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106-541: The shell mounds had been known to the local people for a long time. In 1877, the American zoologist Edward S. Morse (who would later teach zoology at the University of Tokyo ) came to Japan, and, on the second day of his arrival while riding Japan's first railway from Yokohama to Tokyo (founded in 1872), "found" these mounds and instantly felt their importance. Dr. Morse with his assistants later excavated these mounds from

212-775: A 23,000-years-old fisher-hunter-gatherers' camp on the shore of the Sea of Galilee , Northern Israel , provides the earliest evidence for the use of composite cereal harvesting tools. The Ohalo site is at the junction of the Upper Paleolithic and the Early Epipaleolithic , and has been attributed to both periods. The wear traces indicate that tools were used for harvesting near-ripe semi-green wild cereals, shortly before grains are ripe and disperse naturally. The studied tools were not used intensively, and they reflect two harvesting modes: flint knives held by hand and inserts hafted in

318-612: A corridor between the millet and rice cultivation centres where both rice and millet were cultivated. At around 5,500 to 4,000 BP, there was increasing migration into Taiwan from the early Austronesian Dapenkeng culture , bringing rice and millet cultivation technology with them. During this period, there is evidence of large settlements and intensive rice cultivation in Taiwan and the Penghu Islands , which may have resulted in overexploitation . Bellwood (2011) proposes that this may have been

424-589: A different problem: most of the wild seeds do not germinate in the first year; the first evidence of lentil domestication, breaking dormancy in their first year, appears in the early Neolithic at Jerf el Ahmar (in modern Syria), and lentils quickly spread south to the Netiv HaGdud site in the Jordan Valley . The process of domestication allowed the founder crops to adapt and eventually become larger, more easily harvested, more dependable in storage and more useful to

530-405: A food source, also had to be taken into account. Besides being a direct source of food, certain animals could provide leather, wool, hides, and fertilizer. Some of the earliest domesticated animals included dogs ( East Asia , about 15,000 years ago), sheep, goats, cows, and pigs. West Asia was the source for many animals that could be domesticated, such as sheep, goats and pigs. This area was also

636-619: A handle. The finds shed new light on cereal harvesting techniques some 8,000 years before the Natufian and 12,000 years before the establishment of sedentary farming communities in the Near East. Furthermore, the new finds accord well with evidence for the earliest ever cereal cultivation at the site and the use of stone-made grinding implements. Agriculture appeared first in West Asia about 2,000 years later, around 10,000–9,000 years ago. The region

742-746: A highly advantageous geographical location that afforded them a head start in the Neolithic Revolution. Both shared the temperate climate ideal for the first agricultural settings, and both were near a number of easily domesticable plant and animal species. In areas where continents aligned north–south such as the Americas and Africa, crops—and later domesticated animals—could not spread across tropical zones. Agriculture in Neolithic China can be separated into two broad regions, Northern China and Southern China. The agricultural centre in northern China

848-601: A laboratory setting. Ethologists have been particularly concerned with the evolution of behavior and the understanding of behavior in terms of the theory of natural selection . In one sense, the first modern ethologist was Charles Darwin , whose book, The Expression of the Emotions in Man and Animals , influenced many future ethologists. A subfield of ethology is behavioral ecology which attempts to answer Nikolaas Tinbergen 's four questions with regard to animal behavior: what are

954-473: A lifestyle of hunting and gathering to one of agriculture and settlement , making an increasingly large population possible. These settled communities permitted humans to observe and experiment with plants, learning how they grew and developed. This new knowledge led to the domestication of plants into crops . Archaeological data indicate that the domestication of various types of plants and animals happened in separate locations worldwide, starting in

1060-479: A narrow optimal climatic range outside of which they cannot grow for reasons of light or rain changes. For instance, wheat does not normally grow in tropical climates, just like tropical crops such as bananas do not grow in colder climates. Some authors, like Jared Diamond , have postulated that this east–west axis is the main reason why plant and animal domestication spread so quickly from the Fertile Crescent to

1166-413: A new footing, by explaining the processes by which it can occur, and providing observational evidence that it had done so. Darwin's theory was rapidly accepted by the scientific community and soon became a central axiom of the rapidly developing science of biology. The basis for modern genetics began with the work of Gregor Mendel on peas in 1865, although the significance of his work was not realized at

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1272-406: A new perspective on the fundamental basis of life. Having previously been the realm of gentlemen naturalists, over the 18th, 19th and 20th centuries, zoology became an increasingly professional scientific discipline . Explorer-naturalists such as Alexander von Humboldt investigated the interaction between organisms and their environment, and the ways this relationship depends on geography, laying

1378-709: A process involving the union of a male and female haploid gamete , each gamete formed by meiosis . Ordinarily, gametes produced by separate individuals unite by a process of fertilization to form a diploid zygote that can then develop into a genetically unique individual progeny. However, some animals are also capable, as an alternative reproductive process, to reproduce parthenogenetically. Parthenogenesis has been described in snakes and lizards (see Misplaced Pages Parthenogenesis in squamates ), in amphibians (see Misplaced Pages Parthenogenesis in amphibians ) and in numerous other species (see Misplaced Pages Parthenogenesis ). Generally, meiosis in parthanogenetically reproducing animals occurs by

1484-514: A region of the Near East known as the Fertile Crescent. According to the archaeological record this phenomenon, known as "Neolithic", rapidly expanded from these territories into Europe. However, whether this diffusion was accompanied or not by human migrations is greatly debated. Mitochondrial DNA – a type of maternally inherited DNA located in the cell cytoplasm – was recovered from the remains of Pre-Pottery Neolithic B (PPNB) farmers in

1590-537: A relatively short period of between 20 and 200 years. Some of the pioneering attempts failed at first and crops were abandoned, sometimes to be taken up again and successfully domesticated thousands of years later: rye , tried and abandoned in Neolithic Anatolia , made its way to Europe as weed seeds and was successfully domesticated in Europe, thousands of years after the earliest agriculture. Wild lentils presented

1696-601: A similar process to that in sexually reproducing animals, but the diploid zygote nucleus is generated by the union of two haploid genomes from the same individual rather than from different individuals. Neolithic Revolution The Neolithic Revolution , also known as the First Agricultural Revolution , was the wide-scale transition of many human cultures during the Neolithic period in Afro-Eurasia from

1802-446: A variety of other crops, dating back to 11,000 BP. Two potentially significant economic species, taro ( Colocasia esculenta ) and yam ( Dioscorea sp.), have been identified dating at least to 10,200 calibrated years before present (cal BP). Further evidence of bananas and sugarcane dates to 6,950 to 6,440 BCE. This was at the altitudinal limits of these crops, and it has been suggested that cultivation in more favourable ranges in

1908-453: Is a form of scientific taxonomy . Modern biological classification has its root in the work of Carl Linnaeus , who grouped species according to shared physical characteristics. These groupings have since been revised to improve consistency with the Darwinian principle of common descent . Molecular phylogenetics , which uses nucleic acid sequence as data, has driven many recent revisions and

2014-810: Is a highly resilient crop, able to grow in varied and marginal environments, such as in regions of high altitude and latitude. Archaeobotanical evidence shows that barley had spread throughout Eurasia by 2,000 BCE. To further elucidate the routes by which barley cultivation was spread through Eurasia, genetic analysis was used to determine genetic diversity and population structure in extant barley taxa. Genetic analysis shows that cultivated barley spread through Eurasia via several different routes, which were most likely separated in both time and space. When hunter-gathering began to be replaced by sedentary food production it became more efficient to keep animals close at hand. Therefore, it became necessary to bring animals permanently to their settlements, although in many cases there

2120-657: Is administered by the International Code of Zoological Nomenclature . A merging draft, BioCode, was published in 1997 in an attempt to standardize nomenclature, but has yet to be formally adopted. Vertebrate zoology is the biological discipline that consists of the study of vertebrate animals, that is animals with a backbone , such as fish , amphibians , reptiles , birds and mammals . The various taxonomically oriented disciplines i.e. mammalogy , biological anthropology , herpetology , ornithology , and ichthyology seek to identify and classify species and study

2226-480: Is an integrative field of study, uniting concepts and information from evolutionary biology , taxonomy , ecology , physical geography , geology , paleontology and climatology . The origin of this field of study is widely accredited to Alfred Russel Wallace , a British biologist who had some of his work jointly published with Charles Darwin . Molecular biology studies the common genetic and developmental mechanisms of animals and plants, attempting to answer

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2332-402: Is being studied. For example, what is learned about the physiology of yeast cells can also apply to human cells. The field of animal physiology extends the tools and methods of human physiology to non-human species. Physiology studies how, for example, the nervous , immune , endocrine , respiratory , and circulatory systems function and interact. Developmental biology is the study of

2438-539: Is believed to be the homelands of the early Sino-Tibetan -speakers, associated with the Houli , Peiligang , Cishan , and Xinglongwa cultures , clustered around the Yellow River basin. It was the domestication centre for foxtail millet ( Setaria italica ) and broomcorn millet ( Panicum miliaceum ), with early evidence of domestication approximately 8,000 years ago, and widespread cultivation 7,500 years ago. ( Soybean

2544-406: Is done on both the microscopic and molecular levels for single-celled organisms such as bacteria as well as the specialized cells in multicellular organisms such as humans . Understanding the structure and function of cells is fundamental to all of the biological sciences. The similarities and differences between cell types are particularly relevant to molecular biology. Anatomy considers

2650-452: Is in the lower Yangtze River , believed to be the homelands of pre-Austronesians and associated with the Kauhuqiao , Hemudu , Majiabang , and Songze cultures . It is characterized by typical pre-Austronesian features, including stilt houses, jade carving, and boat technologies. Their diet were also supplemented by acorns , water chestnuts , foxnuts , and pig domestication. The second

2756-605: Is in the middle Yangtze River, believed to be the homelands of the early Hmong-Mien -speakers and associated with the Pengtoushan and Daxi cultures . Both of these regions were heavily populated and had regular trade contacts with each other, as well as with early Austroasiatic speakers to the west, and early Kra-Dai speakers to the south, facilitating the spread of rice cultivation throughout southern China. The millet and rice-farming cultures also first came into contact with each other at around 9,000 to 7,000 BP, resulting in

2862-443: Is likely to continue to do so. Biological classification belongs to the science of zoological systematics . Many scientists now consider the five-kingdom system outdated. Modern alternative classification systems generally start with the three-domain system : Archaea (originally Archaebacteria); Bacteria (originally Eubacteria); Eukaryota (including protists , fungi , plants , and animals ) These domains reflect whether

2968-548: Is limited to Northern Levant and Southern Turkey. A detailed satellite map study of a few archaeological sites in the Baluchistan and Khybar Pakhtunkhwa regions also suggests similarities in early phases of farming with sites in Western Asia. Pottery prepared by sequential slab construction, circular fire pits filled with burnt pebbles, and large granaries are common to both Mehrgarh and many Mesopotamian sites. The postures of

3074-607: Is not customarily used in describing cultures in the Americas. However, a broad similarity exists between Eastern Hemisphere cultures of the Neolithic and cultures in the Americas. Maize (corn), beans and squash were among the earliest crops domesticated in Mesoamerica : squash as early as 6000 BCE, beans no later than 4000 BCE, and maize beginning about 7000 BCE. Potatoes and manioc were domesticated in South America . In what

3180-496: Is now the eastern United States, Native Americans domesticated sunflower , sumpweed and goosefoot c.  2500 BCE . In the highlands of central Mexico, sedentary village life based on farming did not develop until the "formative period" in the second millennium BCE. Evidence of drainage ditches at Kuk Swamp on the borders of the Western and Southern Highlands of Papua New Guinea indicates cultivation of taro and

3286-415: Is partly based on paleontology , which uses the fossil record to answer questions about the mode and tempo of evolution, and partly on the developments in areas such as population genetics and evolutionary theory. Following the development of DNA fingerprinting techniques in the late 20th century, the application of these techniques in zoology has increased the understanding of animal populations. In

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3392-406: Is quantitative, and recently a significant amount of work has been done using computer science techniques such as bioinformatics and computational biology . Molecular genetics , the study of gene structure and function, has been among the most prominent sub-fields of molecular biology since the early 2000s. Other branches of biology are informed by molecular biology, by either directly studying

3498-698: Is reflected in zoology by the presence of animals in Egyptian hieroglyphics. Although the concept of zoology as a single coherent field arose much later, the zoological sciences emerged from natural history reaching back to the biological works of Aristotle and Galen in the ancient Greco-Roman world . In the fourth century BC, Aristotle looked at animals as living organisms, studying their structure, development and vital phenomena. He divided them into two groups: animals with blood, equivalent to our concept of vertebrates , and animals without blood, invertebrates . He spent two years on Lesbos , observing and describing

3604-466: Is the earliest Neolithic site in the north-west Indian subcontinent, dated as early as 8500 BCE. Neolithic domesticated crops in Mehrgarh include more than 90% barley and a small amount of wheat. There is good evidence for the local domestication of barley and the zebu cattle at Mehrgarh, but the wheat varieties are suggested to be of Near-Eastern origin, as the modern distribution of wild varieties of wheat

3710-559: Is the genus, and sapiens the specific epithet, both of them combined make up the species name. When writing the scientific name of an organism, it is proper to capitalize the first letter in the genus and put all of the specific epithet in lowercase. Additionally, the entire term may be italicized or underlined. The dominant classification system is called the Linnaean taxonomy . It includes ranks and binomial nomenclature . The classification, taxonomy , and nomenclature of zoological organisms

3816-614: Is the subfield of biology that studies the evolutionary processes (natural selection, common descent, speciation) that produced the diversity of life on Earth. Evolutionary research is concerned with the origin and descent of species , as well as their change over time, and includes scientists from many taxonomically oriented disciplines. For example, it generally involves scientists who have special training in particular organisms such as mammalogy , ornithology , herpetology , or entomology , but use those organisms as systems to answer general questions about evolution. Evolutionary biology

3922-604: The Aegean have been carbon-dated to c.  6500 BCE at Knossos , Franchthi Cave , and a number of mainland sites in Thessaly . Neolithic groups appear soon afterwards in the Balkans and south-central Europe. The Neolithic cultures of southeastern Europe (the Balkans and the Aegean ) show some continuity with groups in southwest Asia and Anatolia (e.g., Çatalhöyük ). Current evidence suggests that Neolithic material culture

4028-598: The Fertile Crescent , and perhaps 8000 BCE in the Kuk Early Agricultural Site of Papua New Guinea in Melanesia . Everywhere, this transition is associated with a change from a largely nomadic hunter-gatherer way of life to a more settled , agrarian one, with the domestication of various plant and animal species – depending on the species locally available, and influenced by local culture. Archaeological research in 2003 suggests that in some regions, such as

4134-537: The Fertile Crescent . Many grinding stones are found with the early Egyptian Sebilian and Mechian cultures and evidence has been found of a neolithic domesticated crop-based economy dating around 7,000 BP. Unlike the Middle East, this evidence appears as a "false dawn" to agriculture, as the sites were later abandoned, and permanent farming then was delayed until 6,500 BP with the Tasian culture and Badarian culture and

4240-657: The Jōmon Period and published his paper in 1879, which launched Japan's modern archeology . These shell mounds are now part of Ōmori Shell Mounds Garden, open to the public. In 1955, the sites were registered as Japan's Historic Sites . The shells, pottery, and other items that Dr. Morse excavated are mostly stored at the University Museum of the University of Tokyo and are Japan's Important Cultural Property . The Ōmori Shell Mounds are located within five-minutes' walk from

4346-695: The Kachi plain of Balochistan , Pakistan; the site has evidence of farming (wheat and barley) and herding (cattle, sheep and goats). There is strong evidence for causal connections between the Near-Eastern Neolithic and that further east, up to the Indus Valley. There are several lines of evidence that support the idea of connection between the Neolithic in the Near East and in the Indian subcontinent. The prehistoric site of Mehrgarh in Baluchistan (modern Pakistan)

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4452-755: The Levantine corridor that show early evidence of agriculture include Wadi Faynan 16 and Netiv Hagdud . Jacques Cauvin noted that the settlers of Aswad did not domesticate on site, but "arrived, perhaps from the neighbouring Anti-Lebanon , already equipped with the seed for planting" . In the Eastern Fertile Crescent, evidence of cultivation of wild plants has been found in Choga Gholan in Iran dated to 12,000 BP, with domesticated emmer wheat appearing in 9,800 BP, suggesting there may have been multiple regions in

4558-767: The Near East and then compared to available data from other Neolithic populations in Europe and also to modern populations from South Eastern Europe and the Near East. The obtained results show that substantial human migrations were involved in the Neolithic spread and suggest that the first Neolithic farmers entered Europe following a maritime route through Cyprus and the Aegean Islands . The earliest Neolithic sites in South Asia are Bhirrana in Haryana dated to 7570–6200  BCE , and Mehrgarh , dated to between 6500 and 5500 BP, in

4664-550: The Neolithic package , provided the basis for centralized administrations and political structures, hierarchical ideologies , depersonalized systems of knowledge (e.g. writing ), densely populated settlements, specialization and division of labour , more trade , the development of non-portable art and architecture , and greater property ownership. The earliest known civilization developed in Sumer in southern Mesopotamia ( c.  6,500 BP ); its emergence also heralded

4770-545: The anatomy of different groups. It is closely related to evolutionary biology and phylogeny (the evolution of species). Physiology studies the mechanical, physical, and biochemical processes of living organisms by attempting to understand how all of the structures function as a whole. The theme of "structure to function" is central to biology. Physiological studies have traditionally been divided into plant physiology and animal physiology , but some principles of physiology are universal, no matter what particular organism

4876-404: The genes of domesticated plants, he preferred theories of a single, or at most a very small number of domestication events for each taxon that spread in an arc from the Levantine corridor around the Fertile Crescent and later into Europe. Gordon Hillman and Stuart Davies carried out experiments with varieties of wild wheat to show that the process of domestication would have occurred over

4982-531: The geological epoch of the Holocene 11,700 years ago, after the end of the last Ice Age. It was humankind's first historically verifiable transition to agriculture. The Neolithic Revolution greatly narrowed the diversity of foods available, resulting in a decrease in the quality of human nutrition compared with that obtained previously from foraging , but because food production became more efficient, it released humans to invest their efforts in other activities and

5088-550: The proximate causes of the behavior, the developmental history of the organism, the survival value and phylogeny of the behavior? Another area of study is animal cognition , which uses laboratory experiments and carefully controlled field studies to investigate an animal's intelligence and learning. Biogeography studies the spatial distribution of organisms on the Earth , focusing on topics like dispersal and migration , plate tectonics , climate change , and cladistics . It

5194-413: The structure , embryology , classification , habits , and distribution of all animals, both living and extinct , and how they interact with their ecosystems . Zoology is one of the primary branches of biology . The term is derived from Ancient Greek ζῷον , zōion ('animal'), and λόγος , logos ('knowledge', 'study'). Although humans have always been interested in the natural history of

5300-399: The taxonomist . Originally it was thought that species were immutable, but with the arrival of Darwin's theory of evolution, the field of cladistics came into being, studying the relationships between the different groups or clades . Systematics is the study of the diversification of living forms, the evolutionary history of a group is known as its phylogeny , and the relationship between

5406-542: The 16th century. During the Renaissance and early modern period, zoological thought was revolutionized in Europe by a renewed interest in empiricism and the discovery of many novel organisms. Prominent in this movement were Andreas Vesalius and William Harvey , who used experimentation and careful observation in physiology , and naturalists such as Carl Linnaeus , Jean-Baptiste Lamarck , and Buffon who began to classify

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5512-470: The 1980s, developmental biology re-entered evolutionary biology from its initial exclusion from the modern synthesis through the study of evolutionary developmental biology . Related fields often considered part of evolutionary biology are phylogenetics , systematics , and taxonomy . Ethology is the scientific and objective study of animal behavior under natural conditions, as opposed to behaviorism , which focuses on behavioral response studies in

5618-689: The 1st millennium CE, they also colonized Madagascar and the Comoros , bringing Southeast Asian food plants, including rice, to East Africa . On the African continent, three areas have been identified as independently developing agriculture: the Ethiopian highlands , the Sahel and West Africa . By contrast, Agriculture in the Nile River Valley is thought to have developed from the original Neolithic Revolution in

5724-600: The Ethiopian highlands. Crops domesticated in the Sahel region include sorghum and pearl millet . The kola nut was first domesticated in West Africa. Other crops domesticated in West Africa include African rice , yams and the oil palm . Agriculture spread to Central and Southern Africa in the Bantu expansion during the 1st millennium BCE to 1st millennium CE. The term "Neolithic"

5830-549: The Fertile Crescent where cereal domestication evolved roughly contemporaneously. The Heavy Neolithic Qaraoun culture has been identified at around fifty sites in Lebanon around the source springs of the River Jordan , but never reliably dated. In his book Guns, Germs, and Steel , Jared Diamond argues that the vast continuous east–west stretch of temperate climatic zones of Eurasia and North Africa gave peoples living there

5936-563: The Neolithic advanced from one patch of fertile alluvial soil to another, bypassing mountainous areas. Analysis of radiocarbon dates show clearly that Mesolithic and Neolithic populations lived side by side for as much as a millennium in many parts of Europe, especially in the Iberian peninsula and along the Atlantic coast. The spread of the Neolithic from the Near East Neolithic to Europe

6042-650: The Southeast Asian peninsula, the transition from hunter-gatherer to agriculturalist was not linear, but region-specific. Once agriculture started gaining momentum, around 9000 BP, human activity resulted in the selective breeding of cereal grasses (beginning with emmer , einkorn and barley ), and not simply of those that favoured greater caloric returns through larger seeds. Plants with traits such as small seeds or bitter taste were seen as undesirable. Plants that rapidly shed their seeds on maturity tended not to be gathered at harvest, therefore not stored and not seeded

6148-449: The animals and plants, considering the adaptations of different organisms and the function of their parts. Four hundred years later, Roman physician Galen dissected animals to study their anatomy and the function of the different parts, because the dissection of human cadavers was prohibited at the time. This resulted in some of his conclusions being false, but for many centuries it was considered heretical to challenge any of his views, so

6254-808: The animals they saw around them, and used this knowledge to domesticate certain species, the formal study of zoology can be said to have originated with Aristotle . He viewed animals as living organisms, studied their structure and development, and considered their adaptations to their surroundings and the function of their parts. Modern zoology has its origins during the Renaissance and early modern period, with Carl Linnaeus , Antonie van Leeuwenhoek , Robert Hooke , Charles Darwin , Gregor Mendel and many others. The study of animals has largely moved on to deal with form and function, adaptations, relationships between groups, behaviour and ecology. Zoology has increasingly been subdivided into disciplines such as classification , physiology , biochemistry and evolution . With

6360-561: The arrival of crops and animals from the Near East. Bananas and plantains , which were first domesticated in Southeast Asia , most likely Papua New Guinea , were re-domesticated in Africa possibly as early as 5,000 years ago. Asian yams and taro were also cultivated in Africa. The most famous crop domesticated in the Ethiopian highlands is coffee . In addition, khat , ensete , noog , teff and finger millet were also domesticated in

6466-729: The beginning of the Bronze Age . The relationship of the aforementioned Neolithic characteristics to the onset of agriculture, their sequence of emergence, and their empirical relation to each other at various Neolithic sites remains the subject of academic debate. It is usually understood to vary from place to place, rather than being the outcome of universal laws of social evolution . Prehistoric hunter-gatherers had different subsistence requirements and lifestyles from agriculturalists. Hunter-gatherers were often highly mobile and migratory, living in temporary shelters and in small tribal groups, and having limited contact with outsiders. Their diet

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6572-427: The cells have nuclei or not, as well as differences in the chemical composition of the cell exteriors. Further, each kingdom is broken down recursively until each species is separately classified. The order is: Domain ; kingdom ; phylum ; class ; order ; family ; genus ; species . The scientific name of an organism is generated from its genus and species. For example, humans are listed as Homo sapiens . Homo

6678-434: The clades can be shown diagrammatically in a cladogram . Although someone who made a scientific study of animals would historically have described themselves as a zoologist, the term has come to refer to those who deal with individual animals, with others describing themselves more specifically as physiologists, ethologists, evolutionary biologists, ecologists, pharmacologists, endocrinologists or parasitologists. Although

6784-514: The determination of the double helical structure of the DNA molecule by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The study of systematics was transformed as DNA sequencing elucidated the degrees of affinity between different organisms. Zoology is the branch of science dealing with animals . A species can be defined as

6890-643: The discovery of the structure of DNA by Francis Crick and James Watson in 1953, the realm of molecular biology opened up, leading to advances in cell biology , developmental biology and molecular genetics . The history of zoology traces the study of the animal kingdom from ancient to modern times. Prehistoric people needed to study the animals and plants in their environment to exploit them and survive. Cave paintings, engravings and sculptures in France dating back 15,000 years show bison, horses, and deer in carefully rendered detail. Similar images from other parts of

6996-456: The diversity of life and the fossil record , as well as studying the development and behavior of organisms. Antonie van Leeuwenhoek did pioneering work in microscopy and revealed the previously unknown world of microorganisms , laying the groundwork for cell theory . van Leeuwenhoek's observations were endorsed by Robert Hooke ; all living organisms were composed of one or more cells and could not generate spontaneously. Cell theory provided

7102-463: The earliest evidence for advanced planning of plants for food consumption and suggests that humans at Ohalo II processed the grain before consumption. Tell Aswad is the oldest site of agriculture, with domesticated emmer wheat dated to 10,800 BP. Soon after came hulled, two-row barley – found domesticated earliest at Jericho in the Jordan valley and at Iraq ed-Dubb in Jordan . Other sites in

7208-663: The first region to domesticate the dromedary . Henri Fleisch discovered and termed the Shepherd Neolithic flint industry from the Bekaa Valley in Lebanon and suggested that it could have been used by the earliest nomadic shepherds . He dated this industry to the Epipaleolithic or Pre-Pottery Neolithic as it is evidently not Paleolithic , Mesolithic or even Pottery Neolithic . The presence of these animals gave

7314-420: The following season; successive years of harvesting spontaneously selected for strains that retained their edible seeds longer. Daniel Zohary identified several plant species as "pioneer crops" or Neolithic founder crops . He highlighted the importance of wheat, barley and rye, and suggested that domestication of flax , peas , chickpeas , bitter vetch and lentils came a little later. Based on analysis of

7420-404: The forms of macroscopic structures such as organs and organ systems. It focuses on how organs and organ systems work together in the bodies of humans and other animals, in addition to how they work independently. Anatomy and cell biology are two studies that are closely related, and can be categorized under "structural" studies. Comparative anatomy is the study of similarities and differences in

7526-431: The foundations for biogeography , ecology and ethology . Naturalists began to reject essentialism and consider the importance of extinction and the mutability of species . These developments, as well as the results from embryology and paleontology , were synthesized in the 1859 publication of Charles Darwin 's theory of evolution by natural selection ; in this Darwin placed the theory of organic evolution on

7632-473: The health of early agriculturalists and their domesticated livestock would have been increased numbers of parasites and disease-bearing pests associated with human waste and contaminated food and water supplies. Fertilizers and irrigation may have increased crop yields but also would have promoted proliferation of insects and bacteria in the local environment while grain storage attracted additional insects and rodents . The term 'neolithic revolution'

7738-555: The human population. Selectively propagated figs , wild barley and wild oats were cultivated at the early Neolithic site of Gilgal I , where in 2006 archaeologists found caches of seeds of each in quantities too large to be accounted for even by intensive gathering , at strata datable to c. 11,000 years ago. Some of the plants tried and then abandoned during the Neolithic period in the Ancient Near East, at sites like Gilgal, were later successfully domesticated in other parts of

7844-617: The impetus of the Austronesian expansion which started with the migration of the Austronesian-speakers from Taiwan to the Philippines at around 5,000 BP. Austronesians carried rice cultivation technology to Island Southeast Asia along with other domesticated species. The new tropical island environments also had new food plants that they exploited. They carried useful plants and animals during each colonization voyage, resulting in

7950-476: The interactions of molecules in their own right such as in cell biology and developmental biology , or indirectly, where molecular techniques are used to infer historical attributes of populations or species , as in fields in evolutionary biology such as population genetics and phylogenetics . There is also a long tradition of studying biomolecules "from the ground up", or molecularly, in biophysics . Animals generally reproduce by sexual reproduction ,

8056-404: The largest group of organisms in which any two individuals of the appropriate sex can produce fertile offspring; about 1.5 million species of animal have been described and it has been estimated that as many as 8 million animal species may exist. An early necessity was to identify the organisms and group them according to their characteristics, differences and relationships, and this is the field of

8162-588: The lowlands may have been even earlier. CSIRO has found evidence that taro was introduced into the Solomon Islands for human use, from 28,000 years ago, making taro the earliest cultivated crop in the world. It seems to have resulted in the spread of the Trans–New Guinea languages from New Guinea east into the Solomon Islands and west into Timor and adjacent areas of Indonesia . This seems to confirm

8268-491: The north entrance of Ōmori Railway Station of the JR Keihin–Tōhoku Line . 35°35′26.8″N 139°43′45.0″E  /  35.590778°N 139.729167°E  / 35.590778; 139.729167 Zoologist Zoology ( UK : / z u ˈ ɒ l ə dʒ i / zoo- OL -ə-jee , US : / z oʊ ˈ ɒ l ə dʒ i / zoh- OL -ə-jee ) is the scientific study of animals . Its studies include

8374-410: The population expanded and communities developed specialized workers and more advanced tools. The process was not as linear as was once thought, but a more complicated effort, which was undertaken by different human populations in different regions in many different ways. One of the world's most important crops, barley , was domesticated in the Near East around 11,000 years ago (c. 9,000 BCE). Barley

8480-517: The processes by which animals and plants reproduce and grow. The discipline includes the study of embryonic development , cellular differentiation , regeneration , asexual and sexual reproduction , metamorphosis , and the growth and differentiation of stem cells in the adult organism. Development of both animals and plants is further considered in the articles on evolution , population genetics , heredity , genetic variability , Mendelian inheritance , and reproduction . Evolutionary biology

8586-401: The production of surplus food. Other developments that are found very widely during this era are the domestication of animals , pottery , polished stone tools, and rectangular houses. In many regions, the adoption of agriculture by prehistoric societies caused episodes of rapid population growth, a phenomenon known as the Neolithic demographic transition . These developments, sometimes called

8692-493: The questions regarding the mechanisms of genetic inheritance and the structure of the gene . In 1953, James Watson and Francis Crick described the structure of DNA and the interactions within the molecule, and this publication jump-started research into molecular biology and increased interest in the subject. While researchers practice techniques specific to molecular biology, it is common to combine these with methods from genetics and biochemistry . Much of molecular biology

8798-540: The rapid introduction of domesticated and semi-domesticated species throughout Oceania . They also came into contact with the early agricultural centres of Papuan -speaking populations of New Guinea as well as the Dravidian -speaking regions of South India and Sri Lanka by around 3,500 BP. They acquired further cultivated food plants like bananas and pepper from them, and in turn introduced Austronesian technologies like wetland cultivation and outrigger canoes . During

8904-429: The region a large advantage in cultural and economic development. As the climate in the Middle East changed and became drier, many of the farmers were forced to leave, taking their domesticated animals with them. It was this massive emigration from the Middle East that later helped distribute these animals to the rest of Afroeurasia . This emigration was mainly on an east–west axis of similar climates, as crops usually have

9010-609: The rest of Eurasia and North Africa, while it did not reach through the north–south axis of Africa to reach the Mediterranean climates of South Africa , where temperate crops were successfully imported by ships in the last 500 years. Similarly, the African Zebu of central Africa and the domesticated bovines of the fertile-crescent – separated by the dry sahara desert – were not introduced into each other's region. Use-wear analysis of five glossed flint blades found at Ohalo II ,

9116-494: The results of his own investigations. His general approach was surprisingly modern, and he wrote, "For it is [the task] of natural science not simply to accept what we are told but to inquire into the causes of natural things." An early pioneer was Conrad Gessner , whose monumental 4,500-page encyclopedia of animals, Historia animalium , was published in four volumes between 1551 and 1558. In Europe, Galen's work on anatomy remained largely unsurpassed and unchallenged up until

9222-509: The rise of civilizations and technological evolutions . However, higher population and food abundance did not necessarily correlate with improved health. Reliance on a very limited variety of staple crops can adversely affect health even while making it possible to feed more people. Maize is deficient in certain essential amino acids ( lysine and tryptophan ) and is a poor source of iron . The phytic acid it contains may inhibit nutrient absorption . Other factors that likely affected

9328-503: The rise of the germ theory of disease , though the mechanism of inheritance remained a mystery. In the early 20th century, the rediscovery of Mendel's work led to the rapid development of genetics , and by the 1930s the combination of population genetics and natural selection in the modern synthesis created evolutionary biology . Research in cell biology is interconnected to other fields such as genetics, biochemistry , medical microbiology , immunology , and cytochemistry . With

9434-481: The speed of 0.6–1.3 km/yr (at 95% confidence level). Since the original human expansions out of Africa 200,000 years ago, different prehistoric and historic migration events have taken place in Europe. Considering that the movement of the people implies a consequent movement of their genes, it is possible to estimate the impact of these migrations through the genetic analysis of human populations. Agricultural and husbandry practices originated 10,000 years ago in

9540-499: The structures and mechanisms specific to those groups. The rest of the animal kingdom is dealt with by invertebrate zoology , a vast and very diverse group of animals that includes sponges , echinoderms , tunicates , worms , molluscs , arthropods and many other phyla , but single-celled organisms or protists are not usually included. Cell biology studies the structural and physiological properties of cells , including their behavior , interactions, and environment . This

9646-529: The study of anatomy stultified. During the post-classical era , Middle Eastern science and medicine was the most advanced in the world, integrating concepts from Ancient Greece, Rome, Mesopotamia and Persia as well as the ancient Indian tradition of Ayurveda , while making numerous advances and innovations. In the 13th century, Albertus Magnus produced commentaries and paraphrases of all Aristotle's works; his books on topics like botany , zoology, and minerals included information from ancient sources, but also

9752-565: The study of animal life is ancient, its scientific incarnation is relatively modern. This mirrors the transition from natural history to biology at the start of the 19th century. Since Hunter and Cuvier , comparative anatomical study has been associated with morphography , shaping the modern areas of zoological investigation: anatomy , physiology , histology , embryology , teratology and ethology . Modern zoology first arose in German and British universities. In Britain, Thomas Henry Huxley

9858-454: The theories of Carl Sauer who, in "Agricultural Origins and Dispersals", suggested as early as 1952 that this region was a centre of early agriculture. Archaeologists trace the emergence of food-producing societies in the Levantine region of southwest Asia at the close of the last glacial period around 12,000 BCE, and developed into a number of regionally distinctive cultures by the eighth millennium BCE. Remains of food-producing societies in

9964-433: The time. Darwin gave a new direction to morphology and physiology , by uniting them in a common biological theory: the theory of organic evolution. The result was a reconstruction of the classification of animals upon a genealogical basis, fresh investigation of the development of animals, and early attempts to determine their genetic relationships. The end of the 19th century saw the fall of spontaneous generation and

10070-437: The world illustrated mostly the animals hunted for food and the savage animals. The Neolithic Revolution , which is characterized by the domestication of animals , continued throughout Antiquity. Ancient knowledge of wildlife is illustrated by the realistic depictions of wild and domestic animals in the Near East, Mesopotamia, and Egypt, including husbandry practices and techniques, hunting and fishing. The invention of writing

10176-532: The world. Once early farmers perfected their agricultural techniques like irrigation (traced as far back as the 6th millennium BCE in Khuzistan ), their crops yielded surpluses that needed storage. Most hunter-gatherers could not easily store food for long due to their migratory lifestyle, whereas those with a sedentary dwelling could store their surplus grain. Eventually granaries were developed that allowed villages to store their seeds longer. So with more food,

10282-612: Was broomcorn millet , domesticated in East Asia. The earliest evidence of cheese -making dates to 5500 BCE in Kujawy , Poland . The diffusion across Europe, from the Aegean to Britain, took about 2,500 years (8500–6000 BP). The Baltic region was penetrated a bit later, around 5500 BP, and there was also a delay in settling the Pannonian plain . In general, colonization shows a "saltatory" pattern, as

10388-419: Was a distinction between relatively sedentary farmers and nomadic herders. The animals' size, temperament, diet, mating patterns, and life span were factors in the desire and success in domesticating animals. Animals that provided milk, such as cows and goats, offered a source of protein that was renewable and therefore quite valuable. The animal's ability as a worker (for example ploughing or towing), as well as

10494-502: Was a prominent figure. His ideas were centered on the morphology of animals. Many consider him the greatest comparative anatomist of the latter half of the 19th century. Similar to Hunter , his courses were composed of lectures and laboratory practical classes in contrast to the previous format of lectures only. Scientific classification in zoology , is a method by which zoologists group and categorize organisms by biological type , such as genus or species . Biological classification

10600-557: Was also domesticated in northern China 4,500 years ago. Orange and peach also originated in China, being cultivated c.  2500 BCE . ) The agricultural centres in southern China are clustered around the Yangtze River basin. Rice was domesticated in this region, together with the development of paddy field cultivation, between 13,500 and 8,200 years ago. There are two possible centres of domestication for rice. The first

10706-488: Was first studied quantitatively in the 1970s, when a sufficient number of Carbon 14 age determinations for early Neolithic sites had become available. In 1973, Ammerman and Cavalli-Sforza discovered a linear relationship between the age of an Early Neolithic site and its distance from the conventional source in the Near East ( Jericho ), demonstrating that the Neolithic spread at an average speed of about 1 km/yr. More recent studies (2005) confirm these results and yield

10812-643: Was introduced to Europe via western Anatolia. All Neolithic sites in Europe contain ceramics , and contain the plants and animals domesticated in Southwest Asia: einkorn , emmer , barley , lentils , pigs , goats , sheep , and cattle . Genetic data suggest that no independent domestication of animals took place in Neolithic Europe, and that all domesticated animals were originally domesticated in Southwest Asia. The only domesticate not from Southwest Asia

10918-469: Was invented by V. Gordon Childe in his book Man Makes Himself (1936). Childe introduced it as the first in a series of agricultural revolutions in Middle Eastern history, calling it a "revolution" to denote its significance, the degree of change to communities adopting and refining agricultural practices. The beginning of this process in different regions has been dated from 10,000 to 8,000 BCE in

11024-517: Was the centre of domestication for three cereals (einkorn wheat, emmer wheat and barley), four legumes (lentil, pea, bitter vetch and chickpea), and flax. Domestication was a slow process that unfolded across multiple regions, and was preceded by centuries if not millennia of pre-domestication cultivation. Finds of large quantities of seeds and a grinding stone at the Epipalaeolithic site of Ohalo II , dating to around 19,400 BP, has shown some of

11130-680: Was thus "ultimately necessary to the rise of modern civilization by creating the foundation for the later process of industrialization and sustained economic growth". The Neolithic Revolution involved much more than the adoption of a limited set of food-producing techniques. During the next millennia, it transformed the small and mobile groups of hunter-gatherers that had hitherto dominated human prehistory into sedentary (non- nomadic ) societies based in built-up villages and towns. These societies radically modified their natural environment by means of specialized food-crop cultivation, with activities such as irrigation and deforestation which allowed

11236-794: Was well-balanced though heavily dependent on what the environment could provide each season. In contrast, because the surplus and plannable supply of food provided by agriculture made it possible to support larger population groups, agriculturalists lived in more permanent dwellings in more densely populated settlements than what could be supported by a hunter-gatherer lifestyle. The agricultural communities' seasonal need to plan and coordinate resource and manpower encouraged division of labour , which gradually led to specialization of labourers and complex societies . The subsequent development of trading networks to exchange surplus commodities and services brought agriculturalists into contact with outside groups, which promoted cultural exchanges that led to

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