Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.
92-617: Montanazhdarcho is a genus of azhdarchoid pterosaur from the Late Cretaceous period ( Campanian stage) of what is now the state of Montana , United States . Montanazhdarcho is known from only one species, M. minor . The holotype specimen, MOR 691 ( Museum of the Rockies ), was found by Robert W. Harmon in Glacier County , in the territory of the Blackfoot , in sandstone of
184-542: A "disagreement" with Rudbeck's wife and had to move out of his mentor's house; his relationship with Rudbeck did not appear to suffer. That Christmas, Linnaeus returned home to Stenbrohult to visit his parents for the first time in about three years. His mother had disapproved of his failing to become a priest, but she was pleased to learn he was teaching at the University. During a visit with his parents, Linnaeus told them about his plan to travel to Lapland ; Rudbeck had made
276-479: A catalogue of the botanical holdings in the herbarium and botanical garden of Hartekamp. He wrote it in nine months (completed in July 1737), but it was not published until 1738. It contains the first use of the name Nepenthes , which Linnaeus used to describe a genus of pitcher plants . Linnaeus stayed with Clifford at Hartekamp until 18 October 1737 (new style), when he left the house to return to Sweden. Illness and
368-466: A child's talents than develop them". Two years after his tutoring had begun, he was sent to the Lower Grammar School at Växjö in 1717. Linnaeus rarely studied, often going to the countryside to look for plants. At some point, his father went to visit him and, after hearing critical assessments by his preceptors, he decided to put the youth as an apprentice to some honest cobbler. He reached
460-643: A collector of natural history, and to see his cabinet , as well as to visit the Chelsea Physic Garden and its keeper, Philip Miller . He taught Miller about his new system of subdividing plants, as described in Systema Naturae . At first, Miller was reluctant to use the new binomial nomenclature , preferring instead the classifications of Joseph Pitton de Tournefort and John Ray . Nevertheless, Linnaeus applauded Miller's Gardeners Dictionary . The conservative Miller actually retained in his dictionary
552-430: A copy of Sir Hans Sloane's Natural History of Jamaica , a rare book, if he let Linnaeus stay with him, and Burman accepted. On 24 September 1735, Linnaeus moved to Hartekamp to become personal physician to Clifford, and curator of Clifford's herbarium. He was paid 1,000 florins a year, with free board and lodging. Though the agreement was only for a winter of that year, Linnaeus practically stayed there until 1738. It
644-555: A curriculum designed for boys preparing for the priesthood. In the last year at the gymnasium, Linnaeus's father visited to ask the professors how his son's studies were progressing; to his dismay, most said that the boy would never become a scholar. Rothman believed otherwise, suggesting Linnaeus could have a future in medicine. The doctor offered to have Linnaeus live with his family in Växjö and to teach him physiology and botany. Nils accepted this offer. Rothman showed Linnaeus that botany
736-560: A director of the Dutch East India Company and the owner of a rich botanical garden at the estate of Hartekamp in Heemstede . Clifford was very impressed with Linnaeus's ability to classify plants, and invited him to become his physician and superintendent of his garden. Linnaeus had already agreed to stay with Burman over the winter, and could thus not accept immediately. However, Clifford offered to compensate Burman by offering him
828-529: A dissertation, written back in Sweden, entitled Dissertatio medica inauguralis in qua exhibetur hypothesis nova de febrium intermittentium causa , in which he laid out his hypothesis that malaria arose only in areas with clay-rich soils. Although he failed to identify the true source of disease transmission, (i.e., the Anopheles mosquito ), he did correctly predict that Artemisia annua ( wormwood ) would become
920-456: A family, he received permission to marry his fiancée, Sara Elisabeth Moræa. Their wedding was held 26 June 1739. Seventeen months later, Sara gave birth to their first son, Carl . Two years later, a daughter, Elisabeth Christina , was born, and the subsequent year Sara gave birth to Sara Magdalena, who died when 15 days old. Sara and Linnaeus would later have four other children: Lovisa, Sara Christina, Johannes and Sophia. In May 1741, Linnaeus
1012-407: A genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of
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#17329273472391104-643: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as
1196-621: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,
1288-476: A naturalist." Linnaeus has been called Princeps botanicorum (Prince of Botanists) and "The Pliny of the North". He is also considered one of the founders of modern ecology . In botany, the abbreviation L. is used to indicate Linnaeus as the authority for a species' name. In zoology, the abbreviation Linnaeus is generally used; the abbreviations L., Linnæus and Linné are also used. In older publications,
1380-472: A new benefactor, Olof Celsius , who was a professor of theology and an amateur botanist. He received Linnaeus into his home and allowed him use of his library, which was one of the richest botanical libraries in Sweden. In 1729, Linnaeus wrote a thesis, Praeludia Sponsaliorum Plantarum on plant sexual reproduction . This attracted the attention of Rudbeck; in May 1730, he selected Linnaeus to give lectures at
1472-564: A number of pre-Linnaean binomial signifiers discarded by Linnaeus but which have been retained by modern botanists. He only fully changed to the Linnaean system in the edition of The Gardeners Dictionary of 1768. Miller ultimately was impressed, and from then on started to arrange the garden according to Linnaeus's system. Linnaeus also travelled to Oxford University to visit the botanist Johann Jacob Dillenius . He failed to make Dillenius publicly fully accept his new classification system, though
1564-453: A partial left wing, lacking the outer three wing finger phalanges, a complete shoulder girdle, a crushed cervical vertebra and two fragments of the symphysis of the mandible . The jaws were edentulous, i.e. they lacked teeth. Montanazhdarcho was by the authors assigned to the Azhdarchidae , mainly based on the elongated form of the neck vertebra. Compared to other azhdarchids however, it
1656-611: A physician, and thus to make it possible to support a family. Once again, Linnaeus found a patron; he became acquainted with Count Carl Gustav Tessin , who helped him get work as a physician at the Admiralty. During this time in Stockholm, Linnaeus helped found the Royal Swedish Academy of Science ; he became the first Praeses of the academy by drawing of lots. Because his finances had improved and were now sufficient to support
1748-556: A practical way, making this the first proto-modern Flora . The account covered 534 species, used the Linnaean classification system and included, for the described species, geographical distribution and taxonomic notes. It was Augustin Pyramus de Candolle who attributed Linnaeus with Flora Lapponica as the first example in the botanical genre of Flora writing. Botanical historian E. L. Greene described Flora Lapponica as "the most classic and delightful" of Linnaeus's works. It
1840-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for
1932-566: A small group of students to Dalarna . Funded by the Governor of Dalarna, the expedition was to catalogue known natural resources and discover new ones, but also to gather intelligence on Norwegian mining activities at Røros . His relations with Nils Rosén having worsened, Linnaeus accepted an invitation from Claes Sohlberg, son of a mining inspector, to spend the Christmas holiday in Falun , where Linnaeus
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#17329273472392024-462: A source of antimalarial medications. Within two weeks he had completed his oral and practical examinations and was awarded a doctoral degree. That summer Linnaeus reunited with Peter Artedi , a friend from Uppsala with whom he had once made a pact that should either of the two predecease the other, the survivor would finish the decedent's work. Ten weeks later, Artedi drowned in the canals of Amsterdam , leaving behind an unfinished manuscript on
2116-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;
2208-452: A thesis along with Frederick Lindberg, a physician student, based on their experiences. In the tradition of the period, this dissertation was essentially an idea of the presiding reviewer ( prases ) expounded upon by the student. Linnaeus's dissertation was translated into French by J. E. Gilibert in 1770 as La Nourrice marâtre, ou Dissertation sur les suites funestes du nourrisage mercénaire . Linnaeus suggested that children might absorb
2300-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and
2392-596: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms
2484-606: Is known as the "father of modern taxonomy ". Many of his writings were in Latin; his name is rendered in Latin as Carolus Linnæus and, after his 1761 ennoblement , as Carolus a Linné . Linnaeus was the son of a curate and was born in Råshult , in the countryside of Småland , southern Sweden . He received most of his higher education at Uppsala University and began giving lectures in botany there in 1730. He lived abroad between 1735 and 1738, where he studied and also published
2576-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,
2668-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with
2760-446: The Beast of Revelation . Even at the risk of incurring the mayor's wrath, Linnaeus made his observations public, dashing the mayor's dreams of selling the hydra for an enormous sum. Linnaeus and Sohlberg were forced to flee from Hamburg. Linnaeus began working towards his degree as soon as he reached Harderwijk , a university known for awarding degrees in as little as a week. He submitted
2852-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to
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2944-799: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;
3036-490: The curate of the small village of Stenbrohult in Småland. Christina was the daughter of the rector of Stenbrohult, Samuel Brodersonius. A year after Linnaeus's birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the rectory from the curate's house. Even in his early years, Linnaeus seemed to have a liking for plants, flowers in particular. Whenever he
3128-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,
3220-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,
3312-469: The Azhdarchidae again, though placed in the basalmost position. Their phylogenetic analysis is shown below: Montanazhdarcho Azhdarcho Phosphatodraco Aralazhdarcho Eurazhdarcho aff. Quetzalcoatlus Zhejiangopterus Arambourgiania Hatzegopteryx Quetzalcoatlus [REDACTED] [REDACTED] [REDACTED] Genus The composition of
3404-567: The Scottish doctor Isaac Lawson , the manuscript was published as Systema Naturae (1735). Linnaeus became acquainted with one of the most respected physicians and botanists in the Netherlands, Herman Boerhaave , who tried to convince Linnaeus to make a career there. Boerhaave offered him a journey to South Africa and America, but Linnaeus declined, stating he would not stand the heat. Instead, Boerhaave convinced Linnaeus that he should visit
3496-547: The Swedish province of Västergötland . He set out from Uppsala on 12 June and returned on 11 August. On the expedition his primary companion was Erik Gustaf Lidbeck, a student who had accompanied him on his previous journey. Linnaeus described his findings from the expedition in the book Wästgöta-Resa , published the next year. After he returned from the journey, the Government decided Linnaeus should take on another expedition to
3588-490: The University although the young man was only a second-year student. His lectures were popular, and Linnaeus often addressed an audience of 300 people. In June, Linnaeus moved from Celsius's house to Rudbeck's to become the tutor of the three youngest of his 24 children. His friendship with Celsius did not wane and they continued their botanical expeditions. Over that winter, Linnaeus began to doubt Tournefort's system of classification and decided to create one of his own. His plan
3680-519: The Upper Two Medicine Formation , a layer about 74 million years old. The genus name Montanazhdarcho was first given informally in 1993 by Kevin Padian , Armand de Ricqlès , and Jack Horner . It was then named formally in 1995 by the same researchers, and was fully described in 2002. The type species is Montanazhdarcho minor . The generic name refers to the state of Montana and to
3772-461: The abbreviation "Linn." is found. Linnaeus's remains constitute the type specimen for the species Homo sapiens following the International Code of Zoological Nomenclature , since the sole specimen that he is known to have examined was himself. Linnaeus was born in the village of Råshult in Småland , Sweden, on 23 May 1707. He was the first child of Nicolaus (Nils) Ingemarsson (who later adopted
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3864-534: The age of 17, Linnaeus had become well acquainted with the existing botanical literature. He remarks in his journal that he "read day and night, knowing like the back of my hand, Arvidh Månsson's Rydaholm Book of Herbs, Tillandz's Flora Åboensis, Palmberg's Serta Florea Suecana, Bromelii's Chloros Gothica and Rudbeckii's Hortus Upsaliensis". Linnaeus entered the Växjö Katedralskola in 1724, where he studied mainly Greek , Hebrew , theology and mathematics ,
3956-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of
4048-442: The botanist Johannes Burman . After his visit, Burman, impressed with his guest's knowledge, decided Linnaeus should stay with him during the winter. During his stay, Linnaeus helped Burman with his Thesaurus Zeylanicus . Burman also helped Linnaeus with the books on which he was working: Fundamenta Botanica and Bibliotheca Botanica . In August 1735, during Linnaeus's stay with Burman, he met George Clifford III ,
4140-424: The class of organisms. Linnaeus published Species Plantarum , the work which is now internationally accepted as the starting point of modern botanical nomenclature , in 1753. The first volume was issued on 24 May, the second volume followed on 16 August of the same year. The book contained 1,200 pages and was published in two volumes; it described over 7,300 species. The same year the king dubbed him knight of
4232-409: The classification of fish. One of the first scientists Linnaeus met in the Netherlands was Johan Frederik Gronovius , to whom Linnaeus showed one of the several manuscripts he had brought with him from Sweden. The manuscript described a new system for classifying plants. When Gronovius saw it, he was very impressed, and offered to help pay for the printing. With an additional monetary contribution by
4324-440: The expedition were later published in Öländska och Gothländska Resa , written in Swedish. Like Flora Lapponica , it contained both zoological and botanical observations, as well as observations concerning the culture in Öland and Gotland. During the summer of 1745, Linnaeus published two more books: Flora Suecica and Fauna Suecica . Flora Suecica was a strictly botanical book, while Fauna Suecica
4416-411: The family name Linnaeus) and Christina Brodersonia. His siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus (who would eventually succeed their father as rector of Stenbrohult and write a manual on beekeeping ), and Emerentia Linnæa. His father taught him Latin as a small child. One of a long line of peasants and priests, Nils was an amateur botanist , a Lutheran minister , and
4508-422: The first edition of his Systema Naturae in the Netherlands. He then returned to Sweden where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on several journeys through Sweden to find and classify plants and animals. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, while publishing several volumes. By the time of his death in 1778, he
4600-544: The flora and fauna in the vicinity of Uppsala. Linnaeus published Philosophia Botanica in 1751. The book contained a complete survey of the taxonomy system he had been using in his earlier works. It also contained information of how to keep a journal on travels and how to maintain a botanical garden. During Linnaeus's time it was normal for upper class women to have wet nurses for their babies. Linnaeus joined an ongoing campaign to end this practice in Sweden and promote breast-feeding by mothers. In 1752 Linnaeus published
4692-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this
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#17329273472394784-727: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,
4876-877: The journey in 1695, but the detailed results of his exploration were lost in a fire seven years afterwards. Linnaeus's hope was to find new plants, animals and possibly valuable minerals. He was also curious about the customs of the native Sami people , reindeer-herding nomads who wandered Scandinavia's vast tundras. In April 1732, Linnaeus was awarded a grant from the Royal Society of Sciences in Uppsala for his journey. Linnaeus began his expedition from Uppsala on 12 May 1732, just before he turned 25. He travelled on foot and horse, bringing with him his journal, botanical and ornithological manuscripts and sheets of paper for pressing plants. Near Gävle he found great quantities of Campanula serpyllifolia , later known as Linnaea borealis ,
4968-577: The kindness of Dutch friends obliged him to stay some months longer in Holland. In May 1738, he set out for Sweden again. On the way home, he stayed in Paris for about a month, visiting botanists such as Antoine de Jussieu . After his return, Linnaeus never again left Sweden. When Linnaeus returned to Sweden on 28 June 1738, he went to Falun , where he entered into an engagement to Sara Elisabeth Moræa . Three months later, he moved to Stockholm to find employment as
5060-628: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,
5152-524: The last year of the Lower School when he was fifteen, which was taught by the headmaster, Daniel Lannerus, who was interested in botany. Lannerus noticed Linnaeus's interest in botany and gave him the run of his garden. He also introduced him to Johan Rothman, the state doctor of Småland and a teacher at Katedralskolan (a gymnasium ) in Växjö. Also a botanist, Rothman broadened Linnaeus's interest in botany and helped him develop an interest in medicine. By
5244-432: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné , was a Swedish biologist and physician who formalised binomial nomenclature , the modern system of naming organisms. He
5336-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus
5428-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this
5520-455: The personality of their wet nurse through the milk. He admired the child care practices of the Lapps and pointed out how healthy their babies were compared to those of Europeans who employed wet nurses. He compared the behaviour of wild animals and pointed out how none of them denied their newborns their breastmilk. It is thought that his activism played a role in his choice of the term Mammalia for
5612-526: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for
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#17329273472395704-455: The quality of its ferruginous water. The journey was successful, and Linnaeus's observations were published the next year in Skånska Resa . In 1750, Linnaeus became rector of Uppsala University, starting a period where natural sciences were esteemed. Perhaps the most important contribution he made during his time at Uppsala was to teach; many of his students travelled to various places in
5796-453: The related species Azhdarcho . The specific name means "the smaller one" in Latin , a reference to the relatively small size in comparison to closely related forms. Montanazhdarcho was a relatively small pterosaur, reaching 2.5 m (8.2 ft) in wingspan. The fossil is largely uncompressed and that of an adult exemplar, as established by a study of the bone by de Ricqlès. It consists of
5888-466: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or
5980-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in
6072-561: The southernmost province Scania . This journey was postponed, as Linnaeus felt too busy. In 1747, Linnaeus was given the title archiater , or chief physician, by the Swedish king Adolf Frederick —a mark of great respect. The same year he was elected member of the Academy of Sciences in Berlin . In the spring of 1749, Linnaeus could finally journey to Scania , again commissioned by the government. With him he brought his student Olof Söderberg. On
6164-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,
6256-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being
6348-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for
6440-744: The twinflower that would become his favourite. He sometimes dismounted on the way to examine a flower or rock and was particularly interested in mosses and lichens , the latter a main part of the diet of the reindeer , a common and economically important animal in Lapland. Linnaeus travelled clockwise around the coast of the Gulf of Bothnia , making major inland incursions from Umeå , Luleå and Tornio . He returned from his six-month-long, over 2,000 kilometres (1,200 mi) expedition in October, having gathered and observed many plants, birds and rocks. Although Lapland
6532-636: The two men remained in correspondence for many years afterwards. Linnaeus dedicated his Critica Botanica to him, as " opus botanicum quo absolutius mundus non-vidit ". Linnaeus would later name a genus of tropical tree Dillenia in his honour. He then returned to Hartekamp, bringing with him many specimens of rare plants. The next year, 1737, he published Genera Plantarum , in which he described 935 genera of plants, and shortly thereafter he supplemented it with Corollarium Generum Plantarum , with another sixty ( sexaginta ) genera. His work at Hartekamp led to another book, Hortus Cliffortianus ,
6624-469: The two professors who taught at the medical faculty at Uppsala: Olof Rudbeck the Younger and Lars Roberg . Although Rudbeck and Roberg had undoubtedly been good professors, by then they were older and not so interested in teaching. Rudbeck no longer gave public lectures, and had others stand in for him. The botany, zoology, pharmacology and anatomy lectures were not in their best state. In Uppsala, Linnaeus met
6716-455: The use of his library, which included many books about botany. He also gave the student free admission to his lectures. In his spare time, Linnaeus explored the flora of Skåne, together with students sharing the same interests. In August 1728, Linnaeus decided to attend Uppsala University on the advice of Rothman, who believed it would be a better choice if Linnaeus wanted to study both medicine and botany. Rothman based this recommendation on
6808-566: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up
6900-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in
6992-539: The way to Scania, he made his last visit to his brothers and sisters in Stenbrohult since his father had died the previous year. The expedition was similar to the previous journeys in most aspects, but this time he was also ordered to find the best place to grow walnut and Swedish whitebeam trees; these trees were used by the military to make rifles. While there, they also visited the Ramlösa mineral spa , where he remarked on
7084-471: The way, the pair stopped in Hamburg , where they met the mayor, who proudly showed them a supposed wonder of nature in his possession: the taxidermied remains of a seven-headed hydra . Linnaeus quickly discovered the specimen was a fake , cobbled together from the jaws and paws of weasels and the skins of snakes. The provenance of the hydra suggested to Linnaeus that it had been manufactured by monks to represent
7176-477: The world to collect botanical samples. Linnaeus called the best of these students his "apostles". His lectures were normally very popular and were often held in the Botanical Garden. He tried to teach the students to think for themselves and not trust anybody, not even him. Even more popular than the lectures were the botanical excursions made every Saturday during summer, where Linnaeus and his students explored
7268-528: The æ ligature, both in handwritten documents and in publications. Carl's patronymic would have been Nilsson, as in Carl Nilsson Linnæus. Linnaeus's father began teaching him basic Latin, religion, and geography at an early age. When Linnaeus was seven, Nils decided to hire a tutor for him. The parents picked Johan Telander, a son of a local yeoman . Linnaeus did not like him, writing in his autobiography that Telander "was better calculated to extinguish
7360-433: Was zoological . Anders Celsius had created the temperature scale named after him in 1742. Celsius's scale was originally inverted compared to the way it is used today, with water boiling at 0 °C and freezing at 100 °C. Linnaeus was the one who inverted the scale to its present usage, in 1745. In the summer of 1746, Linnaeus was once again commissioned by the Government to carry out an expedition, this time to
7452-413: Was a region with limited biodiversity , Linnaeus described about 100 previously unidentified plants. These became the basis of his book Flora Lapponica . However, on the expedition to Lapland, Linnaeus used Latin names to describe organisms because he had not yet developed the binomial system. In Flora Lapponica Linnaeus's ideas about nomenclature and classification were first used in
7544-629: Was a serious subject. He taught Linnaeus to classify plants according to Tournefort's system. Linnaeus was also taught about the sexual reproduction of plants, according to Sébastien Vaillant . In 1727, Linnaeus, age 21, enrolled in Lund University in Skåne . He was registered as Carolus Linnæus , the Latin form of his full name, which he also used later for his Latin publications. Professor Kilian Stobæus , natural scientist, physician and historian, offered Linnaeus tutoring and lodging, as well as
7636-441: Was appointed Professor of Medicine at Uppsala University, first with responsibility for medicine-related matters. Soon, he changed place with the other Professor of Medicine, Nils Rosén, and thus was responsible for the Botanical Garden (which he would thoroughly reconstruct and expand), botany and natural history , instead. In October that same year, his wife and nine-month-old son followed him to live in Uppsala. Ten days after he
7728-482: Was appointed professor, he undertook an expedition to the island provinces of Öland and Gotland with six students from the university to look for plants useful in medicine. They stayed on Öland until 21 June, then sailed to Visby in Gotland. Linnaeus and the students stayed on Gotland for about a month, and then returned to Uppsala. During this expedition, they found 100 previously unrecorded plants. The observations from
7820-448: Was during this expedition that Linnaeus had a flash of insight regarding the classification of mammals. Upon observing the lower jawbone of a horse at the side of a road he was travelling, Linnaeus remarked: "If I only knew how many teeth and of what kind every animal had, how many teats and where they were placed, I should perhaps be able to work out a perfectly natural system for the arrangement of all quadrupeds." In 1734, Linnaeus led
7912-612: Was here that he wrote a book Hortus Cliffortianus , in the preface of which he described his experience as "the happiest time of my life". (A portion of Hartekamp was declared as public garden in April 1956 by the Heemstede local authority, and was named "Linnaeushof". It eventually became, as it is claimed, the biggest playground in Europe. ) In July 1736, Linnaeus travelled to England, at Clifford's expense. He went to London to visit Sir Hans Sloane,
8004-547: Was named Ingemarsson after his father Ingemar Bengtsson. When Nils was admitted to the University of Lund , he had to take on a family name. He adopted the Latinate name Linnæus after a giant linden tree (or lime tree), lind in Swedish, that grew on the family homestead. This name was spelled with the æ ligature . When Carl was born, he was named Carl Linnæus, with his father's family name. The son also always spelled it with
8096-420: Was one of the most acclaimed scientists in Europe. Philosopher Jean-Jacques Rousseau sent him the message: "Tell him I know no greater man on Earth." Johann Wolfgang von Goethe wrote: "With the exception of Shakespeare and Spinoza , I know no one among the no longer living who has influenced me more strongly." Swedish author August Strindberg wrote: "Linnaeus was in reality a poet who happened to become
8188-538: Was permitted to visit the mines. In April 1735, at the suggestion of Sohlberg's father, Linnaeus and Sohlberg set out for the Dutch Republic , where Linnaeus intended to study medicine at the University of Harderwijk while tutoring Sohlberg in exchange for an annual salary. At the time, it was common for Swedes to pursue doctoral degrees in the Netherlands , then a highly revered place to study natural history. On
8280-454: Was small; the fragments of humerus , radius , and carpal suggest an animal with a 2.5 m (8.2 ft) wingspan. Its ulna was longer than the wing metacarpal, which is atypical for azhdarchids. In addition to this, a phylogenetic study in 2015 has disagreed to its phylogenetic position, and reassigned Montanazhdarcho as a non-azhdarchid azhdarchoid. However, in 2018, Nicholas Longrich and colleagues had recovered Montanazhdarcho within
8372-738: Was to divide the plants by the number of stamens and pistils . He began writing several books, which would later result in, for example, Genera Plantarum and Critica Botanica . He also produced a book on the plants grown in the Uppsala Botanical Garden , Adonis Uplandicus . Rudbeck's former assistant, Nils Rosén , returned to the University in March 1731 with a degree in medicine. Rosén started giving anatomy lectures and tried to take over Linnaeus's botany lectures, but Rudbeck prevented that. Until December, Rosén gave Linnaeus private tutoring in medicine. In December, Linnaeus had
8464-414: Was upset, he was given a flower, which immediately calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus and told him their names. Soon Linnaeus was given his own patch of earth where he could grow plants. Carl's father was the first in his ancestry to adopt a permanent surname. Before that, ancestors had used the patronymic naming system of Scandinavian countries: his father
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