An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree that contains any multifurcations can be described as a multifurcating tree.
57-622: 34; see text Cistus (from the Greek kistos ) is a genus of flowering plants in the rockrose family Cistaceae , containing about 20 species (Ellul et al. 2002). They are perennial shrubs found on dry or rocky soils throughout the Mediterranean region , from Morocco and Portugal through to the Middle East , and also on the Canary Islands . Cistus , with its many hybrids and cultivars,
114-471: A synonym of Cistus . Molecular phylogenetic analyses conducted between 2005 and 2011 confirm that Cistus species divide into two well-defined clades , neither of which was fully resolved internally. The first clade consists of those with purple and pink flowers (the "purple pink clade" or PPC). The second clade consists of those with white flowers or, in the case of Cistus parviflorus , pale pink flowers (the "white or whitish pink clade" or WWPC). Although
171-570: A basis for coinages: anthropology , photography , telephony , isomer , biomechanics , cinematography , etc. Together with Latin words , they form the foundation of international scientific and technical vocabulary ; for example, all words ending in -logy ('discourse'). There are many English words of Greek origin . Greek is an independent branch of the Indo-European language family. The ancient language most closely related to it may be ancient Macedonian , which, by most accounts,
228-552: A fairly stable set of consonantal contrasts . The main phonological changes occurred during the Hellenistic and Roman period (see Koine Greek phonology for details): In all its stages, the morphology of Greek shows an extensive set of productive derivational affixes , a limited but productive system of compounding and a rich inflectional system. Although its morphological categories have been fairly stable over time, morphological changes are present throughout, particularly in
285-556: A faster, more convenient cursive writing style with the use of ink and quill . The Greek alphabet consists of 24 letters, each with an uppercase ( majuscule ) and lowercase ( minuscule ) form. The letter sigma has an additional lowercase form (ς) used in the final position of a word: In addition to the letters, the Greek alphabet features a number of diacritical signs : three different accent marks ( acute , grave , and circumflex ), originally denoting different shapes of pitch accent on
342-540: A few species (notably C. ladanifer ), the leaves are coated with a highly aromatic resin called labdanum . They have showy 5-petaled flowers ranging from white to purple and dark pink, in a few species with a conspicuous dark red spot at the base of each petal. Cistus and Halimium form a cohesive and the most derived clade within Cistaceae . Plants of the World Online and World Flora Online treat Halimium as
399-531: A foreign language. It is also often stated that the historical changes have been relatively slight compared with some other languages. According to one estimation, " Homeric Greek is probably closer to Demotic than 12-century Middle English is to modern spoken English ". Greek is spoken today by at least 13 million people, principally in Greece and Cyprus along with a sizable Greek-speaking minority in Albania near
456-463: A nearly frost-free Mediterranean climate . The hardiest of the species is C. laurifolius , which survived the hard frost at Royal Botanical Gardens at Kew in 1895 that eliminated all the cistuses save this and two white-flowered natural hybrids, C. × corbariensis , already grown by John Tradescant the Elder , and C. × loretii , a 19th-century introduction. Cultivars (those marked agm have gained
513-448: A short period of time when in flower. The presence of the parasite does not seem to harm the host population. Cistus species are used as food plants by the larvae of some Lepidoptera species including Coleophora confluella and Coleophora helianthemella , the latter recorded on Cistus monspeliensis . Various Cistus species are known to emit volatile oils, rendering the plants flammable. Their emission of these essential oils
570-430: A single gene in an ancestral organism. In contrast, a soft polytomy stems from branches on gene trees of finite temporal duration but for which no substitutions have occurred. As DNA sequence evolution is usually much faster than evolution of complex phenotypic traits, it may be that genetic lineages diverge a short time apart from each other, while the actual organism has not changed if the whole ancestral population
627-404: A true divergence event of three or more lineages. Interpretations for a polytomy depend on the individuals that are represented in the phylogenetic tree. If the lineages in the phylogenetic tree stand for species, a polytomy shows the simultaneous speciation of three or more species. In particular situations, they may be common, for example when a species that has rapidly expanded its range or
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#1732851630928684-508: Is a genetic adaptation for the species intended to actually promote flammability and even cause spontaneous combustion. Cistus plants are therefore considered pyrophytes , or plants which rely upon fire as a part of their life cycle to encourage spread within their biome. Cistus seeds are resistant to the fires allowing them to germinate once fires have subsided and allowing them to benefit from an environment with less competition from other species. Cistuses are suitable for sunny gardens with
741-766: Is also found in Bulgaria near the Greek-Bulgarian border. Greek is also spoken worldwide by the sizable Greek diaspora which has notable communities in the United States , Australia , Canada , South Africa , Chile , Brazil , Argentina , Russia , Ukraine , the United Kingdom , and throughout the European Union , especially in Germany . Historically, significant Greek-speaking communities and regions were found throughout
798-474: Is an Indo-European language, but also includes a number of borrowings from the languages of the populations that inhabited Greece before the arrival of Proto-Greeks, some documented in Mycenaean texts ; they include a large number of Greek toponyms . The form and meaning of many words have changed. Loanwords (words of foreign origin) have entered the language, mainly from Latin, Venetian , and Turkish . During
855-471: Is commonly encountered as a garden flower. The common name rockrose ( rock rose in the UK) is applied to the species, a name also shared by the related genera Helianthemum and Tuberaria , all in the family Cistaceae. The common name gum cistus is applied to resin-bearing species, especially C. ladanifer . The leaves are evergreen , opposite, simple, usually slightly rough-surfaced, 2–8 cm long. In
912-501: Is considered. Since few if any individuals in a population are genetically alike in any one population – especially if lineage sorting has not widely progressed – it may be that hard polytomies are indeed rare or nonexistent if the entire genome of each individual organism is considered, but rather widespread on the population genetical level if entire species are considered as interbreeding populations (see also species concept ). "Speciation or lineage divergence events occurring at
969-514: Is highly panmictic undergoes peripatric speciation in different regions. An example is the Drosophila simulans species complex . Here, the ancestor seems to have colonized two islands at the same time but independently, yielding two equally old but divergently evolved daughter species If a phylogenetic tree is reconstructed from DNA sequence data of a particular gene, a hard polytomy arises when three or more sampled genes trace their ancestry to
1026-415: Is matched only by the now-extinct Anatolian languages . The Greek language is conventionally divided into the following periods: In the modern era, the Greek language entered a state of diglossia : the coexistence of vernacular and archaizing written forms of the language. What came to be known as the Greek language question was a polarization between two competing varieties of Modern Greek: Dimotiki ,
1083-422: Is peculiar in that it has developed a range of specific adaptations to resist summer drought and frequent disturbance events, such as fire and grazing. In addition, it can form both ectomycorrhizas and arbuscular mycorrhizas . More than 200 ectomycorrhiza-forming fungal species belonging to 40 genera have been reported so far to be associated with Cistus . As with many other Cistaceae, the species of Cistus have
1140-510: Is protected and promoted officially as a regional and minority language in Armenia, Hungary , Romania, and Ukraine. It is recognized as a minority language and protected in Turkey by the 1923 Treaty of Lausanne . The phonology , morphology , syntax , and vocabulary of the language show both conservative and innovative tendencies across the entire attestation of the language from the ancient to
1197-842: Is sometimes called aljamiado , as when Romance languages are written in the Arabic alphabet. Article 1 of the Universal Declaration of Human Rights in Greek: Transcription of the example text into Latin alphabet : Article 1 of the Universal Declaration of Human Rights in English: Proto-Greek Mycenaean Ancient Koine Medieval Modern Polytomy Two types of polytomies are recognized, soft and hard polytomies. Soft polytomies are
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#17328516309281254-758: Is still used internationally for the writing of Ancient Greek . In Greek, the question mark is written as the English semicolon, while the functions of the colon and semicolon are performed by a raised point (•), known as the ano teleia ( άνω τελεία ). In Greek the comma also functions as a silent letter in a handful of Greek words, principally distinguishing ό,τι ( ó,ti , 'whatever') from ότι ( óti , 'that'). Ancient Greek texts often used scriptio continua ('continuous writing'), which means that ancient authors and scribes would write word after word with no spaces or punctuation between words to differentiate or mark boundaries. Boustrophedon , or bi-directional text,
1311-490: Is weak support for a clade consisting of C. clusii and C. munbyi ; the other species either formed part of a polytomy or resolved differently in different analyses. Halimium and Cistus were regularly shown to be paraphyletic with respect to one another. The following species are recognised in the genus Cistus : In addition a large number of hybrids have been recorded, including: They are thermophilous plants, which require open, sunny places. This plant genus
1368-650: The Balkans , Caucasus , the Black Sea coast, Asia Minor , and the Eastern Mediterranean . It has the longest documented history of any Indo-European language, spanning at least 3,400 years of written records. Its writing system is the Greek alphabet , which has been used for approximately 2,800 years; previously, Greek was recorded in writing systems such as Linear B and the Cypriot syllabary . The alphabet arose from
1425-609: The Byzantine Empire and developed into Medieval Greek . In its modern form , Greek is the official language of Greece and Cyprus and one of the 24 official languages of the European Union . It is spoken by at least 13.5 million people today in Greece, Cyprus, Italy, Albania, Turkey , and the many other countries of the Greek diaspora . Greek roots have been widely used for centuries and continue to be widely used to coin new words in other languages; Greek and Latin are
1482-550: The Christian Bible was also originally written in Greek. Together with the Latin texts and traditions of the Roman world , the Greek texts and Greek societies of antiquity constitute the objects of study of the discipline of Classics . During antiquity , Greek was by far the most widely spoken lingua franca in the Mediterranean world . It eventually became the official language of
1539-486: The Eastern Mediterranean , in what are today Southern Italy , Turkey , Cyprus , Syria , Lebanon , Israel , Palestine , Egypt , and Libya ; in the area of the Black Sea , in what are today Turkey, Bulgaria , Romania , Ukraine , Russia , Georgia , Armenia , and Azerbaijan ; and, to a lesser extent, in the Western Mediterranean in and around colonies such as Massalia , Monoikos , and Mainake . It
1596-510: The Indo-Iranian languages (see Graeco-Aryan ), but little definitive evidence has been found. In addition, Albanian has also been considered somewhat related to Greek and Armenian, and it has been proposed that they all form a higher-order subgroup along with other extinct languages of the ancient Balkans; this higher-order subgroup is usually termed Palaeo-Balkan , and Greek has a central position in it. Linear B , attested as early as
1653-641: The Phoenician script and was in turn the basis of the Latin , Cyrillic , Coptic , Gothic , and many other writing systems. The Greek language holds a very important place in the history of the Western world. Beginning with the epics of Homer , ancient Greek literature includes many works of lasting importance in the European canon . Greek is also the language in which many of the foundational texts in science and philosophy were originally composed. The New Testament of
1710-672: The Royal Horticultural Society 's Award of Garden Merit ) include: Greek language Greek ( Modern Greek : Ελληνικά , romanized : Elliniká , [eliniˈka] ; Ancient Greek : Ἑλληνική , romanized : Hellēnikḗ ) is an Indo-European language, constituting an independent Hellenic branch within the Indo-European language family. It is native to Greece , Cyprus , Italy (in Calabria and Salento ), southern Albania , and other regions of
1767-492: The nominal and verbal systems. The major change in the nominal morphology since the classical stage was the disuse of the dative case (its functions being largely taken over by the genitive ). The verbal system has lost the infinitive , the synthetically -formed future, and perfect tenses and the optative mood . Many have been replaced by periphrastic ( analytical ) forms. Pronouns show distinctions in person (1st, 2nd, and 3rd), number (singular, dual , and plural in
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1824-486: The Greek alphabet since approximately the 9th century BC. It was created by modifying the Phoenician alphabet , with the innovation of adopting certain letters to represent the vowels. The variant of the alphabet in use today is essentially the late Ionic variant, introduced for writing classical Attic in 403 BC. In classical Greek, as in classical Latin, only upper-case letters existed. The lower-case Greek letters were developed much later by medieval scribes to permit
1881-491: The Greek language was the Cypriot syllabary (also a descendant of Linear A via the intermediate Cypro-Minoan syllabary ), which is closely related to Linear B but uses somewhat different syllabic conventions to represent phoneme sequences. The Cypriot syllabary is attested in Cyprus from the 11th century BC until its gradual abandonment in the late Classical period, in favor of the standard Greek alphabet. Greek has been written in
1938-629: The Greek verb have likewise remained largely the same over the course of the language's history but with significant changes in the number of distinctions within each category and their morphological expression. Greek verbs have synthetic inflectional forms for: Many aspects of the syntax of Greek have remained constant: verbs agree with their subject only, the use of the surviving cases is largely intact (nominative for subjects and predicates, accusative for objects of most verbs and many prepositions, genitive for possessors), articles precede nouns, adpositions are largely prepositional, relative clauses follow
1995-661: The Greek-Albanian border. A significant percentage of Albania's population has knowledge of the Greek language due in part to the Albanian wave of immigration to Greece in the 1980s and '90s and the Greek community in the country. Prior to the Greco-Turkish War and the resulting population exchange in 1923 a very large population of Greek-speakers also existed in Turkey , though very few remain today. A small Greek-speaking community
2052-410: The ability to form mycorrhizal associations with truffles ( Tuber ) and are thus able to thrive on poor sandy soils or rocks. Cistus ladanifer has been found to have mycorrhizal associations with Boletus edulis , Boletus rhodoxanthus , and Laccaria laccata . Cistus are the only host of Cytinus hypocistis , a small parasitic plant that lives on the roots and is noticeable only for
2109-404: The acute during the late 20th century, and it has only been retained in typography . After the writing reform of 1982, most diacritics are no longer used. Since then, Greek has been written mostly in the simplified monotonic orthography (or monotonic system), which employs only the acute accent and the diaeresis. The traditional system, now called the polytonic orthography (or polytonic system),
2166-402: The ancient language; singular and plural alone in later stages), and gender (masculine, feminine, and neuter), and decline for case (from six cases in the earliest forms attested to four in the modern language). Nouns, articles, and adjectives show all the distinctions except for a person. Both attributive and predicative adjectives agree with the noun. The inflectional categories of
2223-1145: The flower colour of C. parviflorus is anomalous, it has very short styles , otherwise characteristic of WWPC species. A hybrid origin has been suggested. A simplified cladogram is shown below: Halimium spp. Halimium spp. Cistus crispus Cistus heterophyllus Cistus albidus Cistus creticus Cistus asper Cistus chinamadensis Cistus horrens Cistus ocreatus Cistus osbeckiifolius Cistus palmensis Cistus symphytifolius Cistus clusii Cistus munbyi Cistus inflatus Cistus ladanifer Cistus laurifolius Cistus libanotis Cistus monspeliensis Cistus parviflorus Cistus populifolius Cistus pouzolzii Cistus salviifolius Cistus sintenisii Within
2280-439: The infinitive entirely (employing a raft of new periphrastic constructions instead) and uses participles more restrictively. The loss of the dative led to a rise of prepositional indirect objects (and the use of the genitive to directly mark these as well). Ancient Greek tended to be verb-final, but neutral word order in the modern language is VSO or SVO. Modern Greek inherits most of its vocabulary from Ancient Greek, which in turn
2337-463: The late 15th century BC, was the first script used to write Greek. It is basically a syllabary , which was finally deciphered by Michael Ventris and John Chadwick in the 1950s (its precursor, Linear A , has not been deciphered and most likely encodes a non-Greek language). The language of the Linear B texts, Mycenaean Greek , is the earliest known form of Greek. Another similar system used to write
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2394-564: The membership of Greece and Cyprus in the European Union, Greek is one of the organization's 24 official languages . Greek is recognized as a minority language in Albania, and used co-officially in some of its municipalities, in the districts of Gjirokastër and Sarandë . It is also an official minority language in the regions of Apulia and Calabria in Italy. In the framework of the European Charter for Regional or Minority Languages , Greek
2451-441: The modern period. The division into conventional periods is, as with all such periodizations, relatively arbitrary, especially because, in all periods, Ancient Greek has enjoyed high prestige, and the literate borrowed heavily from it. Across its history, the syllabic structure of Greek has varied little: Greek shows a mixed syllable structure, permitting complex syllabic onsets but very restricted codas. It has only oral vowels and
2508-399: The noun they modify and relative pronouns are clause-initial. However, the morphological changes also have their counterparts in the syntax, and there are also significant differences between the syntax of the ancient and that of the modern form of the language . Ancient Greek made great use of participial constructions and of constructions involving the infinitive, and the modern variety lacks
2565-706: The older periods of Greek, loanwords into Greek acquired Greek inflections, thus leaving only a foreign root word. Modern borrowings (from the 20th century on), especially from French and English, are typically not inflected; other modern borrowings are derived from Albanian , South Slavic ( Macedonian / Bulgarian ) and Eastern Romance languages ( Aromanian and Megleno-Romanian ). Greek words have been widely borrowed into other languages, including English. Example words include: mathematics , physics , astronomy , democracy , philosophy , athletics , theatre, rhetoric , baptism , evangelist , etc. Moreover, Greek words and word elements continue to be productive as
2622-532: The predominant sources of international scientific vocabulary . Greek has been spoken in the Balkan peninsula since around the 3rd millennium BC, or possibly earlier. The earliest written evidence is a Linear B clay tablet found in Messenia that dates to between 1450 and 1350 BC, making Greek the world's oldest recorded living language . Among the Indo-European languages, its date of earliest written attestation
2679-469: The purple pink clade (PPC), C. crispus is consistently the first diverging species. C. albidus , C. creticus and C. heterophyllus form a well supported clade. Seven species endemic to the Canary Islands form a polytomy , resolved differently in different analyses, in which subtaxa of some species do not always cluster together. Within the white and whitish pink clade (WWPC), there
2736-406: The result of insufficient phylogenetic information: though the lineages diverged at different times – meaning that some of these lineages are closer relatives than others, and the available data does not allow recognition of this. Most polytomies are soft, meaning that they would be resolved into a typical tree of dichotomies if better data were available. In contrast, a hard polytomy represents
2793-415: The same time" refers to evolutionary time measured in generations , as this is the only means that novel traits (e.g. germline point mutations ) can be passed on. In practical terms, the ability to distinguish between hard and soft polytomies is limited: if for example a kilobase of DNA sequences that mutate approximately 1% per million years is analyzed, lineages diverging from the same ancestor within
2850-437: The stressed vowel; the so-called breathing marks ( rough and smooth breathing ), originally used to signal presence or absence of word-initial /h/; and the diaeresis , used to mark the full syllabic value of a vowel that would otherwise be read as part of a diphthong. These marks were introduced during the course of the Hellenistic period. Actual usage of the grave in handwriting saw a rapid decline in favor of uniform usage of
2907-447: The various stages of the Greek language are often emphasized. Although Greek has undergone morphological and phonological changes comparable to those seen in other languages, never since classical antiquity has its cultural, literary, and orthographic tradition been interrupted to the extent that one can speak of a new language emerging. Greek speakers today still tend to regard literary works of ancient Greek as part of their own rather than
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#17328516309282964-529: The vernacular form of Modern Greek proper, and Katharevousa , meaning 'purified', a compromise between Dimotiki and Ancient Greek developed in the early 19th century that was used for literary and official purposes in the newly formed Greek state. In 1976, Dimotiki was declared the official language of Greece, after having incorporated features of Katharevousa and thus giving birth to Standard Modern Greek , used today for all official purposes and in education . The historical unity and continuing identity between
3021-568: Was a distinct dialect of Greek itself. Aside from the Macedonian question, current consensus regards Phrygian as the closest relative of Greek, since they share a number of phonological, morphological and lexical isoglosses , with some being exclusive between them. Scholars have proposed a Graeco-Phrygian subgroup out of which Greek and Phrygian originated. Among living languages, some Indo-Europeanists suggest that Greek may be most closely related to Armenian (see Graeco-Armenian ) or
3078-549: Was also used as the official language of government and religion in the Christian Nubian kingdoms , for most of their history. Greek, in its modern form, is the official language of Greece, where it is spoken by almost the entire population. It is also the official language of Cyprus (nominally alongside Turkish ) and the British Overseas Territory of Akrotiri and Dhekelia (alongside English ). Because of
3135-546: Was also used in Ancient Greek. Greek has occasionally been written in the Latin script , especially in areas under Venetian rule or by Greek Catholics . The term Frankolevantinika / Φραγκολεβαντίνικα applies when the Latin script is used to write Greek in the cultural ambit of Catholicism (because Frankos / Φράγκος is an older Greek term for West-European dating to when most of (Roman Catholic Christian) West Europe
3192-566: Was under the control of the Frankish Empire ). Frankochiotika / Φραγκοχιώτικα (meaning 'Catholic Chiot') alludes to the significant presence of Catholic missionaries based on the island of Chios . Additionally, the term Greeklish is often used when the Greek language is written in a Latin script in online communications. The Latin script is nowadays used by the Greek-speaking communities of Southern Italy . The Yevanic dialect
3249-653: Was written by Romaniote and Constantinopolitan Karaite Jews using the Hebrew Alphabet . In a tradition, that in modern time, has come to be known as Greek Aljamiado , some Greek Muslims from Crete wrote their Cretan Greek in the Arabic alphabet . The same happened among Epirote Muslims in Ioannina . This also happened among Arabic-speaking Byzantine rite Christians in the Levant ( Lebanon , Palestine, and Syria ). This usage
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