settlement, the British ceded most Indian lands to the new United States. Because New York made a treaty with the Iroquois without getting Congressional approval, some of the land purchases have been subject to land claim suits since
11648-832: The Carcinosomatoidea) in a 2007 study by eurypterid researcher O. Erik Tetlie, which was in turn based on results from various phylogenetic analyses on eurypterids conducted between 2004 and 2007. The second cladogram below is simplified from a study by Lamsdell et al. (2015). Tetlie (2007) Carcinosoma newlini Eusarcana scorpionis Rhinocarcinosoma vaningeni Carcinosoma scoticus Carcinosoma scorpiodes Mixopterus multispinosus Mixopterus simonsoni Mixopterus kiaeri Lamsdell et al. (2015) Megalograptidae Mixopteridae Holmipterus suecicus Rhinocarcinosoma vaningeni Carcinosoma newlini Carcinosoma libertyi Eusarcana acrocephalus Eusarcana scorpionis Carcinosomatid eurypterids such as Rhinocarcinosoma were among
11830-420: The Devonian, large two meter (6.5+ ft) pterygotids such as Acutiramus were already present during the Late Silurian. Their ecology ranged from generalized predatory behavior to ambush predation and some, such as Pterygotus itself, were active apex predators in Late Silurian marine ecosystems. The pterygotids were also evidently capable of crossing oceans, becoming one of only two eurypterid groups to achieve
12012-544: The Greek ῥινός (rhinós, "nose"), referring to the unusual shovel-shaped protrusion on the front of the carapace (head plate) of Rhinocarcinosoma , its most distinctive feature. Other than the protrusion, Rhinocarcinosoma was anatomically very similar to its close relative, Eusarcana , though it lacked the scorpion-like telson (the posteriormost division of the body) of that genus. Further distinguishing features include more slender appendages and slightly different ornamentation of scales. In terms of size, Rhinocarcinosoma
12194-466: The Hudson River flood plain, the rudimentary fort was washed away by flooding in 1617, and abandoned for good after Fort Orange (New Netherland) was built nearby in 1623. Henry Hudson's 1609 voyage marked the beginning of European involvement in the area. Sailing for the Dutch East India Company and looking for a passage to Asia, he entered the Upper New York Bay on September 11 of that year. Word of his findings encouraged Dutch merchants to explore
12376-482: The Iroquois Confederacy include the Seneca, Cayuga, Onondaga, Oneida, Tuscarora and Mohawk. The Iroquois fought for both sides during the Revolutionary War ; afterwards many pro-British Iroquois migrated to Canada. Today, the Iroquois still live in several enclaves across New York and Ontario. Meanwhile, the Lenape formed a close relationship with William Penn . However, upon Penn's death, his sons managed to take over much of their lands and banish them to Ohio. When
12558-414: The Iroquois. In the 1700s, Iroquoian peoples would take in the remaining Susquehannock of Pennsylvania after they were decimated in the French and Indian War . Most of these other groups assimilated and eventually ceased to exist as separate tribes. Then, after the American Revolution , a large group of Seneca split off and returned to Ohio, becoming known as the Mingo Seneca. The current Six Nations of
12740-506: The King of France's sister . Verrazzano described it as "a vast coastline with a deep delta in which every kind of ship could pass" and he adds: "that it extends inland for a league and opens up to form a beautiful lake. This vast sheet of water swarmed with native boats." He landed on the tip of Manhattan and possibly on the furthest point of Long Island. Verrazzano's stay was interrupted by a storm which pushed him north towards Martha's Vineyard . In 1540, French traders from New France built
12922-423: The Lenape. In 1524, Giovanni da Verrazzano , an Italian explorer in the service of the French crown , explored the Atlantic coast of North America between the Carolinas and Newfoundland , including New York Harbor and Narragansett Bay . On April 17, 1524, Verrazzano entered New York Bay , by way of the strait now called the Narrows into the northern bay which he named Santa Margherita, in honor of
13104-804: The Middle Ordovician suggests that eurypterids either originated during the Early Ordovician and experienced a rapid and explosive radiation and diversification soon after the first forms evolved, or that the group originated much earlier, perhaps during the Cambrian period. As such, the exact eurypterid time of origin remains unknown. Though fossils referred to as "primitive eurypterids" have occasionally been described from deposits of Cambrian or even Precambrian age, they are not recognized as eurypterids, and sometimes not even as related forms, today. Some animals previously seen as primitive eurypterids, such as
13286-590: The Middle Ordovician, 467.3 million years ago . There are also reports of even earlier fossil eurypterids in the Fezouata Biota of Late Tremadocian (Early Ordovician) age in Morocco , but these have yet to be thoroughly studied, and are likely to be peytoiid appendages. Pentecopterus was a relatively derived eurypterid, part of the megalograptid family within the carcinosomatoid superfamily. Its derived position suggests that most eurypterid clades, at least within
13468-697: The Middle Silurian and the Early Devonian, with an absolute peak in diversity during the Pridoli epoch , 423 to 419.2 million years ago, of the very latest Silurian. This peak in diversity has been recognized since the early twentieth century; of the approximately 150 species of eurypterids known in 1916, more than half were from the Silurian and a third were from the Late Silurian alone. Though stylonurine eurypterids generally remained rare and low in number, as had been
13650-517: The Ngọc Xuyên hill. Though no new species was named at the time, the eurypterid fossils could quickly be identified as belonging to Rhinocarcinosoma . The new species R. dosonensis , named after the Dô Son Peninsula, was described in 2002 based on the Vietnamese fossils. Although the Vietnamese fossils did not preserve the diagnostic feature of Rhinocarcinosoma , the shovel-shaped protrusion at
13832-633: The Permian. New York (state) New York , also called New York State , is a state in the Northeastern United States . One of the Mid-Atlantic states, it borders the Atlantic Ocean , New England , Canada, and the Great Lakes . With almost 19.6 million residents, it is the fourth-most populous state in the United States , and the 27th-largest state by area , with a total area of 54,556 square miles (141,300 km ). New York has
14014-549: The Pterygotidae, is noted for several unusually large species. Both Acutiramus , whose largest member A. bohemicus measured 2.1 meters (6.9 ft), and Pterygotus , whose largest species P. grandidentatus measured 1.75 meters (5.7 ft), were gigantic. Several different contributing factors to the large size of the pterygotids have been suggested, including courtship behaviour, predation and competition over environmental resources. Giant eurypterids were not limited to
14196-586: The Stylonurina, this appendage takes the form of a long and slender walking leg, while in the Eurypterina, the leg is modified and broadened into a swimming paddle. Other than the swimming paddle, the legs of many eurypterines were far too small to do much more than allow them to crawl across the sea floor . In contrast, a number of stylonurines had elongated and powerful legs that might have allowed them to walk on land (similar to modern crabs ). A fossil trackway
14378-573: The Twin Towers of the original World Trade Center in Lower Manhattan , and the towers collapsed . 7 World Trade Center also collapsed due to damage from fires. The other buildings of the World Trade Center complex were damaged beyond repair and demolished soon thereafter. The collapse of the Twin Towers caused extensive damage and resulted in the deaths of 2,753 victims, including 147 aboard
14560-603: The U.S. drafted the Indian Removal Act , the Lenape were further moved to Missouri, whereas their cousins, the Mohicans, were sent to Wisconsin. Also, in 1778, the United States relocated the Nanticoke from the Delmarva Peninsula to the former Iroquois lands south of Lake Ontario, though they did not stay long. Mostly, they chose to migrate into Canada and merge with the Iroquois, although some moved west and merged with
14742-467: The United States . In the United States, the federal government did not assume direct jurisdiction for immigration until 1890. Prior to this time, the matter was delegated to the individual states, then via contract between the states and the federal government. Most immigrants to New York would disembark at the bustling docks along the Hudson and East Rivers , in the eventual Lower Manhattan . On May 4, 1847,
14924-456: The United States can trace their ancestry to these immigrants. Ellis Island was the subject of a contentious and long-running border and jurisdictional dispute between the State of New York and the State of New Jersey, as both claimed it. The issue was officially settled in 1998 by the U.S. Supreme Court which ruled that the original 3.3-acre (1.3 ha) island was New York state territory and that
15106-485: The United States. The state built its political, cultural, and economic ascendancy over the next century, earning it the nickname of the " Empire State ". Although deindustrialization eroded a portion of the state's economy in the second half of the 20th century, New York in the 21st century continues to be considered as a global node of creativity and entrepreneurship , social tolerance , and environmental sustainability . The state attracts visitors from all over
15288-538: The abundance and diversity previously seen within the eurypterids. A major decline in diversity had already begun during the Early Devonian and eurypterids were rare in marine environments by the Late Devonian. During the Frasnian stage four families went extinct, and the later Famennian saw an additional five families going extinct. As marine groups were the most affected, the eurypterids were primarily impacted within
15470-530: The ancient continent of Laurentia , and demersal (living on the seafloor ) and basal animals from the continents Avalonia and Gondwana. The Laurentian predators, classified in the family Megalograptidae (comprising the genera Echinognathus , Megalograptus and Pentecopterus ), are likely to represent the first truly successful eurypterid group, experiencing a small radiation during the Late Ordovician. Eurypterids were most diverse and abundant between
15652-442: The animal in question could possibly have measured just short of 2 meters (6.6 ft) in length. More robust than the pterygotids, this giant Hibbertopterus would possibly have rivalled the largest pterygotids in weight, if not surpassed them, and as such be among the heaviest arthropods. The two eurypterid suborders, Eurypterina and Stylonurina , are distinguished primarily by the morphology of their final pair of appendages. In
15834-454: The appendage via tracts, but these supposed tracts remain unpreserved in available fossil material. Type B appendages, assumed male, would have produced, stored and perhaps shaped spermatophore in a heart-shaped structure on the dorsal surface of the appendage. A broad genital opening would have allowed large amounts of spermatophore to be released at once. The long furca associated with type B appendages, perhaps capable of being lowered like
16016-599: The balance of the 27.5 acres (11 ha) added after 1834 by landfill was in New Jersey. In May 1964, Ellis Island was added to the National Park Service by President Lyndon B. Johnson and is still owned by the federal government as part of the Statue of Liberty National Monument . In 1990, Ellis Island was opened to the public as a museum of immigration. On September 11, 2001, two of four hijacked planes were flown into
16198-414: The body. The primary analogy used in previous studies has been horseshoe crabs, though their gill structure and that of eurypterids are remarkably different. In horseshoe crabs, the gills are more complex and composed of many lamellae (plates) which give a larger surface area used for gas exchange. In addition, the gill tract of eurypterids is proportionally much too small to support them if it is analogous to
16380-502: The canal carrying passengers and freight. Farm products came in from the Midwest , and finished manufactured goods moved west. It was an engineering marvel which opened up vast areas of New York to commerce and settlement. It enabled Great Lakes port cities such as Buffalo and Rochester to grow and prosper. It also connected the burgeoning agricultural production of the Midwest and shipping on
16562-458: The case during the preceding Ordovician, eurypterine eurypterids experienced a rapid rise in diversity and number. In most Silurian fossil beds, eurypterine eurypterids account for 90% of all eurypterids present. Though some were likely already present by the Late Ordovician (simply missing from the fossil record so far), a vast majority of eurypterid groups are first recorded in strata of Silurian age. These include both stylonurine groups such as
16744-621: The city on October 19 of that year, composed of representatives from across the Thirteen Colonies who set the stage for the Continental Congress to follow. The Stamp Act Congress resulted in the Declaration of Rights and Grievances , which was the first written expression by representatives of the Americans of many of the rights and complaints later expressed in the United States Declaration of Independence . This included
16926-578: The coast in search of profitable fur trading with local Native American tribes. During the 17th century, Dutch trading posts established for the trade of pelts from the Lenape, Iroquois, and other tribes were founded in the colony of New Netherland . The first of these trading posts were Fort Nassau (1614, near present-day Albany ); Fort Orange (1624, on the Hudson River just south of the current city of Albany and created to replace Fort Nassau), developing into settlement Beverwijck (1647), and into what became Albany; Fort Amsterdam (1625, to develop into
17108-420: The coastlines and shallow inland seas of Euramerica. During the Late Silurian the pterygotid eurypterids, large and specialized forms with several new adaptations, such as large and flattened telsons capable of being used as rudders, and large and specialized chelicerae with enlarged pincers for handling (and potentially in some cases killing) prey appeared. Though the largest members of the family appeared in
17290-608: The colony from the Dutch in 1664, renaming it the Province of New York . During the American Revolutionary War , a group of colonists eventually succeeded in establishing independence, and the state ratified the then new United States Constitution in 1788. From the early 19th century, New York's development of its interior, beginning with the construction of the Erie Canal , gave it incomparable advantages over other regions of
17472-695: The cuticle) after which they underwent rapid and immediate growth. Some arthropods, such as insects and many crustaceans, undergo extreme changes over the course of maturing. Chelicerates, including eurypterids, are in general considered to be direct developers, undergoing no extreme changes after hatching (though extra body segments and extra limbs may be gained over the course of ontogeny in some lineages, such as xiphosurans and sea spiders ). Whether eurypterids were true direct developers (with hatchlings more or less being identical to adults) or hemianamorphic direct developers (with extra segments and limbs potentially being added during ontogeny) has been controversial in
17654-566: The deposits is difficult. In what is presumably the lower member of the formation, fossils have been found of fishes Bothriolepis and Vietnamaspis and plants Colpodexylon and Lepidodendropsis . These fossils suggest a Givetian - Frasnian (Middle to Late Devonian ) age. The presumed middle member of the formation, which has yielded the eurypterid fossils, also preserves brachiopod fossils ( Lingula ), bivalves ( Ptychoparia and Modiolopsis ) and fish remains. In addition to R. dosonensis , some fragmentary eurypterid fossils of
17836-427: The discussion of constructing seawalls and other coastal barriers around the shorelines of New York City and Long Island to minimize the risk from another such future event. Such risk is considered highly probable due to global warming and rising sea levels . On March 1, 2020, New York had its first confirmed case of COVID-19 after Washington (state) , two months prior. From May 19–20, Western New York and
18018-533: The downstate region's lower elevation, proximity to the Atlantic Ocean, and relatively lower latitude . Upstate New York experiences warm summers, marred by only occasional, brief intervals of sultry conditions, with long and cold winters. Western New York, particularly the Tug Hill region, receives heavy lake-effect snows , especially during the earlier portions of winter, before the surface of Lake Ontario itself
18200-402: The duration of the conflict, and consequently the focus of General George Washington 's intelligence network . On the notorious British prison ships of Wallabout Bay , more American combatants died than were killed in combat in every battle of the war combined. Both sides of combatants lost more soldiers to disease than to outright wounds. The first of two major British armies were captured by
18382-642: The eastern part of the state, without draining Lakes George or Champlain . Lake George empties at its north end into Lake Champlain, whose northern end extends into Canada, where it drains into the Richelieu River and then ultimately the Saint Lawrence River . The western section of the state is drained by the Allegheny River and rivers of the Susquehanna and Delaware River systems. Niagara Falls
18564-507: The environment was a river delta in an otherwise semi-arid environment. Eurypterid Eurypterids , often informally called sea scorpions , are a group of extinct arthropods that form the order Eurypterida . The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago . The group is likely to have appeared first either during
18746-409: The eurypterine suborder, had already been established at this point during the Middle Ordovician. The earliest known stylonurine eurypterid, Brachyopterus , is also Middle Ordovician in age. The presence of members of both suborders indicates that primitive stem-eurypterids would have preceded them, though these are so far unknown in the fossil record. The presence of several eurypterid clades during
18928-411: The eurypterine suborder. Only one group of stylonurines (the family Parastylonuridae ) went extinct in the Early Devonian. Only two families of eurypterines survived into the Late Devonian at all ( Adelophthalmidae and Waeringopteridae). The eurypterines experienced their most major declines in the Early Devonian, during which over 50% of their diversity was lost in just 10 million years. Stylonurines, on
19110-508: The eurypterine swimming paddles varied from group to group. In the Eurypteroidea , the paddles were similar in shape to oars. The condition of the joints in their appendages ensured their paddles could only be moved in near-horizontal planes, not upwards or downwards. Some other groups, such as the Pterygotioidea, would not have possessed this condition and were probably able to swim faster. Most eurypterines are generally agreed to have utilized
19292-512: The expansive State University of New York , which is among the largest university systems in the nation. New York City is home to the headquarters of the United Nations , and it is sometimes described as the world's most important city, the cultural , financial , and media epicenter, and the capital of the world. The Native American tribes in what is now New York were predominantly Iroquois and Algonquian . Long Island
19474-454: The family Pterygotidae. An isolated 12.7 centimeters (5.0 in) long fossil metastoma of the carcinosomatoid eurypterid Carcinosoma punctatum indicates the animal would have reached a length of 2.2 meters (7.2 ft) in life, rivalling the pterygotids in size. Another giant was Pentecopterus decorahensis , a primitive carcinosomatoid, which is estimated to have reached lengths of 1.7 meters (5.6 ft). Typical of large eurypterids
19656-552: The federal government assumed control over immigration. During that period, more than eight million immigrants passed through its doors (two of every three U.S. immigrants). When the federal government assumed control, it established the Bureau of Immigration , which chose the three-acre (1.2 ha) Ellis Island in Upper New York Harbor for an entry depot. Already federally controlled, the island had served as an ammunition depot. It
19838-476: The found tracks each being about 7.6 centimeters (3.0 in) in diameter. Other eurypterid ichnogenera include Merostomichnites (though it is likely that many specimens actually represent trackways of crustaceans) and Arcuites (which preserves grooves made by the swimming appendages). In eurypterids, the respiratory organs were located on the ventral body wall (the underside of the opisthosoma). Blattfüsse , evolved from opisthosomal appendages, covered
20020-410: The front of the carapace (although some juvenile specimens preserve a slight protrusion), they could be referred to Rhinocarcinosoma on account of the rounded type A genital appendage, the triangular shape of the carapace and the slender shape of the more anterior appendages. The fossil material of R. dosonensis is the most complete of any species of Rhinocarcinosoma . The new species R. dosonensis
20202-568: The gathering of food. In some groups, these spiny appendages became heavily specialized. In some eurypterids in the Carcinosomatoidea, forward-facing appendages were large and possessed enormously elongated spines (as in Mixopterus and Megalograptus ). In derived members of the Pterygotioidea, the appendages were completely without spines, but had specialized claws instead. Other eurypterids, lacking these specialized appendages, likely fed in
20384-651: The genus Strabops from the Cambrian of Missouri , are now classified as aglaspidids or strabopids . The aglaspidids, once seen as primitive chelicerates, are now seen as a group more closely related to trilobites. The fossil record of Ordovician eurypterids is quite poor. The majority of eurypterids once reportedly known from the Ordovician have since proven to be misidentifications or pseudofossils . Today only 11 species can be confidently identified as representing Ordovician eurypterids. These taxa fall into two distinct ecological categories; large and active predators from
20566-447: The genus Vernonaspis . The environment in which R. cicerops lived was a marginal marine one (influenced by both salt and fresh water, such as a lagoon or delta ). The eurypterid-bearing deposits of Oneida County, which yielded the fossils of R. vaningeni , have also yielded other eurypterids, such as Eurypterus remipes . The Dô Son Formation, where R. dosonensis was discovered, very rarely preserves fossils meaning that dating
20748-404: The genus Hughmilleria are also known from the sites. Because the rock layers are relatively homogenous, researchers first assumed that the entire formation was Givetian-Frasnian in age. However, it is now believed that the eurypterid-bearing deposits are of Late Silurian age given that Rhinocarcinosoma is otherwise exclusively known from Late Silurian deposits (though it is not impossible for
20930-513: The genus (of which the most common is the type species, E. remipes ) account for more than 90% (perhaps as many as 95%) of all known fossil eurypterid specimens. Despite their vast number, Eurypterus are only known from a relatively short temporal range, first appearing during the Late Llandovery epoch (around 432 million years ago) and being extinct by the end of the Pridoli epoch. Eurypterus
21112-505: The genus to have survived unnoticed until the Late Devonian), fish remains from the same layer are similar to Late Silurian fish and palynological samples collected suggest a Late Silurian age. As the deposits have not been precisely dated, R. dosonensis and the fauna it co-occurred with have not been dated to a more precise timespan than 'Late Silurian'. The sedimentology of the fossil deposits containing R. dosonensis suggest that
21294-429: The giant millipede Arthropleura , and are possibly vital for the evolution of giant size in arthropods. In addition to the lightweight giant eurypterids, some deep-bodied forms in the family Hibbertopteridae were also very large. A carapace from the Carboniferous of Scotland referred to the species Hibbertoperus scouleri measures 65 cm (26 in) wide. As Hibbertopterus was very wide compared to its length,
21476-412: The gills of other groups. To be functional gills, they would have to have been highly efficient and would have required a highly efficient circulatory system. It is considered unlikely, however, that these factors would be enough to explain the large discrepancy between gill tract size and body size. It has been suggested instead that the "gill tract" was an organ for breathing air, perhaps actually being
21658-405: The globe, with the highest count of any U.S. state in 2022. Many of its landmarks are well known, including four of the world's ten most-visited tourist attractions in 2013: Times Square , Central Park , Niagara Falls , and Grand Central Terminal . New York is home to approximately 200 colleges and universities , including Ivy League members Columbia University and Cornell University , and
21840-587: The group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event . They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event (or sometime shortly before) 251.9 million years ago. Although popularly called "sea scorpions", only the earliest eurypterids were marine ; many later forms lived in brackish or fresh water , and they were not true scorpions . Some studies suggest that
22022-432: The group lived primarily in the waters around and within the ancient supercontinent of Euramerica . Only a handful of eurypterid groups spread beyond the confines of Euramerica and a few genera, such as Adelophthalmus and Pterygotus , achieved a cosmopolitan distribution with fossils being found worldwide. Like all other arthropods , eurypterids possessed segmented bodies and jointed appendages (limbs) covered in
22204-580: The invaginations leading to asphyxiation . Furthermore, most eurypterids would have been aquatic their entire lives. No matter how much time was spent on land, organs for respiration in underwater environments must have been present. True gills, expected to have been located within the branchial chamber within the Blattfüssen , remain unknown in eurypterids. Like all arthropods, eurypterids matured and grew through static developmental stages referred to as instars . These instars were punctuated by periods during which eurypterids went through ecdysis (molting of
22386-491: The larger sizes of adults mean a higher drag coefficient , using this type of propulsion is more energy-efficient. Some eurypterines, such as Mixopterus (as inferred from attributed fossil trackways), were not necessarily good swimmers. It likely kept mostly to the bottom, using its swimming paddles for occasional bursts of movements vertically, with the fourth and fifth pairs of appendages positioned backwards to produce minor movement forwards. While walking, it probably used
22568-495: The largest known arthropod ever to have lived, is Jaekelopterus rhenaniae . A chelicera from the Emsian Klerf Formation of Willwerath, Germany measured 36.4 centimeters (14.3 in) in length, but is missing a quarter of its length, suggesting that the full chelicera would have been 45.5 centimeters (17.9 in) long. If the proportions between body length and chelicerae match those of its closest relatives, where
22750-516: The largest known arthropods ever to have lived. The largest, Jaekelopterus , reached 2.5 meters (8.2 ft) in length. Eurypterids were not uniformly large and most species were less than 20 centimeters (8 in) long; the smallest eurypterid, Alkenopterus , was only 2.03 centimeters (0.80 in) long. Eurypterid fossils have been recovered from every continent. A majority of fossils are from fossil sites in North America and Europe because
22932-497: The last ever radiation within the eurypterids, which gave rise to several new forms capable of "sweep-feeding" (raking through the substrate in search of prey). Only three eurypterid families—Adelophthalmidae, Hibbertopteridae and Mycteroptidae—survived the extinction event in its entirety. It was assumed that these were all freshwater animals, which would have rendered the eurypterids extinct in marine environments, and with marine eurypterid predators gone, sarcopterygians , such as
23114-582: The late 20th century by the federally recognized tribes. New York put up more than 5 million acres (20,000 km ) of former Iroquois territory for sale in the years after the Revolutionary War, leading to rapid development in Upstate New York. As per the Treaty of Paris , the last vestige of British authority in the former Thirteen Colonies —their troops in New York City—departed in 1783, which
23296-480: The more anterior (forwards) appendages of Rhinocarcinosoma were more slender than those of Eusarcana . Although historically assumed to have had a scorpion-like telson (the posteriormost division of the body), like Eusarcana , more complete fossils of R. dosonensis revealed that this was not the case, although the telson did slightly curve upwards. Like Eusarcana , Rhinocarcinosoma possessed an ornamentation of scales on its carapace (head plate), though
23478-465: The most marine eurypterids, known from deposits that were once reefs, some in lagoonal settings, and deeper waters. In contrast to other carcinosomatids, Rhinocarcinosoma fossils are also known from non-marine environments, such as fluvial (river) and lacustrine (lake) settings. The shovel-like development in the front of the carapace of Rhinocarcinosoma suggests a "mud-grubbing" and bottom-dwelling lifestyle. More evidence for Rhinocarcinosoma being
23660-516: The ocellar mound (the raised surface where the ocelli , smaller eyes, were located) being placed centrally and being the highest point of the carapace. The species R. cicerops shares the forward and prominent position of the ocelli with R. vaningeni , but only has a slight development of the snout compared to the others. Its fossils are also the smallest of any species, at only 4 centimetres (1.6 in) in length, but all known fossil specimens of R. cicerops are of immature individuals, meaning that it
23842-428: The ocellar mound was fully developed despite the specimen being immature, that he named a new species. Clarke and Ruedemann referred Clarke's species to the genus Eusarcus (now known as Eusarcana ) in 1912, designating it as Eusarcus (?) cicerops . Accompanying the reclassification were the assignment of several more fossils of varying sizes to the species. The reclassification to Eusarcus , although provisional,
24024-501: The only pair placed before the mouth, is called the chelicerae ( homologous to the fangs of spiders). They were equipped with small pincers used to manipulate food fragments and push them into the mouth. In one lineage, the Pterygotidae , the chelicerae were large and long, with strong, well-developed teeth on specialised chelae (claws). The subsequent pairs of appendages, numbers II to VI, possessed gnathobases (or "tooth-plates") on
24206-808: The opisthosoma was covered in structures evolved from modified opisthosomal appendages. Throughout the opisthosoma, these structures formed plate-like structures termed Blattfüsse ( lit. ' leaf-feet ' in German). These created a branchial chamber (gill tract) between preceding Blattfüsse and the ventral surface of the opisthosoma itself, which contained the respiratory organs. The second to sixth opisthosomal segments also contained oval or triangular organs that have been interpreted as organs that aid in respiration. These organs, termed Kiemenplatten or "gill tracts", would potentially have aided eurypterids to breathe air above water, while Blattfüssen , similar to organs in modern horseshoe crabs , would cover
24388-424: The other hand, persisted through the period with more or less consistent diversity and abundance but were affected during the Late Devonian, when many of the older groups were replaced by new forms in the families Mycteroptidae and Hibbertopteridae. It is possible that the catastrophic extinction patterns seen in the eurypterine suborder were related to the emergence of more derived fish. Eurypterine decline began at
24570-548: The parts that serve for underwater respiration . The appendages of opisthosomal segments 1 and 2 (the seventh and eighth segments overall) were fused into a structure termed the genital operculum, occupying most of the underside of the opisthosomal segment 2. Near the anterior margin of this structure, the genital appendage (also called the Zipfel or the median abdominal appendage) protruded. This appendage, often preserved very prominently, has consistently been interpreted as part of
24752-433: The past. Hemianamorphic direct development has been observed in many arthropod groups, such as trilobites , megacheirans , basal crustaceans and basal myriapods . True direct development has on occasion been referred to as a trait unique to arachnids . There have been few studies on eurypterid ontogeny as there is a general lack of specimens in the fossil record that can confidently be stated to represent juveniles. It
24934-611: The people for ratification. On July 30, 1777, George Clinton was inaugurated as the first Governor of New York at Kingston. Approximately a third of the battles of the American Revolutionary War took place in New York; the first major one and largest of the entire war was the Battle of Long Island , also known as the Battle of Brooklyn, in August 1776. After their victory, the British occupied present-day New York City, making it their military and political base of operations in North America for
25116-434: The point when jawless fish first became more developed and coincides with the emergence of placoderms (armored fish) in both North America and Europe. Stylonurines of the surviving hibbertopterid and mycteroptid families completely avoided competition with fish by evolving towards a new and distinct ecological niche. These families experienced a radiation and diversification through the Late Devonian and Early Carboniferous,
25298-591: The provinces of Ontario and Quebec in Canada, with New York and Ontario sharing the Thousand Islands archipelago within the Saint Lawrence River, while most of its border with Quebec is on land; it shares Lake Champlain with the New England state of Vermont ; the New England state of Massachusetts has mostly a land border; New York extends into Long Island Sound and the Atlantic Ocean , sharing
25480-452: The ratio between claw size and body length is relatively consistent, the specimen of Jaekelopterus that possessed the chelicera in question would have measured between 233 and 259 centimeters (7.64 and 8.50 ft), an average 2.5 meters (8.2 ft), in length. With the chelicerae extended, another meter (3.28 ft) would be added to this length. This estimate exceeds the maximum body size of all other known giant arthropods by almost half
25662-400: The region. Maritime or shore birds of New York are great blue heron , killdeers , northern cardinals , American herring gulls , and common terns . Reptile and amphibian species in land areas of New York include queen snakes , hellbenders , diamondback terrapins , timber rattlesnakes , eastern fence lizards , spotted turtles , and Blanding's turtles . Sea turtles that can be found in
25844-686: The related genus Carcinosoma and the Greek ῥινός (rhinós, "nose"). The holotype specimen of R. dosonensis was discovered in a quarry in 1989 on the Ngọc Xuyên hill on the Dô Son Peninsula in northern Vietnam, part of the Dô Son Formation . Initially believed to be the remains of a chasmataspidid , it was recognised as probably being a carcinosomatid fossil in 1993. Further fossils of R. dosonensis were discovered in November 1993 at two quarries on
26026-445: The reproduction and sexual dimorphism of eurypterids is difficult, as they are only known from fossilized shells and carapaces. In some cases, there might not be enough apparent differences to separate the sexes based on morphology alone. Sometimes two sexes of the same species have been interpreted as two different species, as was the case with two species of Drepanopterus ( D. bembycoides and D. lobatus ). The eurypterid prosoma
26208-458: The reproductive system and occurs in two recognized types, assumed to correspond to male and female. Eurypterids were highly variable in size, depending on factors such as lifestyle, living environment and taxonomic affinity . Sizes around 100 centimeters (3.3 ft) are common in most eurypterid groups. The smallest eurypterid, Alkenopterus burglahrensis , measured just 2.03 centimeters (0.80 in) in length. The largest eurypterid, and
26390-592: The rest of the State of New York. The placement of a boundary between the two is a matter of great contention. Unofficial and loosely defined regions of Upstate New York include from the Southern Tier, which includes many of the counties along the border with Pennsylvania, to the North Country region, above or sometimes including parts of the Adirondack region. Among the total area of New York state, 13.6% consists of water. Much of New York's boundaries are in water, as
26572-435: The rest of the former supercontinent Gondwana , the discoveries of trackways both predate and outnumber eurypterid body fossils. Eurypterid trackways have been referred to several ichnogenera, most notably Palmichnium (defined as a series of four tracks often with an associated drag mark in the mid-line), wherein the holotype of the ichnospecies P. kosinkiorum preserves the largest eurypterid footprints known to date with
26754-452: The right to representative government . At the same time, given strong commercial, personal and sentimental links to Britain , many New York residents were Loyalists . The Capture of Fort Ticonderoga provided the cannon and gunpowder necessary to force a British withdrawal from the siege of Boston in 1775. New York was the only colony not to vote for independence , as the delegates were not authorized to do so. New York then endorsed
26936-425: The same degree, as well as the position of the ocelli being similar, was noted by Clarke and Ruedemann as similarities between the two species. E. vaningeni and E. cicerops were ultimately placed in their own genus, Rhinocarcinosoma , by Nestor Ivanovich Novozhilov in 1962, based on the placement of the eyes, the shape of the carapace and the protrusion at the front of the carapace. The genus name derives from
27118-405: The same genera. The primary function of the long, assumed female, type A appendages was likely to take up spermatophore from the substrate into the reproductive tract rather than to serve as an ovipositor, as arthropod ovipositors are generally longer than eurypterid type A appendages. By rotating the sides of the operculum, it would have been possible to lower the appendage from the body. Due to
27300-447: The scales of Rhinocarcinosoma were smaller and more closely arranged. The type A genital appendage (female reproductive organ) of Rhinocarcinosoma was broader and more rounded than that of Eusarcana . The most distinctive feature of Rhinocarcinosoma was the shovel-shaped protrusion developed from the most forward-facing portion of the carapace. It is possible that the snout was used for digging purposes. As of yet, this feature
27482-413: The short stride length indicates that Hibbertopterus crawled with an exceptionally slow speed, at least on land. The large telson was dragged along the ground and left a large central groove behind the animal. Slopes in the tracks at random intervals suggest that the motion was jerky. The gait of smaller stylonurines, such as Parastylonurus , was probably faster and more precise. The functionality of
27664-700: The site on March 21, 2014. Upon its completion in 2014, the new One World Trade Center became the tallest skyscraper in the Western Hemisphere, at 1,776 feet (541 m), meant to symbolize the year America gained its independence , 1776. From 2006 to 2018, 3 World Trade Center , 4 World Trade Center , 7 World Trade Center, the World Trade Center Transportation Hub , Liberty Park , and Fiterman Hall were completed. St. Nicholas Greek Orthodox Church and Ronald O. Perelman Performing Arts Center are under construction at
27846-429: The sixth pair of appendages were overlaid by a plate that is referred to as the metastoma, originally derived from a complete exoskeleton segment. The opisthosoma itself can be divided either into a " mesosoma " (comprising segments 1 to 6) and " metasoma " (comprising segments 7 to 12) or into a "preabdomen" (generally comprising segments 1 to 7) and "postabdomen" (generally comprising segments 8 to 12). The underside of
28028-441: The species Lanarkopterus dolichoschelus from the Ordovician of Ohio contain fragments of jawless fish and fragments of smaller specimens of Lanarkopterus itself. Though apex predatory roles would have been limited to the very largest eurypterids, smaller eurypterids were likely formidable predators in their own right just like their larger relatives. As in many other entirely extinct groups, understanding and researching
28210-427: The spongy structure of the eurypterid gill tracts. It is possible the two organs functioned in the same way. Some researchers have suggested that eurypterids may have been adapted to an amphibious lifestyle, using the full gill tract structure as gills and the invaginations within it as pseudotrachea. This mode of life may not have been physiologically possible, however, since water pressure would have forced water into
28392-463: The state are the green sea turtle , loggerhead sea turtle , leatherback sea turtle and Kemp's ridley sea turtle . New York Harbor and the Hudson River constitute an estuary , making the state of New York home to a rich array of marine life including shellfish—such as oysters and clams —as well as fish, microorganisms, and sea-birds. Due to its long history, New York has several overlapping and often conflicting definitions of regions within
28574-492: The structure. Though the Kiemenplatte is referred to as a "gill tract", it may not necessarily have functioned as actual gills. In other animals, gills are used for oxygen uptake from water and are outgrowths of the body wall. Despite eurypterids clearly being primarily aquatic animals that almost certainly evolved underwater (some eurypterids, such as the pterygotids, would even have been physically unable to walk on land), it
28756-480: The town New Amsterdam , which is present-day New York City); and Esopus (1653, now Kingston ). The success of the patroonship of Rensselaerswyck (1630), which surrounded Albany and lasted until the mid-19th century, was also a key factor in the early success of the colony. The English captured the colony during the Second Anglo-Dutch War and governed it as the Province of New York . The city of New York
28938-467: The two planes. Since September 11, most of Lower Manhattan has been restored. In the years since, over 7,000 rescue workers and residents of the area have developed several life-threatening illnesses, and some have died. A memorial at the site, the National September ;11 Memorial & Museum , was opened to the public on September 11, 2011. A permanent museum later opened at
29120-464: The type A appendage is divided into three but the type B appendage into only two. Such division of the genital appendage is common in eurypterids, but the number is not universal; for instance, the appendages of both types in the family Pterygotidae are undivided. The type A appendage is also armed with two curved spines called furca (lit. 'fork' in Latin). The presence of furca in the type B appendage
29302-414: The type A appendage, could have been used to detect whether a substrate was suitable for spermatophore deposition. Until 1882 no eurypterids were known from before the Silurian. Contemporary discoveries since the 1880s have expanded the knowledge of early eurypterids from the Ordovician period. The earliest eurypterids known today, the megalograptid Pentecopterus , date from the Darriwilian stage of
29484-400: The underside and created a gill chamber where the "gill tracts" were located. Depending on the species, the eurypterid gill tract was either triangular or oval in shape and was possibly raised into a cushion-like state. The surface of this gill tract bore several spinules (small spines), which resulted in an enlarged surface area. It was composed of spongy tissue due to many invaginations in
29666-426: The vast majority of the state's geographic area is dominated by meadows , forests , rivers, farms, mountains, and lakes. Most of the southern part of the state rests on the Allegheny Plateau , which extends from the southeastern United States to the Catskill Mountains ; the section in the State of New York is known as the Southern Tier . The rugged Adirondack Mountains , with vast tracts of wilderness, lie west of
29848-429: The way different plates overlay at its location, the appendage would have been impossible to move without muscular contractions moving around the operculum. It would have been kept in place when not it use. The furca on the type A appendages may have aided in breaking open the spermatophore to release the free sperm inside for uptake. The "horn organs," possibly spermathecae, are thought to have been connected directly to
30030-475: Was R. cicerops , described by Clarke in 1907 as Eurypterus ? cicerops . The species was described based on a single carapace discovered in the Shawangunk grit at Otisville, New York , of Llandovery - Ludlow age (dating not entirely certain but likely only Late Silurian, i.e. Ludlow). Though the specimen was immature, Clarke considered the fossil so unusual, in that the compound eyes were so developed and
30212-569: Was a major ocean port and had extensive traffic importing cotton from the South and exporting manufacturing goods. Nearly half of the state's exports were related to cotton. Southern cotton factors, planters and bankers visited so often that they had favorite hotels. At the same time, activism for abolitionism was strong upstate, where some communities provided stops on the Underground Railroad . Upstate, and New York City, gave strong support for
30394-476: Was a medium-sized carcinosomatid eurypterid , with the largest species, R. vaningeni , reaching lengths of 39 centimetres (15.4 in) and the second largest, R. dosonensis , reaching lengths of 22 centimetres (8.7 in). In terms of the outline of the body, with a broad abdomen and a nearly tubular postabdomen (tail), Rhinocarcinosoma was similar to the related genus Eusarcana . Its appendages, large and with spines, are also similar to those of Eusarcana , though
30576-467: Was a medium-sized carcinosomatid eurypterid, with the largest species, R. vaningeni , reaching lengths of 39 centimetres (15.4 in). In contrast to other carcinosomatids, Rhinocarcinosoma is not known only from marine settings, but also from deposits that were once lakes or rivers. It was adapted to a bottom-dwelling lifestyle, as either a burrowing or digging scavenger or top predator , feeding on other invertebrates and small fish. Rhinocarcinosoma
30758-400: Was also restricted to the continent Euramerica (composed of the equatorial continents Avalonia, Baltica and Laurentia), which had been completely colonized by the genus during its merging and was unable to cross the vast expanses of ocean separating this continent from other parts of the world, such as the southern supercontinent Gondwana. As such, Eurypterus was limited geographically to
30940-581: Was based on the head and abdomen of E ?. cicerops resembling species of that genus, in particular in the width of the body, the sub-triangular outline of the carapace and the oval shape and more or less marginal position of the eyes. The species R. vaningeni was also described by Clarke and Ruedemann in The Eurypterida of New York , like R. cicerops designated as a species of Eusarcus (as Eusarcus vaningeni ). Regarded by Clarke and Ruedemann as "very unexpected and peculiar" and "puzzling", E. vaningeni
31122-423: Was by expensive wagons on muddy roads before canals opened up the rich farmlands to long-distance traffic. Governor DeWitt Clinton promoted the Erie Canal , which connected New York City to the Great Lakes by the Hudson River , the new canal, and the rivers and lakes. Work commenced in 1817, and the Erie Canal opened eight years later, in 1825. Packet boats pulled by horses on tow paths traveled slowly over
31304-589: Was chosen due its relative isolation with proximity to New York City and the rail lines of Jersey City, New Jersey , via a short ferry ride. While the island was being developed and expanded via land reclamation , the federal government operated a temporary depot at the Barge Office at the Battery. Ellis Island opened on January 1, 1892, and operated as a central immigration center until the National Origins Act
31486-426: Was created for the fossils given that they differed from both R. cicerops and R. vaningeni . A juvenile specimen of R. dosonensis , of about the same size as specimens of R. cicerops , has a markedly different carapace shape and whereas the metastoma (a plate on the underside of the abdomen) of R. vaningeni expands in size anteriorly, it stays about the same size in R. dosonensis . Though Rhinocarcinosoma
31668-516: Was described based on fossils from Oriskany Creek in Oneida County, New York , of Ludlow age. Although Clarke and Ruedemann considered E. vaningeni to be undoubtedly similar to Eusarcus scorpionis (the generic type species) in general, the position of the ocelli between the eyes and the large shovel-like projection in the front represented considerable differences from the type material. The snout also being developed in R. cicerops , though not to
31850-497: Was discovered in Carboniferous-aged fossil deposits of Scotland in 2005. It was attributed to the stylonurine eurypterid Hibbertopterus due to a matching size (the trackmaker was estimated to have been about 1.6 meters (5.2 ft) long) and inferred leg anatomy. It is the largest terrestrial trackway—measuring 6 meters (20 ft) long and averaging 95 centimeters (3.12 ft) in width—made by an arthropod found thus far. It
32032-638: Was divided roughly in half between the Algonquian Wampanoag and Lenape peoples. The Lenape also controlled most of the region surrounding New York Harbor . North of the Lenape was a third Algonquian nation, the Mohicans . Starting north of them, from east to west, were two Iroquoian nations: the Mohawk —part of the original Iroquois Five Nations, and the Petun . South of them, divided roughly along Appalachia , were
32214-552: Was long afterward celebrated as Evacuation Day . New York City was the national capital under the Articles of Confederation and Perpetual Union , the first national government. That organization was found to be inadequate, and prominent New Yorker Alexander Hamilton advocated for a new government that would include an executive, national courts, and the power to tax. Hamilton led the Annapolis Convention (1786) that called for
32396-452: Was more or less parallel and similar to that of extinct and extant xiphosurans, with the largest exception being that eurypterids hatched with a full set of appendages and opisthosomal segments. Eurypterids were thus not hemianamorphic direct developers, but true direct developers like modern arachnids. The most frequently observed change occurring through ontogeny (except for some genera, such as Eurypterus , which appear to have been static)
32578-612: Was one of the original Thirteen Colonies that went on to form the United States. The area of present-day New York had been inhabited by tribes of the Algonquians and the Iroquois Confederacy Native Americans for several thousand years by the time the earliest Europeans arrived. Stemming from Henry Hudson 's expedition in 1609, the Dutch established the multiethnic colony of New Netherland in 1621. England seized
32760-536: Was passed in 1924, reducing immigration. After that date, the only immigrants to pass through were displaced persons or war refugees . The island ceased all immigration processing on November 12, 1954, when the last person detained on the island, Norwegian seaman Arne Peterssen, was released. He had overstayed his shore leave and left on the 10:15 a.m. Manhattan-bound ferry to return to his ship. More than 12 million immigrants passed through Ellis Island between 1892 and 1954. More than 100 million Americans across
32942-402: Was recaptured by the Dutch in 1673 during the Third Anglo-Dutch War (1672–1674) and renamed New Orange. It was returned to the English under the terms of the Treaty of Westminster a year later. The Sons of Liberty were organized in New York City during the 1760s, largely in response to the oppressive Stamp Act passed by the British Parliament in 1765. The Stamp Act Congress met in
33124-518: Was used as an ovipositor (used to deposit eggs). The different types of genital appendages are not necessarily the only feature that distinguishes between the sexes of eurypterids. Depending on the genus and species in question, other features such as size, the amount of ornamentation and the proportional width of the body can be the result of sexual dimorphism. In general, eurypterids with type B appendages (males) appear to have been proportionally wider than eurypterids with type A appendages (females) of
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