70-399: Family-level: Colymbidae Vigors , 1825 (but see text ) Colymbinae Bonaparte , 1831 (but see text ) Urinatores Vieillot , 1818 Urinatoridae Vieillot, 1818 Urinatorides Vieillot, 1818 Genus-level: Colymbus Linnaeus , 1758 (but see text ) Urinator Lacépède , 1799 Loons ( North American English ) or divers ( British / Irish English ) are
140-1054: A dagger (†); groups of uncertain placement are labelled with a question mark (?) and dashed lines (- - - - -). Jawless fishes (118 species: hagfish , lampreys ) [REDACTED] † Thelodonti , † Conodonta , † Anaspida [REDACTED] [REDACTED] [REDACTED] † Galeaspida [REDACTED] † Osteostraci [REDACTED] † Placodermi [REDACTED] † Acanthodii [REDACTED] (>1,100 species: sharks , rays , chimaeras ) [REDACTED] (2 species: coelacanths ) [REDACTED] Dipnoi (6 species: lungfish ) [REDACTED] Tetrapoda (>38,000 species, not considered fish: amphibians, reptiles, birds, mammals) [REDACTED] (14 species: bichirs , reedfish ) [REDACTED] (27 species: sturgeons , paddlefish ) [REDACTED] Ginglymodi (7 species: gars , alligator gars ) [REDACTED] Halecomorphi (2 species: bowfin , eyetail bowfin ) [REDACTED] (>32,000 species) [REDACTED] Fishes (without tetrapods) are
210-579: A paraphyletic group and for this reason, the class Pisces seen in older reference works is no longer used in formal classifications. Traditional classification divides fish into three extant classes (Agnatha, Chondrichthyes, and Osteichthyes), and with extinct forms sometimes classified within those groups, sometimes as their own classes. Fish account for more than half of vertebrate species. As of 2016, there are over 32,000 described species of bony fish, over 1,100 species of cartilaginous fish, and over 100 hagfish and lampreys. A third of these fall within
280-671: A barrister and became a Doctor of Civil Law in 1832. Vigors was a co-founder of the Zoological Society of London in 1826, and its first secretary until 1833. In that year, he founded what became the Royal Entomological Society of London . He was a fellow of the Linnean Society and the Royal Society . He was the author of 40 papers, mostly on ornithology . He described 110 species of birds, enough to rank him among
350-400: A group of aquatic birds found in much of North America and northern Eurasia . All living species of loons are members of the genus Gavia , family Gaviidae and order Gaviiformes . Loons, which are the size of large ducks or small geese , resemble these birds in shape when swimming. Like ducks and geese, but unlike coots (which are Rallidae ) and grebes ( Podicipedidae ),
420-436: A more spherical lens . Their retinas generally have both rods and cones (for scotopic and photopic vision ); many species have colour vision , often with three types of cone. Teleosts can see polarized light ; some such as cyprinids have a fourth type of cone that detects ultraviolet . Amongst jawless fish , the lamprey has well-developed eyes, while the hagfish has only primitive eyespots. Hearing too
490-754: A mostly terrestrial clade of vertebrates that have dominated the top trophic levels in both aquatic and terrestrial ecosystems since the Late Paleozoic , evolved from lobe-finned fish during the Carboniferous , developing air-breathing lungs homologous to swim bladders. Despite the cladistic lineage, tetrapods are usually not considered to be fish, making "fish" a paraphyletic group. Fish have been an important natural resource for humans since prehistoric times, especially as food . Commercial and subsistence fishers harvest fish in wild fisheries or farm them in ponds or in breeding cages in
560-405: A necropsy of a bald eagle found floating on a Maine lake (beside the floating body of a loon chick) found that the eagle had been stabbed through the heart by an adult loon's beak. Biologists , especially from Chapman University , have extensively studied the mating behaviour of the common loon ( G. immer ). Contrary to popular belief, pairs seldom mate for life . Indeed, a typical adult loon
630-419: A protective bony cover or operculum . They are able to oxygenate their gills using muscles in the head. Some 400 species of fish in 50 families can breathe air, enabling them to live in oxygen-poor water or to emerge on to land. The ability of fish to do this is potentially limited by their single-loop circulation, as oxygenated blood from their air-breathing organ will mix with deoxygenated blood returning to
700-441: A sense of touch and of hearing . Blind cave fish navigate almost entirely through the sensations from their lateral line system. Some fish, such as catfish and sharks, have the ampullae of Lorenzini , electroreceptors that detect weak electric currents on the order of millivolt. Vision is an important sensory system in fish. Fish eyes are similar to those of terrestrial vertebrates like birds and mammals, but have
770-433: A spear-shaped bill. Males are larger on average, but relative size is only apparent when the male and female are together. In winter, plumage is dark grey above, with some indistinct lighter mottling on the wings, and a white chin, throat and underside. The specific species can then be distinguished by certain features, such as the size and colour of the head, neck, back and bill. But reliable identification of loons in winter
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#1732851634649840-485: A tail fin, jaws, skin covered with scales , and lays eggs. Each criterion has exceptions, creating a wide diversity in body shape and way of life. For example, some fast-swimming fish are warm-blooded, while some slow-swimming fish have abandoned streamlining in favour of other body shapes. Fish species are roughly divided equally between freshwater and marine (oceanic) ecosystems; there are some 15,200 freshwater species and around 14,800 marine species. Coral reefs in
910-585: A true "land fish" as this worm-like catfish strictly lives among waterlogged leaf litter . Cavefish of multiple families live in underground lakes , underground rivers or aquifers . Like other animals, fish suffer from parasitism . Some species use cleaner fish to remove external parasites. The best known of these are the bluestreak cleaner wrasses of coral reefs in the Indian and Pacific oceans. These small fish maintain cleaning stations where other fish congregate and perform specific movements to attract
980-422: A typical fish is adapted for efficient swimming by alternately contracting paired sets of muscles on either side of the backbone. These contractions form S-shaped curves that move down the body. As each curve reaches the tail fin, force is applied to the water, moving the fish forward. The other fins act as control surfaces like an aircraft's flaps, enabling the fish to steer in any direction. Since body tissue
1050-606: Is also found in tubenoses ( Procellariiformes ) and penguins (Sphenisciformes), both relatives of the loons. Loons are excellent swimmers, using their feet to propel themselves above and under water. However, since their feet are located far back on the body, loons have difficulty walking on land, though they can effectively run short distances to reach water when frightened. Thus, loons avoid coming to land, except for mating and nesting. Loons fly strongly, though they have high wing loading (mass to wing area ratio), which complicates takeoff. Indeed, most species must run upwind across
1120-465: Is an important sensory system in fish. Fish sense sound using their lateral lines and otoliths in their ears, inside their heads. Some can detect sound through the swim bladder. Some fish, including salmon, are capable of magnetoreception ; when the axis of a magnetic field is changed around a circular tank of young fish, they reorient themselves in line with the field. The mechanism of fish magnetoreception remains unknown; experiments in birds imply
1190-431: Is denser than water, fish must compensate for the difference or they will sink. Many bony fish have an internal organ called a swim bladder that allows them to adjust their buoyancy by increasing or decreasing the amount of gas it contains. The scales of fish provide protection from predators at the cost of adding stiffness and weight. Fish scales are often highly reflective; this silvering provides camouflage in
1260-481: Is likely to have several mates during its lifetime because of territorial takeover. Each breeding pair must frequently defend its territory against "floaters" (territory-less adults) trying to evict at least one owner and seize the breeding site. Territories that have produced chicks in the past year are especially prone to takeovers, because nonbreeding loons use chicks as cues to indicate high-quality territories. One-third of all territorial evictions among males result in
1330-422: Is often difficult even for experts – particularly as the smaller immature birds look similar to winter-plumage adults, making size an unreliable means of identification. Gaviiformes are among the few groups of birds in which the young moult into a second coat of down feathers after shedding the first one, rather than growing juvenile feathers with downy tips that wear off, as is typical in many birds. This trait
1400-460: Is quite unrelated to loons and just happens to be another black-and-white seabird which swims and dives for fish. It is not likely that the ancient Romans had much knowledge of loons, as these are limited to more northern latitudes and since the end of the last glacial period seem to have occurred only as rare winter migrants in the Mediterranean region. The term gavia was transferred from
1470-465: Is the biggest part of the brain; it is small in hagfish and lampreys , but very large in mormyrids , processing their electrical sense . The brain stem or myelencephalon controls some muscles and body organs, and governs respiration and osmoregulation . The lateral line system is a network of sensors in the skin which detects gentle currents and vibrations, and senses the motion of nearby fish, whether predators or prey. This can be considered both
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#17328516346491540-567: The Cambrian as small filter feeders ; they continued to evolve through the Paleozoic , diversifying into many forms. The earliest fish with dedicated respiratory gills and paired fins , the ostracoderms , had heavy bony plates that served as protective exoskeletons against invertebrate predators . The first fish with jaws , the placoderms, appeared in the Silurian and greatly diversified during
1610-514: The Devonian , the "Age of Fishes". Bony fish, distinguished by the presence of swim bladders and later ossified endoskeletons , emerged as the dominant group of fish after the end-Devonian extinction wiped out the apex placoderms. Bony fish are further divided into the lobe-finned and ray-finned fish . About 96% of all living fish species today are teleosts , a crown group of ray-finned fish that can protrude their jaws . The tetrapods ,
1680-606: The Gnathostomata or (for bony fish) Osteichthyes , also contains the clade of tetrapods (four-limbed vertebrates, mostly terrestrial), which are usually not considered fish. Some tetrapods, such as cetaceans and ichthyosaurs , have secondarily acquired a fish-like body shape through convergent evolution . Fishes of the World comments that "it is increasingly widely accepted that tetrapods, including ourselves, are simply modified bony fishes, and so we are comfortable with using
1750-709: The Indo-Pacific constitute the center of diversity for marine fishes, whereas continental freshwater fishes are most diverse in large river basins of tropical rainforests , especially the Amazon , Congo , and Mekong basins. More than 5,600 fish species inhabit Neotropical freshwaters alone, such that Neotropical fishes represent about 10% of all vertebrate species on the Earth. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams (e.g., char and gudgeon ) to
1820-522: The Late Pliocene of Orciano Pisano , Italy , is known from a cervical vertebra that may or may not have been from a loon. If so, it was from a bird slightly smaller than the common loon . Older authors were quite sure the bone was indeed from a Gavia and even considered G. concinna a possibly junior synonym of it. This is now regarded as rather unlikely due to the quite distinct range and age. The Early Pliocene Gavia skull from Empoli (Italy)
1890-561: The Silurian , with giant armoured placoderms such as Dunkleosteus . Jawed fish, too, appeared during the Silurian: the cartilaginous Chondrichthyes and the bony Osteichthyes . During the Devonian , fish diversity greatly increased, including among the placoderms, lobe-finned fishes, and early sharks, earning the Devonian the epithet "the age of fishes". Fishes are a paraphyletic group, since any clade containing all fish, such as
1960-469: The abyssal and even hadal depths of the deepest oceans (e.g., cusk-eels and snailfish ), although none have been found in the deepest 25% of the ocean. The deepest living fish in the ocean so far found is a cusk-eel, Abyssobrotula galatheae , recorded at the bottom of the Puerto Rico Trench at 8,370 m (27,460 ft). In terms of temperature, Jonah's icefish live in cold waters of
2030-400: The intertidal zone , are facultative air breathers, able to breathe air when out of water, as may occur daily at low tide , and to use their gills when in water. Some coastal fish like rockskippers and mudskippers choose to leave the water to feed in habitats temporarily exposed to the air. Some catfish absorb air through their digestive tracts. The digestive system consists of a tube,
2100-425: The stout infantfish . Swimming performance varies from fish such as tuna, salmon , and jacks that can cover 10–20 body-lengths per second to species such as eels and rays that swim no more than 0.5 body-lengths per second. A typical fish is cold-blooded , has a streamlined body for rapid swimming, extracts oxygen from water using gills, has two sets of paired fins, one or two dorsal fins, an anal fin and
2170-1107: The South Baltic and directly over land to the Black Sea or Mediterranean. Loons can live as long as 30 years and can hold their breath for as long as 90 seconds while underwater. Loons are migratory birds, and in the winter months they move from their northern freshwater lake nesting habitats to southern marine coastlines. They are well-adapted to this change in salinity, however, because they have special salt glands located directly above their eyes. These glands filter out salts in their blood and flush this salty solution out through their nasal passages, which allows them to immediately consume fish from oceans and drink saltwater after their long migration. Loons find their prey by sight. They eat mainly fish , supplemented with amphibians , crustaceans and similar mid-sized aquatic fauna . Specifically, they have been noted to feed on crayfish , frogs , snails , salamanders and leeches . They prefer clear lakes because they can more easily see their prey through
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2240-594: The Southern Ocean, including under the Filchner–Ronne Ice Shelf at a latitude of 79°S, while desert pupfish live in desert springs, streams, and marshes, sometimes highly saline, with water temperatures as high as 36 C. A few fish live mostly on land or lay their eggs on land near water. Mudskippers feed and interact with one another on mudflats and go underwater to hide in their burrows. A single undescribed species of Phreatobius has been called
2310-554: The army during the Peninsular War from 1809 to 1811 and was wounded in the Battle of Barossa on 5 March 1811. Though he had not yet completed his studies, he still published "An inquiry into the nature and extent of poetick licence" in London in 1810. He then returned to Oxford to continue his studies and achieved his Bachelor of Arts in 1817 and Master of Arts in 1818. He practiced as
2380-556: The attention of the cleaners. Cleaning behaviors have been observed in a number of fish groups, including an interesting case between two cichlids of the same genus, Etroplus maculatus , the cleaner, and the much larger E. suratensis . Fish occupy many trophic levels in freshwater and marine food webs . Fish at the higher levels are predatory , and a substantial part of their prey consists of other fish. In addition, mammals such as dolphins and seals feed on fish, alongside birds such as gannets and cormorants . The body of
2450-550: The body, and produce a concentrated urine. The reverse happens in freshwater fish : they tend to gain water osmotically, and produce a dilute urine. Some fish have kidneys able to operate in both freshwater and saltwater. Fish have small brains relative to body size compared with other vertebrates, typically one-fifteenth the brain mass of a similarly sized bird or mammal. However, some fish have relatively large brains, notably mormyrids and sharks , which have brains about as large for their body weight as birds and marsupials . At
2520-573: The bones of frogs and salamanders. The gastroliths may also be involved in stomach cleaning as an aid to regurgitation of indigestible food parts. Loons may inadvertently ingest small lead pellets, released by anglers and hunters, that will contribute to lead poisoning and the loon's eventual death. Jurisdictions that have banned the use of lead shot and sinkers include but are not limited to Maine , New Hampshire , Vermont , Michigan , some areas of Massachusetts , Yellowstone National Park , Canada , Great Britain , and Denmark . Loons nest during
2590-565: The death of the owner; in contrast, female loons usually survive. Birds that are displaced from a territory but survive usually try to re-mate and (re)claim a breeding territory later in life. In 2020, a loon hatched for the first time in over a century in Southeastern Massachusetts at Fall River, the Massachusetts Division of Fisheries and Wildlife and Biodiversity Research Institute. The chicks were relocated in 2015 with
2660-498: The ducks to the loons only in the 18th century. Earlier naturalists referred to the loons as mergus (the Latin term for diving seabirds of all sorts) or colymbus , which became the genus name used in the first modern scientific description of a Gavia species (by Carl Linnaeus ) in 1758. Unfortunately, confusion about whether Linnaeus' " wastebin genus " Colymbus referred to loons or grebes abounded. North American ornithologists used
2730-1201: The eggs are lost, the pair may re-nest, usually in a different location. Since the nest is very close to the water, rising water may induce the birds to slowly move the nest upwards, over a metre. Despite the roughly equal participation of the sexes in nest building and incubation, analysis has shown clearly that males alone select the location of the nest. This pattern has the important consequence that male loons, but not females, establish significant site-familiarity with their territories that allows them to produce more chicks there over time. Sex-biased site-familiarity might explain, in part, why resident males fight so hard to defend their territories. Most clutches consist of two eggs, which are laid in May or June, depending upon latitude. Loon chicks are precocial , able to swim and dive right away, but will often ride on their parents' back during their first two weeks to rest, conserve heat, and avoid predators. Chicks are fed mainly by their parents for about six weeks but gradually begin to feed themselves over time. By 11 or 12 weeks, chicks gather almost all of their own food and have begun to fly. In 2019,
2800-511: The exact root is unknown; some authorities reconstruct a Proto-Indo-European root * peysk- , attested only in Italic , Celtic , and Germanic . About 530 million years ago during the Cambrian explosion , fishlike animals with a notochord and eyes at the front of the body, such as Haikouichthys , appear in the fossil record . During the late Cambrian , other jawless forms such as conodonts appear. Jawed vertebrates appear in
2870-449: The extant species has found that the red-throated loons are the most basal of the five species. Gavia stellata G. arctica G. pacifica G. immer G. adamsii † Gavia egeriana † G. schultzi † G. howardae G. stellata † G. brodkorbi † G. paradoxa † G. moldavica † G. concinna G. arctica G. pacifica † G. fortis G. adamsii G. immer Nearly ten prehistoric species have been named to date in
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2940-405: The front of the brain are the olfactory lobes , a pair of structures that receive and process signals from the nostrils via the two olfactory nerves . Fish that hunt primarily by smell, such as hagfish and sharks, have very large olfactory lobes. Behind these is the telencephalon , which in fish deals mostly with olfaction. Together these structures form the forebrain. Connecting the forebrain to
3010-404: The future nest site, and build their nests close to the water, preferring sites that are completely surrounded by water such as islands or emergent vegetation. Loons use a variety of materials to build their nests including aquatic vegetation, pine needles, leaves, grass, moss and mud. Sometimes, nest material is almost lacking. Both male and female build the nest and incubate jointly for 28 days. If
3080-775: The genus Gavia originated in Europe during the Paleogene . The earliest species, G. egeriana , was found in early Miocene deposits in Dolnice in the Czech Republic. During the remainder of the Miocene , Gavia managed to disperse into North America via the Atlantic coastlines, eventually making their way to the continent's Pacific coastlines by the Late Miocene . Study of the interrelationships of
3150-524: The genus Gavia , and about as many undescribed ones await further study. The genus is known from the Early Miocene onwards, and the oldest members are rather small (some are smaller than the red-throated loon ). Throughout the late Neogene , the genus by and large follows Cope's Rule (that population lineages tend to increase in body size over evolutionary time). List of fossil Gavia species List of fossil Gavia specimens "Gavia" portisi from
3220-414: The genus name to refer to grebes, while Europeans used it for loons, following Nicholas Aylward Vigors and Richard Bowdler Sharpe . The International Commission on Zoological Nomenclature tried to settle this issue in 1956 by declaring Colymbus a suppressed name unfit for further use and establishing Gavia , created by Johann Reinhold Forster in 1788, as the valid genus name for the loons. However,
3290-412: The gills flows in the opposite direction to the water, resulting in efficient countercurrent exchange . The gills push the oxygen-poor water out through openings in the sides of the pharynx. Cartilaginous fish have multiple gill openings: sharks usually have five, sometimes six or seven pairs; they often have to swim to oxygenate their gills. Bony fish have a single gill opening on each side, hidden beneath
3360-532: The gills. Oxygen-rich blood then flows without further pumping, unlike in mammals, to the body tissues. Finally, oxygen-depleted blood returns to the heart. Fish exchange gases using gills on either side of the pharynx . Gills consist of comblike structures called filaments. Each filament contains a capillary network that provides a large surface area for exchanging oxygen and carbon dioxide . Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gills. Capillary blood in
3430-447: The gut, leading from the mouth to the anus. The mouth of most fishes contains teeth to grip prey, bite off or scrape plant material, or crush the food. An esophagus carries food to the stomach where it may be stored and partially digested. A sphincter, the pylorus, releases food to the intestine at intervals. Many fish have finger-shaped pouches, pyloric caeca , around the pylorus, of doubtful function. The pancreas secretes enzymes into
3500-494: The head droops more than in similar aquatic birds. Male and female loons have identical plumage , which is largely patterned black-and-white in summer, with grey on the head and neck in some species. All have a white belly. This resembles many sea-ducks (Merginae) – notably the smaller goldeneyes ( Bucephala ) – but is distinct from most cormorants, which rarely have white feathers, and if so, usually as large rounded patches rather than delicate patterns. All species of loons have
3570-452: The heart from the rest of the body. Lungfish, bichirs, ropefish, bowfins, snakefish, and the African knifefish have evolved to reduce such mixing, and to reduce oxygen loss from the gills to oxygen-poor water. Bichirs and lungfish have tetrapod-like paired lungs, requiring them to surface to gulp air, and making them obligate air breathers. Many other fish, including inhabitants of rock pools and
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#17328516346493640-521: The hopes of re-establishing breeding and nesting patterns. The European Anglophone name "diver" comes from the bird's habit of catching fish by swimming calmly along the surface and then abruptly plunging into the water. The North American name "loon" likely comes from either the Old English word lumme , meaning lummox or awkward person, or the Scandinavian word lum meaning lame or clumsy. Either way,
3710-455: The intestine to digest the food; other enzymes are secreted directly by the intestine itself. The liver produces bile which helps to break up fat into an emulsion which can be absorbed in the intestine. Most fish release their nitrogenous wastes as ammonia . This may be excreted through the gills or filtered by the kidneys . Salt is excreted by the rectal gland. Saltwater fish tend to lose water by osmosis ; their kidneys return water to
3780-446: The latter including all living cartilaginous and bony fish , as well as the extinct placoderms and acanthodians . Most fish are cold-blooded , their body temperature varying with the surrounding water, though some large active swimmers like white shark and tuna can hold a higher core temperature . Many fish can communicate acoustically with each other, such as during courtship displays . The earliest fish appeared during
3850-554: The loon's toes are connected by webbing . The loons may be confused with the cormorants (Phalacrocoracidae), but can be distinguished from them by their distinct call. Cormorants are not-too-distant relatives of loons, and like them are heavy-set birds whose bellies, unlike those of ducks and geese, are submerged when swimming. Loons in flight resemble plump geese with seagulls ' wings that are relatively small in proportion to their bulky bodies. The bird points its head slightly upwards while swimming, but less so than cormorants. In flight,
3920-419: The midbrain is the diencephalon ; it works with hormones and homeostasis . The pineal body is just above the diencephalon; it detects light, maintains circadian rhythms, and controls color changes. The midbrain contains the two optic lobes . These are very large in species that hunt by sight, such as rainbow trout and cichlids . The hindbrain controls swimming and balance.The single-lobed cerebellum
3990-549: The name refers to the loon's poor ability to walk on land. Another possible derivation is from the Norwegian word lom for these birds, which comes from Old Norse lómr , possibly cognate with English "lament", referring to the characteristic plaintive sound of the loon. The scientific name Gavia refers to seabirds in general. The scientific name Gavia was the Latin term for the smew ( Mergellus albellus ). This small sea-duck
4060-443: The nine largest families; from largest to smallest, these are Cyprinidae , Gobiidae , Cichlidae , Characidae , Loricariidae , Balitoridae , Serranidae , Labridae , and Scorpaenidae . About 64 families are monotypic , containing only one species. Fish range in size from the huge 16-metre (52 ft) whale shark to some tiny teleosts only 8-millimetre (0.3 in) long, such as the cyprinid Paedocypris progenetica and
4130-449: The ocean. Fish are caught for recreation , or raised by fishkeepers as ornaments for private and public exhibition in aquaria and garden ponds . Fish have had a role in human culture through the ages, serving as deities , religious symbols, and as the subjects of art, books and movies. The word fish is inherited from Proto-Germanic , and is related to German Fisch , the Latin piscis and Old Irish īasc , though
4200-402: The open ocean. Because the water all around is the same colour, reflecting an image of the water offers near-invisibility. Fish have a closed-loop circulatory system . The heart pumps the blood in a single loop throughout the body; for comparison, the mammal heart has two loops, one for the lungs to pick up oxygen, one for the body to deliver the oxygen. In fish, the heart pumps blood through
4270-421: The seats. On the death of one of them, Vigors won the subsequent by-election in 1837 and retained the seat until his own death. Fish A fish ( pl. : fish or fishes ) is an aquatic , anamniotic , gill -bearing vertebrate animal with swimming fins and a hard skull , but lacking limbs with digits . Fish can be grouped into the more basal jawless fish and the more common jawed fish ,
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#17328516346494340-489: The situation was not completely resolved even then, and the following year the ICZN had to act again to prevent Louis Pierre Vieillot 's 1818 almost-forgotten family name Urinatoridae from overruling the much younger Gaviidae. Some eminent ornithologists such as Pierce Brodkorb tried to keep the debate alive, but the ICZN's solution has been satisfactory. All living species are placed in the genus Gavia . It has been suggested that
4410-434: The summer on freshwater lakes and/or large ponds. Smaller bodies of water (up to 0.5 km) will usually only have one pair. Larger lakes may have more than one pair, with each pair occupying a bay or section of the lake. The red-throated loon, however, may nest colonially, several pairs close together, in small Arctic tarns and feed at sea or in larger lakes, ferrying the food in for the young. Loons mate on land, often on
4480-415: The taxon Osteichthyes as a clade, which now includes all tetrapods". The biodiversity of extant fish is unevenly distributed among the various groups; teleosts , bony fishes able to protrude their jaws , make up 96% of fish species. The cladogram shows the evolutionary relationships of all groups of living fishes (with their respective diversity ) and the tetrapods. Extinct groups are marked with
4550-545: The top 30 bird authors historically. He provided the text for John Gould 's A Century of Birds from the Himalaya Mountains (1830–32). One bird that he described was "Sabine's snipe". This was treated as a common snipe by Barrett-Hamilton in 1895 and by Meinertzhagen in 1926 but was thought to be probably a Wilson's snipe in 1945. Vigors lent a skin for later editions of Thomas Bewick 's History of British Birds . Vigors succeeded to his father's estate in 1828. He
4620-487: The water's surface with wings flapping to generate sufficient lift to take flight. Only the red-throated loon ( G. stellata ) can take off from land. Once airborne, loons are capable of long flights during migration. Scientists from the U.S. Geological Survey, who have implanted satellite transmitters in some individuals, have recorded daily flights of up to 1078 km in a 24-hour period, which probably resulted from single movements. North European loons migrate primarily via
4690-419: The water. The loon uses its pointy bill to stab or grasp prey. They eat vertebrate prey headfirst to facilitate swallowing, and swallow all their prey whole. To help digestion, loons swallow small pebbles from the bottoms of lakes. Similar to grit eaten by chickens , these gastroliths may assist the loon's gizzard in crushing the hard parts of the loon's food such as the exoskeletons of crustaceans and
4760-588: Was MP for the borough of Carlow from 1832 until 1835. He briefly represented County Carlow in 1835. Vigors had been elected in a by-election in June after the Conservative MPs originally returned at the 1835 United Kingdom general election were unseated on petition and a new writ issued. On 19 August 1835, Vigors and his running mate, in the two-member county constituency, were unseated on petition. The same two Conservatives who had previously been unseated were awarded
4830-457: Was born at Old Leighlin , County Carlow , in 1785. He was the first son of Capt. Nicholas Aylward Vigors, who served in the 29th (Worcestershire) Regiment , and his first wife, Catherine Vigors, daughter of Solomon Richards of Solsborough. He matriculated at Trinity College, Oxford , in November 1803, and was admitted at Lincoln's Inn in November 1806. Without completing his studies, he served in
4900-429: Was referred to G. concinna , and thus could conceivably have been of "G." portisi if that was indeed a loon. The holotype vertebra may now be lost, which would make "G." portisi a nomen dubium . Nicholas Aylward Vigors Nicholas Aylward Vigors (1785 – 26 October 1840) was an Irish zoologist and politician . He popularized the classification of birds on the basis of the quinarian system . Vigors
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