137-599: Pollachius virens is a species of marine fish in the genus Pollachius . Together with P. pollachius , it is generally referred to in the United States as pollock . It is commonly known in Britain as the coalfish , coley , or saithe ( / s eɪ ð / or / s eɪ θ / ), and the young fish may also be called podleys in Scotland and northern England. This species can be separated from P. pollachius by looking at
274-728: A general armistice ended the fighting. In the German Revolution (November 1918), Wilhelm II and the ruling princes abdicated their positions, and Germany was declared a federal republic . Germany's new leadership signed the Treaty of Versailles in 1919, accepting defeat by the Allies . Germans perceived the treaty as humiliating, which was seen by historians as influential in the rise of Adolf Hitler . Germany lost around 13% of its European territory and ceded all of its colonial possessions in Africa and
411-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
548-505: A sustainable energy supply, the debt brake for balanced budgets, measures to increase the fertility rate ( pronatalism ), and high-tech strategies for the transition of the German economy, summarised as Industry 4.0 . During the 2015 European migrant crisis , the country took in over a million refugees and migrants. Germany is the seventh-largest country in Europe . It borders Denmark to
685-555: A temperate climate, ranging from oceanic in the north and west to continental in the east and southeast. Winters range from the cold in the Southern Alps to cool and are generally overcast with limited precipitation, while summers can vary from hot and dry to cool and rainy. The northern regions have prevailing westerly winds that bring in moist air from the North Sea, moderating the temperature and increasing precipitation. Conversely,
822-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
959-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
1096-455: A NATO effort to provide security in that country after the ousting of the Taliban . In the 2005 elections , Angela Merkel became the first female chancellor. In 2009, the German government approved a €50 billion stimulus plan. Among the major German political projects of the early 21st century are the advancement of European integration , the energy transition ( Energiewende ) for
1233-638: A centralised totalitarian state , withdrew from the League of Nations , and dramatically increased the country's rearmament . A government-sponsored programme for economic renewal focused on public works, the most famous of which was the Autobahn . In 1935, the regime withdrew from the Treaty of Versailles and introduced the Nuremberg Laws which targeted Jews and other minorities. Germany also reacquired control of
1370-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
1507-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
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#17328587864491644-599: A framework laid out in the 1949 constitution known as the Grundgesetz (Basic Law). Amendments generally require a two-thirds majority of both the Bundestag and the Bundesrat ; the fundamental principles of the constitution, as expressed in the articles guaranteeing human dignity, the separation of powers, the federal structure, and the rule of law , are valid in perpetuity. The president , currently Frank-Walter Steinmeier ,
1781-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
1918-430: A higher proportion of fish which are of poor quality. There is a detectable difference in taste between saithe associated and not associated with fish farms, but there are conflicting results on which tastes better and is probably a matter of consumer preference. Saithe is also used as food for domestic cats . Species A species ( pl. : species) is a population of organisms in which any two individuals of
2055-658: A joint responsibility of the international community. It was the world's second-biggest aid donor in 2019 after the United States. Germany's military, the Bundeswehr (Federal Defence), is organised into the Heer (Army and special forces KSK ), Marine (Navy), Luftwaffe (Air Force) and Cyber- und Informationsraum (Cyber and Information Domain Service) branches. In absolute terms, German military spending in 2023
2192-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On
2329-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
2466-399: A noticeable kink over the pectoral fins while that of P. virens is straighter. The flesh of coalfish ( P. virens ) is darkly coloured (hence the common name) while that of P. pollachius is similar to other members of the cod family. This dark colour in the fresh uncooked flesh may have led to the undeserved reputation of this fish as poor for eating. It is common in the northern parts of
2603-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
2740-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that
2877-537: A provisional capital, to emphasise its stance that the two-state solution was temporary. West Germany was established as a federal parliamentary republic with a " social market economy ". Starting in 1948 West Germany became a major recipient of reconstruction aid under the American Marshall Plan . Konrad Adenauer was elected the first federal chancellor of Germany in 1949. The country enjoyed prolonged economic growth ( Wirtschaftswunder ) beginning in
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#17328587864493014-565: A series of Imperial Reforms (approximately 1495–1555) provided for considerable local autonomy and a stronger Imperial Diet . The House of Habsburg held the imperial crown from 1438 until the death of Charles VI in 1740. Following the War of the Austrian Succession and the Treaty of Aix-la-Chapelle , Charles VI's daughter Maria Theresa ruled as empress consort when her husband, Francis I , became emperor. From 1740, dualism between
3151-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
3288-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
3425-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
3562-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
3699-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
3836-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
3973-615: A state, the East German authorities eased border restrictions, but this actually led to an acceleration of the Wende reform process culminating in the Two Plus Four Treaty under which Germany regained full sovereignty. This permitted German reunification on 3 October 1990, with the accession of the five re-established states of the former GDR. The fall of the Wall in 1989 became a symbol of
4110-692: A symbol of the Cold War . Tensions between East and West Germany were reduced in the late 1960s by Chancellor Willy Brandt 's Ostpolitik . In 1989, Hungary decided to dismantle the Iron Curtain and open its border with Austria , causing the emigration of thousands of East Germans to West Germany via Hungary and Austria. This had devastating effects on the GDR, where regular mass demonstrations received increasing support. In an effort to help retain East Germany as
4247-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
Pollachius virens - Misplaced Pages Continue
4384-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
4521-613: A universal health care system , and tuition-free university education . Germany is a member of the United Nations , Council of Europe , NATO and OECD , and a founding member of the European Union, G7 and G20 . It has the third-highest number of UNESCO World Heritage Sites , 54 , of which 51 are cultural. The English word Germany derives from the Latin Germania , which came into use after Julius Caesar adopted it for
4658-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
4795-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
4932-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
5069-541: Is 22.7 kilograms (50 lb 1 oz) which was caught at Saltstraumen in Norway . Saithe reach sexual maturity at 4–9 years old and are iteroparous , batch spawners with determinate fecundity. Females produce, depending on their size, between 500 thousand and 9 million eggs which are 1.0 to 1.3 millimetres ( 5 ⁄ 128 to 7 ⁄ 128 in) in diameter. Saithe is fished year-round using gear such as Danish seine nets , trawlers , long lines and gill nets and
5206-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
5343-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
5480-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
5617-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,
Pollachius virens - Misplaced Pages Continue
5754-470: Is considerably less valuable than premium whitefish such as cod and haddock . To achieve a salmon -like orange color, it can be salted and smoked. In Germany , the fish is commonly sold as Seelachs (literally 'sea salmon'), although it is not closely related to any salmon . While a great deal of saithe consumed in Europe are caught in British waters, it is not a popular fish with consumers there. Most of
5891-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
6028-637: Is devoted to agriculture, while 30% is forested and 14% is covered by settlements or infrastructure. Plants and animals include those generally common to Central Europe. According to the National Forest Inventory, beeches , oaks , and other deciduous trees constitute just over 40% of the forests; roughly 60% are conifers , particularly spruce and pine . There are many species of ferns , flowers , fungi , and mosses . Wild animals include roe deer , wild boar , mouflon (a subspecies of wild sheep), fox , badger , hare , and small numbers of
6165-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
6302-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
6439-599: Is largely autonomous in regard to its internal organisation. As of 2017 , Germany is divided into 401 districts ( Kreise ) at a municipal level; these consist of 294 rural districts and 107 urban districts . Germany has a civil law system based on Roman law with some references to Germanic law . The Bundesverfassungsgericht (Federal Constitutional Court) is the German Supreme Court responsible for constitutional matters, with power of judicial review . Germany's specialized supreme court system includes
6576-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
6713-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
6850-570: Is often caught in mixed species fishery with other groundfish species such as cod and whiting. The main fishing grounds in the eastern Atlantic are in the Barents Sea , around Iceland , around the Faeroe Islands and in the North Sea and Celtic Sea . Landings in the eastern Atlantic have fluctuated around 300–700 thousand metric tons (300,000–690,000 long tons; 330,000–770,000 short tons) in
6987-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
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#17328587864497124-490: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Germany – in Europe (light green & dark grey) – in
7261-627: Is the head of state and invested primarily with representative responsibilities and powers. He is elected by the Bundesversammlung (federal convention), an institution consisting of the members of the Bundestag and an equal number of state delegates. The second-highest official in the German order of precedence is the Bundestagspräsident ( President of the Bundestag ), who is elected by
7398-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
7535-483: Is vested in the parliament consisting of the Bundestag (Federal Diet) and Bundesrat (Federal Council), which together form the legislative body. The Bundestag is elected through direct elections using the mixed-member proportional representation system. The members of the Bundesrat represent and are appointed by the governments of the sixteen federated states. The German political system operates under
7672-457: The Gründerzeit period following the unification of Germany, Bismarck's foreign policy as chancellor of Germany secured Germany's position as a great nation by forging alliances and avoiding war. However, under Wilhelm II , Germany took an imperialistic course, leading to friction with neighbouring countries. A dual alliance was created with the multinational realm of Austria-Hungary ;
7809-484: The Bundestag and responsible for overseeing the daily sessions of the body. The third-highest official and the head of government is the chancellor, who is appointed by the Bundespräsident after being elected by the party or coalition with the most seats in the Bundestag . The chancellor, currently Olaf Scholz , is the head of government and exercises executive power through his Cabinet . Since 1949,
7946-861: The Allies de jure abolished the German state and partitioned Berlin and Germany's remaining territory into four occupation zones. The western sectors, controlled by France, the United Kingdom , and the United States , were merged on 23 May 1949 to form the Federal Republic of Germany ( German : Bundesrepublik Deutschland ); on 7 October 1949, the Soviet Zone became the German Democratic Republic (GDR) ( German : Deutsche Demokratische Republik ; DDR). They were informally known as West Germany and East Germany. East Germany selected East Berlin as its capital, while West Germany chose Bonn as
8083-607: The Battle of Britain in the same year. In 1941, German troops invaded Yugoslavia , Greece and the Soviet Union . By 1942, Germany and its allies controlled most of continental Europe and North Africa , but following the Soviet victory at the Battle of Stalingrad , the Allied reconquest of North Africa and invasion of Italy in 1943, German forces suffered repeated military defeats. In 1944,
8220-578: The Berchtesgaden National Park . In addition, there are 17 Biosphere Reserves , and 105 nature parks . More than 400 zoos and animal parks operate in Germany. The Berlin Zoo , which opened in 1844, is the oldest in Germany, and claims the most comprehensive collection of species in the world. Germany is a federal , parliamentary , representative democratic republic. Federal legislative power
8357-1085: The Eurasian beaver . The blue cornflower was once a German national symbol . The 16 national parks in Germany include the Jasmund National Park , the Vorpommern Lagoon Area National Park , the Müritz National Park , the Wadden Sea National Parks , the Harz National Park , the Hainich National Park , the Black Forest National Park , the Saxon Switzerland National Park , the Bavarian Forest National Park and
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#17328587864498494-605: The European Union (light green) Germany , officially the Federal Republic of Germany , is a country in Central Europe . It lies between the Baltic and North Sea to the north and the Alps to the south. Its sixteen constituent states have a total population of over 82 million in an area of 357,596 km (138,069 sq mi), making it the most populous member state of
8631-643: The Fall of Communism , the dissolution of the Soviet Union , German reunification and Die Wende ("the turning point"). United Germany was considered the enlarged continuation of West Germany so it retained its memberships in international organisations. Based on the Berlin/Bonn Act (1994), Berlin again became the capital of Germany, while Bonn obtained the unique status of a Bundesstadt (federal city) retaining some federal ministries. The relocation of
8768-628: The German Revolution of 1918–1919 , the Empire was replaced by the Weimar Republic . The Nazi rise to power in 1933 led to the establishment of a totalitarian dictatorship , World War II , and the Holocaust . In 1949, after the war and a period of Allied occupation , Germany was organized into two separate polities with limited sovereignty: the Federal Republic of Germany, or West Germany , and
8905-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
9042-694: The Investiture controversy . Under the Hohenstaufen emperors (1138–1254), German princes encouraged German settlement to the south and east ( Ostsiedlung ). Members of the Hanseatic League , mostly north German towns, prospered in the expansion of trade. The population declined starting with the Great Famine in 1315, followed by the Black Death of 1348–1350. The Golden Bull issued in 1356 provided
9179-635: The Jastorf culture during the Nordic Bronze Age or early Iron Age . From southern Scandinavia and northern Germany , they expanded south, east, and west, coming into contact with the Celtic , Iranian , Baltic , and Slavic tribes. Southern Germany was inhabited by Celtic-speaking peoples, who belonged to the wider La Tène culture . They were later assimilated by the Germanic conquerors. Under Augustus ,
9316-408: The Lower Paleolithic , with various tribes inhabiting it from the Neolithic onward, chiefly the Celts . Various Germanic tribes have inhabited the northern parts of modern Germany since classical antiquity . A region named Germania was documented before AD 100. In 962, the Kingdom of Germany formed the bulk of the Holy Roman Empire . During the 16th century, northern German regions became
9453-467: The Molotov–Ribbentrop Pact that divided Eastern Europe into German and Soviet spheres of influence. On 1 September 1939, Germany invaded Poland , beginning World War II in Europe ; Britain and France declared war on Germany on 3 September. In the spring of 1940, Germany conquered Denmark and Norway , the Netherlands , Belgium , Luxembourg , and France , forcing the French government to sign an armistice . The British repelled German air attacks in
9590-432: The Northern Atlantic , including the Bay of Biscay . Adults can typically live up to 16–20 years and grow to 100–120 centimetres (39–47 in) but individuals up to 130 cm (51 in) and weight up to 32 kg (70 lb 9 oz) have been caught. Juveniles tend to be found close to shore, particularly in rocky areas, and tend to move out into deeper waters as they grow. The current IGFA All-Tackle World Record
9727-529: The Roman Empire began to invade lands inhabited by the Germanic tribes, creating a short-lived Roman province of Germania between the Rhine and Elbe rivers. In 9 AD, three Roman legions were defeated by Arminius in the Battle of the Teutoburg Forest . The outcome of this battle dissuaded the Romans from their ambition of conquering Germania and is thus considered one of the most important events in European history . By 100 AD, when Tacitus wrote Germania , Germanic tribes had settled along
9864-692: The Saarland in 1935, remilitarised the Rhineland in 1936, annexed Austria in 1938, annexed the Sudetenland in 1938 with the Munich Agreement , and in violation of the agreement occupied Czechoslovakia in March 1939. Kristallnacht (Night of Broken Glass) saw the burning of synagogues, the destruction of Jewish businesses, and mass arrests of Jewish people. In August 1939, Hitler's government negotiated
10001-616: The Triple Alliance of 1882 included Italy. Britain, France and Russia also concluded alliances to protect against Habsburg interference with Russian interests in the Balkans or German interference against France. At the Berlin Conference in 1884, Germany claimed several colonies including German East Africa , German South West Africa , Togoland , and Kamerun . Later, Germany further expanded its colonial empire to include holdings in
10138-401: The chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part
10275-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
10412-409: The former East German states join the Federal Republic of Germany on 3 October 1990 . Germany has been described as a great power with a strong economy ; it has the largest economy in Europe by nominal GDP . As a global power in industrial, scientific and technological sectors, it is both the world's third-largest exporter and importer . As a developed country , it offers social security ,
10549-595: The inquisitorial Federal Court of Justice for civil and criminal cases, along with the Federal Labour Court , Federal Social Court , Federal Fiscal Court , and Federal Administrative Court for other matters. Criminal and private laws are codified on the national level in the Strafgesetzbuch and the Bürgerliches Gesetzbuch respectively. The German penal system seeks the rehabilitation of
10686-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
10823-488: The revolutions of 1848 in the German states , raising the German question. King Frederick William IV of Prussia was offered the title of emperor, but with a loss of power; he rejected the crown and the proposed constitution, a temporary setback for the movement. King William I appointed Otto von Bismarck as the Minister President of Prussia in 1862. Bismarck successfully concluded the war with Denmark in 1864 ;
10960-593: The "Evangelical" faith ( Lutheranism ), but also decreed that the faith of the prince was to be the faith of his subjects ( cuius regio, eius religio ). From the Cologne War through the Thirty Years' Wars (1618–1648), religious conflict devastated German lands and significantly reduced the population. The Peace of Westphalia ended religious warfare among the Imperial Estates . The legal system initiated by
11097-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
11234-619: The Alps. Subsequently, the Holy Roman Empire emerged from it. The Ottonian rulers (919–1024) consolidated several major duchies . In 996, Gregory V became the first German Pope, appointed by his cousin Otto III , whom he shortly after crowned Holy Roman Emperor. The Holy Roman Empire absorbed northern Italy and Burgundy under the Salian emperors (1024–1125), although the emperors lost power through
11371-865: The Austrian Habsburg monarchy and the Kingdom of Prussia dominated German history. In 1772, 1793, and 1795, Prussia and Austria, along with the Russian Empire , agreed to the Partitions of Poland . During the period of the French Revolutionary Wars , the Napoleonic era and the subsequent final meeting of the Imperial Diet , most of the Free Imperial Cities were annexed by dynastic territories;
11508-515: The British saithe catch is thus exported to France, where it is widely eaten. Saithe often congregate around fish farms and feed on uneaten salmon feeds which have passed through the net walls of the cages. It is the opinion of fishermen that the fish caught close to fish farms is of poor quality. Quality testing shows that the there is relatively minor differences in fillet quality between fish which were caught close to fish farms and those that were not, however for catches taken close to fish farms have
11645-574: The European Union . It borders Denmark to the north, Poland and the Czech Republic to the east, Austria and Switzerland to the south, and France , Luxembourg , Belgium , and the Netherlands to the west. The nation's capital and most populous city is Berlin and its main financial centre is Frankfurt ; the largest urban area is the Ruhr . Settlement in the territory of modern Germany began in
11782-667: The Franks established the Frankish Kingdom and pushed east to subjugate Saxony and Bavaria . Areas of what is today eastern Germany were inhabited by Western Slavic tribes. Charlemagne founded the Carolingian Empire in 800; it was divided in 843 . The eastern successor kingdom of East Francia stretched from the Rhine in the west to the Elbe river in the east and from the North Sea to
11919-496: The German Bundestag , though they have never been part of the federal government. In the 2017 German federal election , the right-wing populist Alternative for Germany gained enough votes to attain representation in the parliament for the first time. Germany is a federation and comprises sixteen constituent states which are collectively referred to as Länder . Each state ( Land ) has its own constitution, and
12056-614: The German Democratic Republic, or East Germany . Berlin continued its de jure Four Power status . The Federal Republic of Germany was a founding member of the European Economic Community and the European Union , while the German Democratic Republic was a communist Eastern Bloc state and member of the Warsaw Pact . After the fall of the communist led-government in East Germany, German reunification saw
12193-486: The Holocaust , the German government persecuted minorities , including interning them in concentration and death camps across Europe. The regime systematically murdered 6 million Jews, at least 130,000 Romani , 275,000 disabled , thousands of Jehovah's Witnesses , thousands of homosexuals , and hundreds of thousands of political and religious opponents . Nazi policies in German-occupied countries resulted in
12330-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
12467-617: The Pacific and China. The colonial government in South West Africa (present-day Namibia ), from 1904 to 1907, carried out the annihilation of the local Herero and Namaqua peoples as punishment for an uprising; this was the 20th century's first genocide . The assassination of Austria's crown prince on 28 June 1914 provided the pretext for Austria-Hungary to attack Serbia and trigger World War I . After four years of warfare, in which approximately two million German soldiers were killed,
12604-664: The Pacific. On 11 August 1919, President Friedrich Ebert signed the democratic Weimar Constitution . In the subsequent struggle for power, communists seized power in Bavaria , but conservative elements elsewhere attempted to overthrow the Republic in the Kapp Putsch . Street fighting in the major industrial centres, the occupation of the Ruhr by Belgian and French troops, and a period of hyperinflation followed. A debt restructuring plan and
12741-583: The Reichstag after a special election in 1932 and Hindenburg appointed Hitler as chancellor of Germany on 30 January 1933. After the Reichstag fire , a decree abrogated basic civil rights and the first Nazi concentration camp opened. On 23 March 1933, the Enabling Act gave Hitler unrestricted legislative power, overriding the constitution, and marked the beginning of Nazi Germany. His government established
12878-571: The Rhine and the Danube (the Limes Germanicus ), occupying most of modern Germany. However, Baden-Württemberg , southern Bavaria , southern Hesse and the western Rhineland had been incorporated into Roman provinces . Around 260, Germanic peoples broke into Roman-controlled lands. After the invasion of the Huns in 375, and with the decline of Rome from 395, Germanic tribes moved farther southwest:
13015-653: The Soviets pushed into Eastern Europe ; the Western allies landed in France and entered Germany despite a final German counteroffensive . Following Hitler's suicide during the Battle of Berlin , Germany signed the surrender document on 8 May 1945, ending World War II in Europe and Nazi Germany. Following the end of the war, surviving Nazi officials were tried for war crimes at the Nuremberg trials . In what later became known as
13152-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
13289-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
13426-404: The appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use the concept of
13563-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
13700-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
13837-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be
13974-803: The centre of the Protestant Reformation . Following the Napoleonic Wars and the dissolution of the Holy Roman Empire in 1806, the German Confederation was formed in 1815. Formal unification of Germany into the modern nation-state commenced on 18 August 1866 with the North German Confederation Treaty establishing the Prussia -led North German Confederation , which became the German Empire in 1871. After World War I and
14111-522: The communist-controlled Socialist Unity Party of Germany , supported by the Stasi , an immense secret service. While East German propaganda was based on the benefits of the GDR's social programmes and the alleged threat of a West German invasion, many of its citizens looked to the West for freedom and prosperity. The Berlin Wall , built in 1961, prevented East German citizens from escaping to West Germany, becoming
14248-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
14385-577: The constitutional structure of the Empire and codified the election of the emperor by seven prince-electors . Johannes Gutenberg introduced moveable-type printing to Europe, laying the basis for the democratization of knowledge . In 1517, Martin Luther incited the Protestant Reformation and his translation of the Bible began the standardization of the language; the 1555 Peace of Augsburg tolerated
14522-462: The creation of a new currency in 1924 ushered in the Golden Twenties , an era of artistic innovation and liberal cultural life. The worldwide Great Depression hit Germany in 1929. Chancellor Heinrich Brüning 's government pursued a policy of fiscal austerity and deflation which caused unemployment of nearly 30% by 1932. The Nazi Party led by Adolf Hitler became the largest party in
14659-466: The creation of a more unified European political, economic and security apparatus. The governments of Germany and the United States are close political allies . Cultural ties and economic interests have crafted a bond between the two countries resulting in Atlanticism . After 1990, Germany and Russia worked together to establish a "strategic partnership" in which energy development became one of
14796-536: The criminal and the protection of the public. With the exceptions of petty crimes, tried by a single professional judge, and of serious political crimes, all charges are adjudicated by mixed tribunals where lay judges ( Schöffen ) and professional judges preside together. As of 2016, Germany's murder rate stood at a low of 1.18 murders per 100,000. In 2018, the overall crime rate fell to its lowest since 1992. Same-sex marriage has been legal in Germany since 2017, and LGBT rights are generally protected in
14933-461: The deaths of an estimated 2.7 million Poles , 1.3 million Ukrainians , 1 million Belarusians and 3.5 million Soviet prisoners of war . German military casualties have been estimated at 5.3 million, and around 900,000 German civilians died. Around 12 million ethnic Germans were expelled from across Eastern Europe, and Germany lost roughly one-quarter of its pre-war territory. After Nazi Germany surrendered,
15070-447: The early 1950s. West Germany joined NATO in 1955 and was a founding member of the European Economic Community . On 1 January 1957, the Saarland joined West Germany. East Germany was an Eastern Bloc state under political and military control by the Soviet Union via occupation forces and the Warsaw Pact . Although East Germany claimed to be a democracy, political power was exercised solely by leading members ( Politbüro ) of
15207-624: The ecclesiastical territories were secularised and annexed. In 1806 the Imperium was dissolved; France, Russia, Prussia, and the Habsburgs (Austria) competed for hegemony in the German states during the Napoleonic Wars . Following the fall of Napoleon , the Congress of Vienna founded the German Confederation, a loose league of 39 sovereign states . The appointment of the emperor of Austria as
15344-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
15481-597: The government was completed in 1999, and modernisation of the East German economy was scheduled to last until 2019. Since reunification, Germany has taken a more active role in the European Union , signing the Maastricht Treaty in 1992 and the Lisbon Treaty in 2007, and co-founding the eurozone . Germany sent a peacekeeping force to secure stability in the Balkans and sent German troops to Afghanistan as part of
15618-469: The most important factors. As a result of the cooperation, Germany imported most of its natural gas and crude oil from Russia. Germany's development policy functions as a distinct sector within its foreign policy framework. It is formulated by the Federal Ministry for Economic Cooperation and Development and carried out by the implementing organisations. The German government sees development policy as
15755-641: The nation. Germany has a network of 227 diplomatic missions abroad and maintains relations with more than 190 countries. Germany is a member of NATO , the OECD , the G7 , the G20 , the World Bank and the IMF . It has played an influential role in the European Union since its inception and has maintained a strong alliance with France and all neighbouring countries since 1990. Germany promotes
15892-539: The north, Poland and Czechia to the east, Austria and Switzerland to the south, and France , Luxembourg , Belgium , and the Netherlands to the west. Germany is also bordered by the North Sea and, at the north-northeast, by the Baltic Sea. German territory covers 357,596 km (138,069 sq mi). Elevation ranges from the mountains of the Alps (highest point: the Zugspitze at 2,963 metres or 9,721 feet) in
16029-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
16166-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
16303-587: The party system has been dominated by the Christian Democratic Union and the Social Democratic Party of Germany . So far every chancellor has been a member of one of these parties. However, the smaller liberal Free Democratic Party and the Alliance 90/The Greens have also been junior partners in coalition governments . Since 2007, the democratic socialist party The Left has been a staple in
16440-661: The people' (see also the Latinised form Theodiscus ), derived from * þeudō , descended from Proto-Indo-European * tewtéh₂- 'people', from which the word Teutons also originates. Pre-human ancestors, the Danuvius guggenmosi , who were present in Germany over 11 million years ago, are theorized to be among the earliest ones to walk on two legs. Ancient humans were present in Germany at least 600,000 years ago. The first non-modern human fossil (the Neanderthal )
16577-503: The peoples east of the Rhine . The German term Deutschland , originally diutisciu land ('the German lands'), is derived from deutsch ( cf. Dutch ), descended from Old High German diutisc 'of the people' (from diot or diota 'people'), originally used to distinguish the language of the common people from Latin and its Romance descendants . This in turn descends from Proto-Germanic * þiudiskaz 'of
16714-555: The period 1980–2017. All the stocks in eastern Atlantic are assessed by International Council for the Exploration of the Sea (ICES) which publish a recommendations on an annual basis for Total Allowable Catch. The commercial catch of saithe in the western Atlantic is taken by USA and Canada and has fluctuated around 5–45 thousand metric tons (4,900–44,300 long tons; 5,500–49,600 short tons) per year between 1980 and 2018. The population in
16851-472: The permanent president reflected the Congress's rejection of Prussia 's rising influence. Disagreement within restoration politics partly led to the rise of liberal movements, followed by new measures of repression by Austrian statesman Klemens von Metternich . The Zollverein , a tariff union, furthered economic unity. In light of revolutionary movements in Europe , intellectuals and commoners started
16988-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
17125-461: The relative lengths of the upper and lower jaws. P. pollachius has a longer underslung lower jaw while P. virens has approximately equal upper and lower jaw lengths. This gives a very different profile to the head. In general, P. pollachius is a brown or golden colour with a dark back while P. virens is bright silver with a very dark green back. P. virens generally appears to have relatively smaller eyes. The lateral line of P. pollachius has
17262-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
17399-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
17536-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
17673-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
17810-549: The south to the shores of the North Sea ( Nordsee ) in the northwest and the Baltic Sea ( Ostsee ) in the northeast. The forested uplands of central Germany and the lowlands of northern Germany (lowest point: in the municipality Neuendorf-Sachsenbande , Wilstermarsch at 3.54 metres or 11.6 feet below sea level ) are traversed by such major rivers as the Rhine, Danube and Elbe. Significant natural resources include iron ore, coal, potash , timber, lignite , uranium , copper, natural gas, salt, and nickel. Most of Germany has
17947-756: The southeast regions have more extreme temperatures. From February 2019 – 2020, average monthly temperatures in Germany ranged from a low of 3.3 °C (37.9 °F) in January 2020 to a high of 19.8 °C (67.6 °F) in June 2019. Average monthly precipitation ranged from 30 litres per square metre in February and April 2019 to 125 litres per square metre in February 2020. Average monthly hours of sunshine ranged from 45 in November 2019 to 300 in June 2019. The territory of Germany can be divided into five terrestrial ecoregions : Atlantic mixed forests , Baltic mixed forests , Central European mixed forests , Western European broadleaf forests , and Alps conifer and mixed forests . As of 2016 , 51% of Germany's land area
18084-456: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
18221-429: The subsequent decisive Prussian victory in the Austro-Prussian War of 1866 enabled him to create the North German Confederation which excluded Austria . After the defeat of France in the Franco-Prussian War , the German princes proclaimed the founding of the German Empire in 1871. Prussia was the dominant constituent state of the new empire; the King of Prussia ruled as its Kaiser, and Berlin became its capital. In
18358-406: The western Atlantic is assessed by USA National Oceanic and Atmospheric Administration (NOAA) and Fisheries and Oceans Canada . All four stocks assessed in the eastern Atlantic and the stock in the western Atlantic are harvested sustainably with many saithe fisheries having been certified as sustainable by the Marine Stewardship Council . Coalfish is edible and has commercial value, although it
18495-494: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
18632-454: Was discovered in the Neander Valley . Similarly dated evidence of modern humans has been found in the Swabian Jura , including 42,000-year-old flutes which are the oldest musical instruments ever found, the 40,000-year-old Lion Man , and the 41,000-year-old Venus of Hohle Fels . The Nebra sky disk , created during the European Bronze Age , has been attributed to a German site. The Germanic peoples are thought to have emerged from
18769-444: Was the seventh-highest in the world . In response to the 2022 Russian invasion of Ukraine , Chancellor Olaf Scholz announced that German military expenditure would be increased past the NATO target of 2%, along with a one-time 2022 infusion of 100 billion euros, representing almost double the 53 billion euro military budget for 2021. In 2023, military spending according to NATO criteria amounted to $ 73.1 billion, or 1.64% of
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