118-558: The raspberry is the edible fruit of several plant species in the genus Rubus of the rose family , most of which are in the subgenus Idaeobatus . The name also applies to these plants themselves. Raspberries are perennial with woody stems . World production of raspberries in 2022 was 947,852 tonnes , led by Russia with 22% of the total. Raspberries are cultivated across northern Europe and North America and are consumed in various ways, including as whole fruit and in preserves , cakes, ice cream, and liqueurs . Raspberries are
236-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
354-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
472-424: A "true" cultivar name – the recognized scientific name in the public domain – and a "commercial synonym" – an additional marketing name that is legally protected. An example would be Rosa Fascination = 'Poulmax', in which Rosa is the genus, Fascination is the trade designation, and 'Poulmax' is scientific cultivar name. Because a name that is attractive in one language may have less appeal in another country,
590-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
708-424: A component of a cultigen can be accepted as a cultivar if it is recognisable and has stable characters. Therefore, all cultivars are cultigens, because they are cultivated, but not all cultigens are cultivars, because some cultigens have not been formally distinguished and named as cultivars. The Cultivated Plant Code notes that the word cultivar is used in two different senses: first, as a "classification category"
826-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
944-436: A constant state of development which makes the naming of such an assemblage as a cultivar a futile exercise." However, retired transgenic varieties such as the fish tomato , which are no longer being developed, do not run into this obstacle and can be given a cultivar name. Cultivars may be selected because of a change in the ploidy level of a plant which may produce more desirable characteristics. Every unique cultivar has
1062-796: A cultivar. Some cultivars "come true from seed", retaining their distinguishing characteristics when grown from seed. Such plants are termed a "variety", "selection", or "strain" but these are ambiguous and confusing words that are best avoided. In general, asexually propagated cultivars grown from seeds produce highly variable seedling plants, and should not be labelled with, or sold under, the parent cultivar's name. Seed-raised cultivars may be produced by uncontrolled pollination when characteristics that are distinct, uniform and stable are passed from parents to progeny. Some are produced as "lines" that are produced by repeated self-fertilization or inbreeding or "multilines" that are made up of several closely related lines. Sometimes they are F1 hybrids which are
1180-783: A database of new cultivars protected by PBR in all countries. An International Cultivar Registration Authority (ICRA) is a voluntary, non-statutory organization appointed by the Commission for Nomenclature and Cultivar Registration of the International Society of Horticultural Science. ICRAs are generally formed by societies and institutions specializing in particular plant genera such as Dahlia or Rhododendron and are currently located in Europe, North America, China, India, Singapore, Australia, New Zealand, South Africa and Puerto Rico. Each ICRA produces an annual report and its reappointment
1298-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
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#17328584446721416-421: A direct threat to raspberry plants but also increase the risk of transmitting various plant diseases. This dual impact can significantly undermine agricultural productivity, making it crucial for raspberry growers to implement effective pest management strategies to mitigate the damage caused by Japanese beetle infestations. In 2022, world production of raspberries was 947,852 tonnes , led by Russia with 22% of
1534-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
1652-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On
1770-1022: A low- glycemic index food, with total sugar content of only 4% and no starch . The aggregate fruit structure contributes to raspberry's nutritional value, as it increases the proportion of dietary fiber, which is among the highest known in whole foods – up to 6% fiber per total weight. Raspberries contain phytochemicals , such as anthocyanin pigments , ellagic acid , ellagitannins , quercetin , gallic acid , cyanidins , pelargonidins , catechins , kaempferol and salicylic acid . Yellow raspberries and others with pale-colored fruits are lower in anthocyanin content. Both yellow and red raspberries contain carotenoids, mostly lutein esters, but these are masked by anthocyanins in red raspberries. Raspberry compounds are under preliminary research for their potential to affect human health. Raspberry leaves can be used fresh or dried in herbal teas , providing an astringent flavor. In herbal and traditional medicine , raspberry leaves are used for some remedies, although there
1888-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
2006-563: A modern vernacular language to distinguish them from botanical epithets. For example, the full cultivar name of the King Edward potato is Solanum tuberosum 'King Edward'. 'King Edward' is the cultivar epithet, which, according to the Rules of the Cultivated Plant Code , is bounded by single quotation marks. For patented or trademarked plant product lines developed from a given cultivar,
2124-413: A name, the first letter of the word "Group" is itself capitalized. Since the 1990s there has been an increasing use of legal protection for newly produced cultivars. Plant breeders expect legal protection for the cultivars they produce. According to proponents of such protections, if other growers can immediately propagate and sell these cultivars as soon as they come on the market, the breeder's benefit
2242-530: A pH between 6 and 7 with ample organic matter to assist in retaining water. While moisture is essential, wet and heavy soils or excess irrigation can bring on Phytophthora root rot, which is one of the most serious pest problems facing the red raspberry. As a cultivated plant in moist, temperate regions, it is easy to grow and tends to spread unless pruned. Escaped raspberries frequently appear as garden weeds, spread by seeds found in bird droppings. An individual raspberry weighs 3–5 g (0.11–0.18 oz) and
2360-491: A particular disease. Genetically modified plants with characteristics resulting from the deliberate implantation of genetic material from a different germplasm may form a cultivar. However, the International Code of Nomenclature for Cultivated Plants notes, "In practice such an assemblage is often marketed from one or more lines or multilines that have been genetically modified. These lines or multilines often remain in
2478-595: A particular part of the plant, such as a lateral branch, or from a particular phase of the life cycle, such as a juvenile leaf, or from aberrant growth as occurs with witch's broom . Plants whose distinctive characters are derived from the presence of an intracellular organism may also form a cultivar provided the characters are reproduced reliably from generation to generation. Plants of the same chimera (which have mutant tissues close to normal tissue) or graft-chimeras (which have vegetative tissue from different kinds of plants and which originate by grafting) may also constitute
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#17328584446722596-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
2714-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that
2832-523: A plant may be given different selling names from country to country. Quoting the original cultivar name allows the correct identification of cultivars around the world. The main body coordinating plant breeders' rights is the International Union for the Protection of New Varieties of Plants ( Union internationale pour la protection des obtentions végétales , UPOV) and this organization maintains
2950-410: A rich source of vitamin C , manganese , and dietary fiber . A raspberry is an aggregate fruit , developing from the numerous distinct carpels of a single flower. What distinguishes the raspberry from its blackberry relatives is whether or not the torus ( receptacle or stem) "picks with" (i.e., stays with) the fruit. When picking a blackberry fruit, the torus stays with the fruit. With a raspberry,
3068-682: A rough surface, related to the Old English rasp or "rough berry". Examples of raspberry species in Rubus subgenus Idaeobatus include: Several species of Rubus , also called raspberries, are classified in other subgenera, including: Various kinds of raspberries can be cultivated from hardiness zones 3 to 9. Raspberries are traditionally planted in the winter as dormant canes, although planting of tender, plug plants produced by tissue culture has become much more common. A specialized production system called "long cane production" involves growing canes for
3186-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
3304-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
3422-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
3540-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
3658-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
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3776-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
3894-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
4012-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
4130-462: A unique name within its denomination class (which is almost always the genus). Names of cultivars are regulated by the International Code of Nomenclature for Cultivated Plants , and may be registered with an International Cultivar Registration Authority (ICRA). There are sometimes separate registration authorities for different plant types such as roses and camellias. In addition, cultivars may be associated with commercial marketing names referred to in
4248-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
4366-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
4484-690: A vernacular language. From circa the 1900s, cultivated plants in Europe were recognised in the Scandinavian, Germanic, and Slavic literature as stamm or sorte , but these words could not be used internationally because, by international agreement, any new denominations had to be in Latin. In the twentieth century an improved international nomenclature was proposed for cultivated plants. Liberty Hyde Bailey of Cornell University in New York , United States created
4602-485: A year in a northern climate such as Scotland or Oregon or Washington , where the chilling requirement for proper bud break is attained, or attained earlier than the ultimate place of planting. These canes are then dug, roots and all, to be replanted in warmer climates such as Spain, where they quickly flower and produce a very early season crop. Plants are typically planted 2–6 per m in fertile, well drained soil; raspberries are usually planted in raised beds/ridges, if there
4720-409: Is capitalized (with some permitted exceptions such as conjunctions). It is permissible to place a cultivar epithet after a common name provided the common name is botanically unambiguous. Cultivar epithets published before 1 January 1959 were often given a Latin form and can be readily confused with the specific epithets in botanical names; after that date, newly coined cultivar epithets must be in
4838-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
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4956-404: Is a common fungal infection of raspberries and other soft fruit under wet conditions. It is seen as a gray mold growing on the raspberries, and particularly affects fruit which are bruised, as the bruises provide an easy entrance point for the spores. Raspberry plants should not be planted where potatoes, tomatoes, peppers, eggplants, or bulbs have previously been grown, without prior fumigation of
5074-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
5192-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
5310-529: Is any plant that is deliberately selected for or altered in cultivation, as opposed to an indigen ; the Cultivated Plant Code states that cultigens are "maintained as recognisable entities solely by continued propagation". Cultigens can have names at any of many taxonomic ranks, including those of grex , species , cultivar group , variety , form , and cultivar; and they may be plants that have been altered in cultivation, including by genetic modification , but have not been formally denominated. A cultigen or
5428-620: Is any question about root rot problems. All cultivars of raspberries have perennial roots, but many do not have perennial shoots. In fact, most raspberries have shoots that are biennial (meaning shoots grow in the first growing season and fruits grow on those shoots during the second growing season). The flowers can be a major nectar source for honeybees and other pollinators. Raspberries are vigorous and can be locally invasive. They propagate using basal shoots (also known as suckers), extended underground shoots that develop roots and individual plants. They can sucker new canes some distance from
5546-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
5664-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,
5782-444: Is considered every four years. The main task is to maintain a register of the names within the group of interest and where possible this is published and placed in the public domain. One major aim is to prevent the duplication of cultivar and Group epithets within a genus, as well as ensuring that names are in accord with the latest edition of the Cultivated Plant Code . In this way, over the last 50 years or so, ICRAs have contributed to
5900-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
6018-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
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#17328584446726136-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
6254-404: Is largely lost. Legal protection for cultivars is obtained through the use of Plant breeders' rights and plant Patents but the specific legislation and procedures needed to take advantage of this protection vary from country to country. The use of legal protection for cultivars can be controversial, particularly for food crops that are staples in developing countries, or for plants selected from
6372-468: Is made up of around 100 drupelets , each of which consists of a juicy pulp and a single central seed. A raspberry bush can yield several hundred berries a year. Raspberry derives its name from raspise , "a sweet rose-colored wine" (mid-15th century), from the Anglo-Latin vinum raspeys , or from raspoie , meaning "thicket", of Germanic origin. The name may have been influenced by its appearance as having
6490-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
6608-571: Is no scientifically valid evidence to support their medicinal use. Species A species ( pl. : species) is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use
6726-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
6844-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
6962-432: Is simply a matter of convenience as the category was created to serve the practical needs of horticulture , agriculture , and forestry . Members of a particular cultivar are not necessarily genetically identical. The Cultivated Plant Code emphasizes that different cultivated plants may be accepted as different cultivars, even if they have the same genome, while cultivated plants with different genomes may be regarded as
7080-853: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Cultivar A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated . Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting , tissue culture , or carefully controlled seed production. Most cultivars arise from deliberate human manipulation , but some originate from wild plants that have distinctive characteristics. Cultivar names are chosen according to rules of
7198-424: Is the sense of cultivar that is most generally understood and which is used as a general definition. A cultivar is an assemblage of plants that (a) has been selected for a particular character or combination of characters, (b) is distinct, uniform and stable in those characters, and (c) when propagated by appropriate means, retains those characters. Which plants are chosen to be named as cultivars
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#17328584446727316-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
7434-440: Is to replace the Latin scientific names on plant labels in retail outlets with appealing marketing names that are easy to use, pronounce, and remember. Marketing names lie outside the scope of the Cultivated Plant Code which refers to them as "trade designations". If a retailer or wholesaler has the sole legal rights to a marketing name then that may offer a sales advantage. Plants protected by plant breeders' rights (PBR) may have
7552-402: The Cultivated Plant Code as "trade designations" (see below). A cultivar name consists of a botanical name (of a genus, species , infraspecific taxon , interspecific hybrid or intergeneric hybrid) followed by a cultivar epithet . The cultivar epithet is enclosed by single quotes; it should not be italicized if the botanical name is italicized; and each of the words within the epithet
7670-417: The Cultivated Plant Code . Each ICRA also ensures that new names are formally established (i.e. published in hard copy, with a description in a dated publication). They record details about the plant, such as parentage, the names of those concerned with its development and introduction, and a basic description highlighting its distinctive characters. ICRAs are not responsible for assessing the distinctiveness of
7788-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
7906-495: The International Code of Nomenclature for Cultivated Plants (ICNCP), and not all cultivated plants qualify as cultivars. Horticulturists generally believe the word cultivar was coined as a term meaning " cultivated variety ". Popular ornamental plants like roses , camellias , daffodils , rhododendrons , and azaleas are commonly cultivars produced by breeding and selection or as sports , for floral colour or size, plant form, or other desirable characteristics. Similarly,
8024-591: The Latin names in Linnaeus ' (1707–1778) Species Plantarum (tenth edition) and Genera Plantarum (fifth edition). In Species Plantarum , Linnaeus enumerated all plants known to him, either directly or from his extensive reading. He recognised the rank of varietas (botanical "variety", a rank below that of species and subspecies ) and he indicated these varieties with letters of the Greek alphabet , such as α, β, and λ, before
8142-514: The cultigen , which is defined as a plant whose origin or selection is primarily due to intentional human activity. A cultivar is not the same as a botanical variety , which is a taxonomic rank below subspecies , and there are differences in the rules for creating and using the names of botanical varieties and cultivars. In recent times, the naming of cultivars has been complicated by the use of statutory patents for plants and recognition of plant breeders' rights . The International Union for
8260-415: The etymology and it has been suggested that the word is actually a blend of culti gen and var iety . The neologism cultivar was promoted as "euphonious" and "free from ambiguity". The first Cultivated Plant Code of 1953 subsequently commended its use, and by 1960 it had achieved common international acceptance. The words cultigen and cultivar may be confused with each other. A cultigen
8378-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
8496-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
8614-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
8732-457: The 1995 edition, it is still widely used and recommended by other authorities. Where several very similar cultivars exist they can be associated into a Group (formerly Cultivar-group ). As Group names are used with cultivar names it is necessary to understand their way of presentation. Group names are presented in normal type and the first letter of each word capitalised as for cultivars, but they are not placed in single quotes. When used in
8850-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
8968-443: The Protection of New Varieties of Plants (UPOV – French : Union internationale pour la protection des obtentions végétales ) offers legal protection of plant cultivars to persons or organisations that introduce new cultivars to commerce. UPOV requires that a cultivar be "distinct", "uniform", and "stable". To be "distinct", it must have characters that easily distinguish it from any other known cultivar. To be "uniform" and "stable",
9086-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
9204-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
9322-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
9440-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
9558-473: The botanical variety except in respect to its origin. In that essay, Bailey used only the rank of species for the cultigen, but it was obvious to him that many domesticated plants were more like botanical varieties than species, and that realization appears to have motivated the suggestion of the new category of cultivar . Bailey created the word cultivar . It is generally assumed to be a blend of culti vated and var iety but Bailey never explicitly stated
9676-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be
9794-421: The commercial product name is typically indicated by the symbols "TM" or "®", or is presented in capital letters with no quotation marks, following the cultivar name, as in the following example, where "Bloomerang" is the commercial name and 'Penda' is the cultivar epithet: Syringa 'Penda' BLOOMERANG. Although "cv." has not been permitted by the International Code of Nomenclature for Cultivated Plants since
9912-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
10030-416: The concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part
10148-491: The cultivar is defined in Article 2 of the International Code of Nomenclature for Cultivated Plants (2009, 8th edition) as follows: The basic category of cultivated plants whose nomenclature is governed by this Code is the cultivar. There are two other classification categories for cultigens, the grex and the group . The Code then defines a cultivar as a "taxonomic unit within the classification category of cultivar". This
10266-472: The cultivar must retain these characters in repeated propagation. The naming of cultivars is an important aspect of cultivated plant taxonomy , and the correct naming of a cultivar is prescribed by the Rules and Recommendations of the International Code of Nomenclature for Cultivated Plants (ICNCP, commonly denominated the Cultivated Plant Code ). A cultivar is given a cultivar name, which consists of
10384-479: The eastern United States are derivatives of red raspberries. Yellow-fruited variants of the black raspberry are sometimes grown in home gardens. Red raspberries have also been crossed with various species in other subgenera of the genus Rubus , resulting in a number of hybrids , the first of which was the loganberry . Later notable hybrids include the olallieberry , boysenberry , marionberry , and tayberry ; all are multi-generational hybrids. Hybridization between
10502-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
10620-468: The familiar cultivated red raspberries and a few Asiatic species of Rubus has also been achieved. Numerous raspberry cultivars have been selected. Two types of raspberry are available for commercial and domestic cultivation; the summer-bearing type produces an abundance of fruit on second-year canes (floricanes) within a relatively short period in midsummer, and double or "everbearing" plants, which also bear some fruit on first-year canes (primocanes) in
10738-622: The late summer and fall, as well as the summer crop on second-year canes. Those marked (AGM) have gained the Royal Horticultural Society 's Award of Garden Merit . Raspberries are sometimes eaten by the larvae of some Lepidoptera species (butterflies and moths). More serious are the raspberry beetle (in Europe) and the raspberry fruitworm (in North America), whose larvae can damage raspberries. Botrytis cinerea , or gray mold,
10856-460: The main plant. For this reason, raspberries spread well, and can take over gardens if left unchecked. Raspberries are often propagated using cuttings, and will root readily in moist soil conditions. The fruit is harvested when it comes off the receptacle easily and has turned a deep color (red, black, purple, or golden yellow, depending on the species and cultivar). This is when the fruits are ripest and sweetest. High tunnel bramble production offers
10974-576: The native North American red raspberries classified as either R. idaeus subsp. strigosus , or R. idaeus var. strigosus . Recent breeding has resulted in cultivars that are thornless and more strongly upright, not needing staking. The black raspberry, Rubus occidentalis , is also cultivated, providing both fresh and frozen fruit, as well as jams, preserves, and other products, all with that species' distinctive flavor. Purple raspberries have been produced by horticultural hybridization of red and black raspberries, and have also been found in
11092-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
11210-408: The opportunity to bridge gaps in availability during late fall and late spring. Furthermore, high tunnels allow less hardy floricane-fruiting raspberries to overwinter in climates where they would not otherwise survive. In the tunnel, plants are established at close spacing usually prior to tunnel construction. Raspberries are an important commercial fruit crop, widely grown in all temperate regions of
11328-525: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
11446-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
11564-428: The result of a deliberate repeatable single cross between two pure lines. A few F2 hybrid seed cultivars also exist, such as Achillea 'Summer Berries'. Some cultivars are agamospermous plants, which retain their genetic composition and characteristics under reproduction. Occasionally cultivars are raised from seed of a specially selected provenance – for example the seed may be taken from plants that are resistant to
11682-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
11800-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
11918-899: The same cultivar. The production of cultivars generally entails considerable human involvement although in a few cases it may be as little as simply selecting variation from plants growing in the wild (whether by collecting growing tissue to propagate from or by gathering seed). Cultivars generally occur as ornamentals and food crops: Malus ' Granny Smith ' and Malus ' Red Delicious ' are cultivars of apples propagated by cuttings or grafting , Lactuca 'Red Sails' and Lactuca 'Great Lakes' are lettuce cultivars propagated by seeds. Named cultivars of Hosta and Hemerocallis plants are cultivars produced by micropropagation or division. Cultivars that are produced asexually are genetically identical and known as clones ; this includes plants propagated by division , layering , cuttings , grafts , and budding . The propagating material may be taken from
12036-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
12154-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
12272-464: The scientific Latin botanical name followed by a cultivar epithet . The cultivar epithet is usually in a vernacular language. The word cultivar originated from the need to distinguish between wild plants and those with characteristics that arose in cultivation, presently denominated cultigens . This distinction dates to the Greek philosopher Theophrastus (370–285 BC), the "Father of Botany", who
12390-450: The soil. These crops are hosts for the disease Verticillium wilt , a fungus that can stay in the soil for many years and can infest the raspberry crop. Raspberries, among other plants with high sugar content like peaches, are prime targets for the Japanese beetle , which relies heavily on these sources as its main food resource. The voracious feeding habits of Japanese beetles not only pose
12508-502: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
12626-475: The stability of cultivated plant nomenclature. In recent times many ICRAs have also recorded trade designations and trademarks used in labelling plant material, to avoid confusion with established names. New names and other relevant data are collected by and submitted to the ICRA and in most cases there is no cost. The ICRA then checks each new epithet to ensure that it has not been used before and that it conforms with
12744-407: The torus remains on the plant, leaving a hollow core in the raspberry fruit. Raspberries are grown for the fresh fruit market and for commercial processing into individually quick frozen (IQF) fruit, purée , juice, or dried fruit used in a variety of grocery products such as raspberry pie . Raspberries need ample sun and water for optimal development. Raspberries thrive in well-drained soil with
12862-640: The total (table). Other major producers were Mexico , Serbia , Poland , and the United States . Raw raspberries are 86% water, 12% carbohydrates , and have about 1% each of protein and fat (table). In a reference amount of 100 g (3.5 oz), raspberries supply 53 kilocalories and 6.5 grams of dietary fiber . Raspberries are a rich source of vitamin C (29% of the Daily Value , DV), manganese (29% DV), and dietary fiber (26% DV), but otherwise have low content of micronutrients (table). Raspberries are
12980-432: The varietal name, rather than using the abbreviation "var." as is the present convention. Most of the varieties that Linnaeus enumerated were of "garden" origin rather than being wild plants. In time the need to distinguish between wild plants and those with variations that had been cultivated increased. In the nineteenth century many "garden-derived" plants were given horticultural names, sometimes in Latin and sometimes in
13098-496: The whitebark raspberry, R. leucodermis . Both the red and the black raspberry species have albino-like pale-yellow natural or horticultural variants, resulting from presence of recessive genes that impede production of anthocyanin pigments. Fruits from such plants are called golden raspberries or yellow raspberries; despite their similar appearance, they retain the distinctive flavor of their respective species (red or black). Most pale-fruited raspberries commercially sold in
13216-428: The wild and propagated for sale without any additional breeding work; some people consider this practice unethical . The formal scientific name of a cultivar, like Solanum tuberosum 'King Edward', is a way of uniquely designating a particular kind of plant. This scientific name is in the public domain and cannot be legally protected. Plant retailers wish to maximize their share of the market and one way of doing this
13334-592: The wild in a few places (for example, in Vermont ) where the American red and the black raspberries both grow naturally. Commercial production of purple-fruited raspberries is rare. Blue raspberry is a local name used in Prince Edward County, Ontario , Canada, for the cultivar 'Columbian', a hybrid (purple raspberry) of R. strigosus and R. occidentalis . Blue raspberry can also refer to
13452-537: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
13570-414: The word cultivar in 1923 when he wrote that: The cultigen is a species, or its equivalent, that has appeared under domestication – the plant is cultigenous. I now propose another name, cultivar, for a botanical variety, or for a race subordinate to species, that has originated under cultivation; it is not necessarily, however, referable to a recognized botanical species. It is essentially the equivalent of
13688-406: The world's agricultural food crops are almost exclusively cultivars that have been selected for characters such as improved yield, flavour, and resistance to disease, and very few wild plants are now used as food sources. Trees used in forestry are also special selections grown for their enhanced quality and yield of timber . Cultivars form a major part of Liberty Hyde Bailey 's broader group,
13806-513: The world. Many of the most important modern commercial red raspberry cultivars derive from hybrids between R. idaeus and R. strigosus . Some botanists consider the Eurasian and American red raspberries to belong to a single, circumboreal species, Rubus idaeus , with the European plants then classified as either R. idaeus subsp. idaeus or R. idaeus var. idaeus , and
13924-477: Was keenly aware of this difference. Botanical historian Alan Morton noted that Theophrastus in his Historia Plantarum ( Enquiry into Plants ) "had an inkling of the limits of culturally induced ( phenotypic ) changes and of the importance of genetic constitution" ( Historia Plantarum , Book 3, 2, 2 and Causa Plantarum , Book 1, 9, 3). The International Code of Nomenclature for algae, fungi, and plants uses as its starting point for modern botanical nomenclature
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