186-721: See text , traditionally 1, but possibly up to 6 The reindeer or caribou ( Rangifer tarandus ) is a species of deer with circumpolar distribution , native to Arctic , subarctic , tundra , boreal , and mountainous regions of Northern Europe, Siberia, and North America. It is the only representative of the genus Rangifer . More recent studies suggest the splitting of reindeer and caribou into six distinct species over their range. Reindeer occur in both migratory and sedentary populations, and their herd sizes vary greatly in different regions. The tundra subspecies are adapted for extreme cold, and some are adapted for long-distance migration. Reindeer vary greatly in size and color from
372-561: A Dene (Athapascan) group, call the Arctic caribou Ɂekwǫ̀ and the boreal woodland caribou tǫdzı . The Gwichʼin (also a Dene group) have over 24 distinct caribou-related words. Reindeer are also called tuttu by the Greenlandic Inuit and hreindýr , sometimes rein , by the Icelanders . The "glacial-interglacial cycles of the upper Pleistocene had a major influence on
558-456: A boreal forest hosting a species assemblage with no modern analogue. These are among the oldest DNA fragments ever sequenced. Carl Linnaeus in 1758 named the Eurasian tundra species Cervus tarandus , the genus Rangifer being credited to Smith, 1827. Rangifer has had a convoluted history because of the similarity in antler architecture (brow tines asymmetrical and often palmate, bez tines,
744-606: A clade sister to Cervidae. According to the study, Cervidae diverged from the Bovidae-Moschidae clade 27 to 28 million years ago. The following cladogram is based on the 2003 study. Tragulidae [REDACTED] Antilocapridae [REDACTED] Giraffidae [REDACTED] Cervidae [REDACTED] Bovidae [REDACTED] Svalbard Svalbard ( / ˈ s v ɑː l b ɑːr ( d )/ SVAHL -bar(d) , Urban East Norwegian: [ˈsvɑ̂ːɫbɑr] ), previously known as Spitsbergen or Spitzbergen ,
930-508: A demilitarized zone , as the treaty prohibits the establishment of military installations on the islands. However, since the treaty recognizes Norway as the sovereign power in the archipelago, the country claims exclusive rights in the maritime zone around the islands; rights which Norway argues permit the Norwegian Coast Guard to conduct fishery and other maritime surveillance and enforcement in these waters. Certain other parties to
1116-719: A liver without a gallbladder . Deer also have a tapetum lucidum , which gives them sufficiently good night vision . All male deer have antlers , with the exception of the water deer , in which males have long tusk-like canines that reach below the lower jaw. Females generally lack antlers, though female reindeer bear antlers smaller and less branched than those of the males. Occasionally females in other species may develop antlers, especially in telemetacarpal deer such as European roe deer, red deer, white-tailed deer and mule deer and less often in plesiometacarpal deer. A study of antlered female white-tailed deer noted that antlers tend to be small and malformed, and are shed frequently around
1302-455: A Captain Craycott had brought a live pair from Greenland to England in 1738. He named it Capra groenlandicus , Greenland reindeer. Linnaeus, in the 12th edition of Systema naturae , gave grœnlandicus as a synonym for Cervus tarandus . Borowski disagreed (and again changed the spelling), saying Cervus grönlandicus was morphologically distinct from Eurasian tundra reindeer. Baird placed it under
1488-519: A Russian coal mining company, owns 0.4%, while other private owners hold 0.4%. As Svalbard is north of the Arctic Circle , it experiences midnight sun in summer and polar night in winter. At 74° north, the midnight sun lasts 99 days and polar night 84 days, while the respective figures at 81° north are 141 and 128 days. In Longyearbyen, midnight sun lasts from 20 April until 23 August, and polar night lasts from 26 October to 15 February. In winter,
1674-543: A Russian crew for an expedition in 1795. After the Anglo-Russian War in 1809, Russian activity on Svalbard diminished, and had ceased by the 1820s. Norwegian whaling was abandoned about the same time as the Russians left, but whaling continued around Spitsbergen until the 1830s, and around Bjørnøya until the 1860s. By the 1890s, Svalbard had become a destination for Arctic tourism , coal deposits had been found, and
1860-448: A back tine sometimes branched, and branched at the distal end, often palmate). Because of individual variability, early taxonomists were unable to discern consistent patterns among populations, nor could they, examining collections in Europe, appreciate the difference in habitats and the differing function they imposed on antler architecture. Comparative morphometrics, the measurement of skulls,
2046-406: A broad central portion), white-tailed deer antlers include a series of tines sprouting upward from a forward-curving main beam, and those of the pudú are mere spikes. Antler development begins from the pedicel, a bony structure that appears on the top of the skull by the time the animal is a year old. The pedicel gives rise to a spiky antler the following year, that is replaced by a branched antler in
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#17328439570682232-775: A broad, high muzzle to increase the volume of the nasal cavity to warm and moisten the air before it enters the throat and lungs, bez tines set close to the brow tines, distinctive coat patterns, short legs and other adaptations for running long distances, and multiple behaviors suited to tundra, but not to forest (such as synchronized calving and aggregation during rutting and post-calving). As well, many genes, including those for vitamin D metabolism, fat metabolism , retinal development, circadian rhythm , and tolerance to cold temperatures, are found in tundra caribou that are lacking or rudimentary in forest types. For this reason, forest-adapted reindeer and caribou could not survive in tundra or polar deserts . The oldest undoubted Rangifer fossil
2418-439: A direct common ancestor , they cannot be Biological specificity#conspecific|conspecific. Similarly, woodland caribou diverged from the ancestors of Arctic caribou before modern barren-ground caribou had evolved, and were more likely related to extinct North American forest reindeer (see Evolution above). Lacking a direct shared ancestor, barren-ground and woodland caribou cannot be conspecific. Molecular data also revealed that
2604-453: A disease which in the UK in 2005 cost £90 million in attempts to eradicate. In New Zealand, deer are thought to be important as vectors picking up M. bovis in areas where brushtail possums Trichosurus vulpecula are infected, and transferring it to previously uninfected possums when their carcasses are scavenged elsewhere. The white-tailed deer Odocoileus virginianus has been confirmed as
2790-592: A forest subspecies, formerly included reindeer west of the Sea of Okhotsk which, however, are indistinguishable genetically from the Jano-Indigirka, East Siberian taiga and Chukotka populations of R. t. sibiricus . Siberian tundra reindeer herds have been in decline but are stable or increasing since 2000. Insular (island) reindeer, classified as the Novaya Zemlya reindeer ( R. t. pearsoni ) occupy several island groups:
2976-726: A full part of the Kingdom of Norway. The Svalbard Treaty established Svalbard as a free economic zone and restricts the military use of the archipelago. The Norwegian Store Norske and the Russian Arktikugol remain the only mining companies in place. Research and tourism have become important supplementary industries, with the University Centre in Svalbard and the Svalbard Global Seed Vault playing critical roles in
3162-546: A garrison in the islands, which was done in May 1942 during Operation Fritham . Meanwhile, the Germans responded to the destruction of the weather station by establishing a reporting station of their own, codenamed "Banso" , in October 1941. They were chased away in October by a visit from what the Germans mistook to be four British warships, but later returned. A second station, "Knospe",
3348-425: A large scale in the world. Both wild and domestic reindeer have been an important source of food, clothing, and shelter for Arctic people from prehistorical times. They are still herded and hunted today. In some traditional Christmas legends, Santa Claus's reindeer pull a sleigh through the night sky to help Santa Claus deliver gifts to good children on Christmas Eve. Names follow international convention before
3534-426: A maximum speed is 22 knots (41 km/h; 25 mph) with more than 60 days endurance and the complement is up to 100 people. The first ship, KV Jan Mayen , was delivered in early 2023. These vessels will complement NoCGV Svalbard which predominantly serves Svalbard and the surrounding waters. In 2023, Norway also announced the acquisition of six MH-60R helicopters which are to be initially deployed with
3720-698: A mtDNA haplotype with Labrador caribou, in the North American lineage (i.e., woodland caribou). Røed et al. (1991) had noted: Among Baffin Island caribou the TFL2 allele was the most common allele (p=0.521), while this allele was absent, or present in very low frequencies, in other caribou populations (Table 1), including the Canadian barren-ground caribou from the Beverly herd. A large genetic difference between Baffin Island caribou and
3906-546: A nearly complete skeleton of Diacodexis discovered in 1982 gave rise to speculation that this ancestor could be closer to the non-ruminants than the ruminants. Andromeryx is another prominent prehistoric ruminant, but appears to be closer to the tragulids . The formation of the Himalayas and the Alps brought about significant geographic changes. This was the chief reason behind the extensive diversification of deer-like forms and
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#17328439570684092-785: A new revision of the genus. Abbreviations: The table above includes, as per the recent revision, R. t. caboti (the Labrador caribou (the Eastern Migratory population DU4)), and R. t. terranovae (the Newfoundland caribou (the Newfoundland population DU5)), which molecular analyses have shown to be of North American (i.e., woodland caribou) lineage; and four mountain ecotypes now known to be of distant Beringia - Eurasia lineage (see Taxonomy above). The scientific name Tarandus rangifer buskensis Millais, 1915 (the Busk Mountains reindeer)
4278-437: A predator-avoidance strategy, which requires large rutting aggregations. Males cannot defend a harem because, while he was busy fighting, they would disappear into the mass of the herd. Males therefore tend individual females; their fights are infrequent and brief. Their antlers are thin, beams round in cross-section, sweep back and then forward with a cluster of branches at the top; these are designed more for visual stimulation of
4464-543: A result of acclimatisation society releases in the 19th century, Australia has six introduced species of deer that have established sustainable wild populations. They are fallow deer, red deer, sambar, hog deer, rusa , and chital. Red deer were introduced into New Zealand in 1851 from English and Scottish stock. Many have been domesticated in deer farms since the late 1960s and are common farm animals there now. Seven other species of deer were introduced into New Zealand but none are as widespread as red deer. Deer constitute
4650-533: A scathing review by Ian McTaggart-Cowan in 1962. Most authorities continued to consider all or most subspecies valid; some were quite distinct. In his chapter in the authoritative 2005 reference work Mammal Species of the World , referenced by the American Society of Mammalogists , English zoologist Peter Grubb agreed with Valerius Geist , a specialist on large mammals, that these subspecies were valid (i.e., before
4836-522: A separate policy called "Regulations relating to rejection and expulsion of persons from Svalbard". Among the requirements is that residents must have the means to be able to reside on Svalbard. These requirements apply to both foreigners and Norwegian citizens, and the Governor of Svalbard may reject persons who do not meet the requirements. Russia retains a consulate in Barentsburg . In September 2010,
5022-485: A smaller form, but a combination of anthropogenic and climatic pressures is now thought to be the most likely culprit. Meanwhile, the moose and reindeer radiated into North America from Siberia. Deer constitute the artiodactyl family Cervidae. This family was first described by German zoologist Georg August Goldfuss in Handbuch der Zoologie (1820). Three subfamilies were recognised: Capreolinae (first described by
5208-501: A source of income can be rejected by the governor. No one is required to have a visa or residence permit on Svalbard. Regardless of citizenship, persons can live and work in Svalbard indefinitely. The Svalbard Treaty grants treaty nationals equal right of abode as Norwegian nationals. So far, non-treaty nationals have been admitted visa-free as well. While there is no visa requirement, everyone must meet certain requirements in order to stay in Svalbard. These requirements are governed by
5394-496: A summer population of 180. The Norwegian Meteorological Institute has outposts at Bjørnøya and Hopen, with ten and four inhabitants, respectively. Both can also house temporary research staff. Poland operates the Polish Polar Station at Hornsund , with ten permanent residents. The Soviet mining settlement of Pyramiden was abandoned in 1998, leaving Barentsburg as the only permanently inhabited Russian settlement. It
5580-418: A terrestrial bird that never leaves Svalbard and many seabirds , that use Svalbard as a breeding ground. The most numerous are Little Auks, Brünnich's Guillemots (Thick-billed Murre) and Black-legged Kittiwakes who breed on the cliffs around Svalbard. The most unique ones are King Eider for their striking appearance, Ivory gulls who like to be around polar bears, Snow Buntings - the only songbird in Svalbard. It
5766-499: A treaty was signed between Russia and Norway fixing the boundary between the Svalbard archipelago and the Novaya Zemlya archipelago . Increased interest in petroleum exploration in the Arctic raised interest in a resolution of the dispute. The agreement takes into account the relative positions of the archipelagos, rather than being based simply on northward extension of the continental border of Norway and Russia. Svalbard constitutes
Reindeer - Misplaced Pages Continue
5952-751: A very interesting history. Allen (1902) named it as a distinct species, R. granti , from the "western end of Alaska Peninsula , opposite Popoff Island " and noting that: Rangifer granti is a representative of the Barren Ground group of Caribou, which includes R. arcticus of the Arctic Coast and R. granlandicus of Greenland. It is not closely related to R. stonei of the Kenai Peninsula, from which it differs not only in its very much smaller size, but in important cranial characters and in coloration. ...The external and cranial differences between R. granti and
6138-651: A view to using them as a base of operations to send supplies to north Russia, but the idea was rejected as impractical. Instead, with the agreement of the Soviets and the Norwegian government in exile, in August 1941 the Norwegian and Soviet settlements on Svalbard were evacuated, and facilities there destroyed, in Operation Gauntlet . However, the Norwegian government in exile decided it would be important politically to establish
6324-783: A wide variety of vegetation. The teeth of deer are adapted to feeding on vegetation, and like other ruminants, they lack upper incisors , instead having a tough pad at the front of their upper jaw. Deer are browsers , and feed primarily on foliage of grasses , sedges , forbs , shrubs and trees , secondarily on lichens in northern latitudes during winter. They have small, unspecialized stomachs by ruminant standards, and high nutrition requirements. Rather than eating and digesting vast quantities of low-grade fibrous food as, for example, sheep and cattle do, deer select easily digestible shoots, young leaves, fresh grasses, soft twigs, fruit, fungi , and lichens . The low-fibered food, after minimal fermentation and shredding, passes rapidly through
6510-492: A wild animal of any kind. Cognates of Old English dēor in other dead Germanic languages have the general sense of animal , such as Old High German tior , Old Norse djur or dȳr , Gothic dius , Old Saxon dier , and Old Frisian diar . This general sense gave way to the modern English sense by the end of the Middle English period, around 1500. All modern Germanic languages save English and Scots retain
6696-499: Is Wijdefjorden (108 km or 67 mi), followed by Isfjorden (107 km or 66 mi), Van Mijenfjorden (83 km or 52 mi), Woodfjorden (64 km or 40 mi), and Wahlenbergfjorden (46 km or 29 mi). Svalbard is part of the High Arctic Large Igneous Province , and experienced Norway's strongest earthquake on 6 March 2009 at magnitude 6.5. The Dutchman Willem Barentsz made
6882-510: Is a Norwegian archipelago in the Arctic Ocean . North of mainland Europe , it lies about midway between the northern coast of Norway and the North Pole . The islands of the group range from 74° to 81° north latitude, and from 10° to 35° east longitude. The largest island is Spitsbergen , followed in size by Nordaustlandet and Edgeøya . The largest settlement is Longyearbyen on
7068-487: Is a company town: all facilities are owned by Arktikugol, which operates a coal mine. In addition to the mining facilities, Arktikugol has opened a hotel and souvenir shop, catering to tourists taking day trips or hikes from Longyearbyen. The village features a school, library, sports center, community center, swimming pool, farm, and greenhouse. Pyramiden features similar facilities; both are built in typical post-World War II Soviet architectural and planning style and contain
7254-509: Is a hoofed ruminant ungulate of the family Cervidae (informally the deer family ). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac , elk (wapiti), red deer , and fallow deer ) and Capreolinae (which includes, among others reindeer (caribou), white-tailed deer , roe deer , and moose ). Male deer of almost all species (except the water deer ), as well as female reindeer, grow and shed new antlers each year. These antlers are bony extensions of
7440-416: Is a permanent research settlement in the northwest of Spitsbergen and the northernmost functional civilian settlement in the world. Formerly a mining town, it is still a company town operated by the Norwegian state-owned Kings Bay company. While some tourism to the outpost is permitted, Norwegian authorities limit access to minimize impact on scientific work. Ny-Ålesund has a winter population of 35 and
7626-508: Is from Latin : cervus , meaning ' stag ' or ' deer ' . Deer live in a variety of biomes , ranging from tundra to the tropical rainforest . While often associated with forests, many deer are ecotone species that live in transitional areas between forests and thickets (for cover) and prairie and savanna (open space). The majority of large deer species inhabit temperate mixed deciduous forest, mountain mixed coniferous forest, tropical seasonal/dry forest, and savanna habitats around
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7812-564: Is from Omsk , Russia, dated to 2.1-1.8 Ma. The oldest North American Rangifer fossil is from the Yukon , 1.6 million years before present (BP). A fossil skull fragment from Süßenborn, Germany, R. arcticus stadelmanni , (which is probably misnamed) with "rather thin and cylinder-shaped" antlers, dates to the Middle Pleistocene (Günz) Period, 680,000-620,000 BP. Rangifer fossils become increasingly frequent in circumpolar deposits beginning with
7998-533: Is home to polar bears , reindeer , the Arctic fox , and certain marine mammals. Seven national parks and 23 nature-reserves cover two-thirds of the archipelago, protecting the largely untouched fragile environment. Norway announced new regulations regarding tourism in February 2024, including a maximum of 200 people on a ship, to protect flora and fauna in Svalbard. While part of the Kingdom of Norway since 1925, Svalbard
8184-471: Is necessary with R. montanus or with any of the woodland forms." Osgood and Murie (1935), agreeing with granti ' s close relationship with the barren-ground caribou, brought it under R. arcticus as a subspecies, R. t. granti . Anderson (1946) and Banfield (1961), based on statistical analysis of cranial, dental and other characters, agreed. But Banfield (1961) also synonymized Alaska's large R. stonei with other mountain caribou of British Columbia and
8370-463: Is not governed by Norway's policies on migration and does not issue visas or residence permits itself. Foreigners do not need a visa or work and residence permits from the Norwegian authorities to travel to Svalbard. However, foreign citizens with a visa requirement for the Schengen Area must have a Schengen visa when travelling to and from Svalbard via mainland Norway. The Svalbard Act established
8556-705: Is not part of geographical Norway; administratively, the archipelago is not part of any Norwegian county , but forms an unincorporated area . This means that it is administered directly by the Norwegian government through an appointed governor , and is a special jurisdiction subject to the Svalbard Treaty that is outside of the Schengen Area , the Nordic Passport Union , and the European Economic Area . Svalbard and Jan Mayen are collectively assigned
8742-662: Is often seen as more objective than description of differences of color or antler patterns, but actually confounds genetic variance with epistatic and statistical variance as well as compounded environment-based variance. For example, woodland caribou males, rutting in boreal forest where only a few females can be found, collect harems and defend them against other males, for which they have short, straight, strong, much-branched antlers, beams flattened in cross-section, designed for combat — and not too large, so as not to impede them in forested winter ranges. By contrast, modern tundra caribou (see Evolution above) have synchronized calving as
8928-452: Is only slightly taller and heavier. Sexual dimorphism is quite pronounced – in most species males tend to be larger than females, and, except for the reindeer, only males have antlers. Coat colour generally varies between red and brown, though it can be as dark as chocolate brown in the tufted deer or have a grayish tinge as in elk. Different species of brocket deer vary from gray to reddish brown in coat colour. Several species such as
9114-659: Is part of University Hospital of North Norway , while the airport is operated by state-owned Avinor . Ny-Ålesund and Barentsburg remain company towns with all infrastructure owned by Kings Bay and Arktikugol. Other public offices with presence on Svalbard are the Norwegian Directorate of Mining , the Norwegian Polar Institute , the Norwegian Tax Administration , and the Church of Norway . Svalbard
9300-623: Is possible to ski from Sørkapp in the south to the north of Spitsbergen, with only a short distance not being covered by snow or glacier. Kvitøya is 99.3% covered by glacier. The landforms of Svalbard were created through repeated ice ages , when glaciers cut the former plateau into fjords, valleys, and mountains. The tallest peak is Newtontoppen (1,717 m or 5,633 ft), followed by Perriertoppen (1,712 m or 5,617 ft), Ceresfjellet (1,675 m or 5,495 ft), Chadwickryggen (1,640 m or 5,380 ft), and Galileotoppen (1,637 m or 5,371 ft). The longest fjord
9486-412: Is strong enough to walk with its mother. The fawn and its mother stay together for about one year. A male usually leaves and never sees his mother again, but females sometimes come back with their own fawns and form small herds. In some areas of the UK, deer (especially fallow deer due to their gregarious behaviour ) have been implicated as a possible reservoir for transmission of bovine tuberculosis ,
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#17328439570689672-585: Is subordinate to Nord-Troms District Court and Hålogaland Court of Appeal , both in Tromsø . Although Norway is part of the European Economic Area (EEA) and the Schengen Agreement , Svalbard is not part of the Schengen Area or the EEA. Non-EU and non-Nordic Svalbard residents do not need Schengen visas for Svalbard itself, but those travelling via mainland Norway require visas to pass through Norway. People without
9858-501: Is the moose , which is nearly 2.6 metres (8 ft 6 in) tall and weighs up to 800 kilograms (1,800 lb). The elk stands 1.4–2 metres (4 ft 7 in – 6 ft 7 in) at the shoulder and weighs 240–450 kilograms (530–990 lb). The northern pudu is the smallest deer in the world; it reaches merely 32–35 centimetres ( 12 + 1 ⁄ 2 –14 in) at the shoulder and weighs 3.3–6 kilograms ( 7 + 1 ⁄ 4 – 13 + 1 ⁄ 4 lb). The southern pudu
10044-629: The Barbary stag , a subspecies of red deer that is confined to the Atlas Mountains in the northwest of the continent. Another extinct species of deer, Megaceroides algericus , was present in North Africa until 6000 years ago. Fallow deer have been introduced to South Africa . Small species of brocket deer and pudús of Central and South America , and muntjacs of Asia generally occupy dense forests and are less often seen in open spaces, with
10230-487: The Caucasus Mountains have forest areas that are not only home to sizable deer populations but also other animals that were once abundant such as the wisent, Eurasian lynx , Iberian lynx , wolves , and brown bears . The highest concentration of large deer species in temperate Asia occurs in the mixed deciduous forests, mountain coniferous forests, and taiga bordering North Korea, Manchuria (Northeastern China), and
10416-573: The Early Pleistocene (2 million years ago) Kap Kobenhavn Formation of northern Greenland identified preserved DNA fragments of Rangifer , identified as basal but potentially ancestral to modern reindeer. This suggests that reindeer have inhabited Greenland since at least the Early Pleistocene. Around this time, northern Greenland was 11–19 °C (20–34 °F) warmer than the Holocene , with
10602-588: The European Space Research Organisation . Petroleum test drilling was started in 1963 and continued until 1984, but no commercially viable fields were found. From 1960, regular charter flights were made from the mainland to a field at Hotellneset ; in 1975, Svalbard Airport, Longyearbyen opened, allowing year-round services. During the Cold War , the Soviet Union comprised about two-thirds of
10788-532: The Heerodden helicopter accident , which killed three people. Longyearbyen remained purely a company town until 1989 when utilities, culture, and education was separated into Svalbard Samfunnsdrift. In 1993, it was sold to the national government and the University Centre was established. Through the 1990s, tourism increased and the town developed an economy independent of Store Norske and mining. Longyearbyen
10974-455: The ISO 3166-1 alpha-2 country-code "SJ". Both areas are administered by Norway, though they are separated by a distance of over 950 kilometres (510 nautical miles) and have very different administrative structures. The name Svalbard was officially adopted for the archipelago by Norway under the 1925 Svalbard Act which formally annexed it. The former name Spitsbergen was thenceforth restricted to
11160-715: The Last Glacial Period until the present day. In the non-forested mountains of central Norway, such as Jotunheimen , it is still possible to find remains of stone-built trapping pits , guiding fences and bow rests, built especially for hunting reindeer. These can, with some certainty, be dated to the Migration Period , although it is not unlikely that they have been in use since the Stone Age . Cave paintings by ancient Europeans include both tundra and forest types of reindeer. A 2022 study of ancient environmental DNA from
11346-781: The Late Miocene , 8.7–9.6 million years ago. Rangifer "evolved as a mountain deer, ...exploiting the subalpine and alpine meadows...". Rangifer originated in the Late Pliocene and diversified in the Early Pleistocene , a 2+ million-year period of multiple glacier advances and retreats. Several named Rangifer fossils in Eurasia and North America predate the evolution of modern tundra reindeer. Archaeologists distinguish "modern" tundra reindeer and barren-ground caribou from primitive forms – living and extinct – that did not have adaptations to extreme cold and to long-distance migration. They include
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#173284395706811532-557: The Mi'kmaq qalipu , meaning "snow shoveler", and refers to its habit of pawing through the snow for food. Because of its importance to many cultures, Rangifer and some of its species and subspecies have names in many languages. Inuvialuit of the western Canadian Arctic and Inuit of the eastern Canadian Arctic, who speak different dialects of the Inuit languages , both call the barren-ground caribou tuktu . The Wekʼèezhìi ( Tłı̨chǫ ) people,
11718-689: The Novaya Zemlya Archipelago (about 5,000 animals at last count, but most of these are either domestic reindeer or domestic-wild hybrids), the New Siberia Archipelago (about 10,000 to 15,000), and Wrangel Island (200 to 300 feral domestic reindeer). What was once the second largest herd is the migratory Labrador caribou ( R. t. caboti ) George River herd in Canada, with former variations between 28,000 and 385,000. As of January 2018, there are fewer than 9,000 animals estimated to be left in
11904-668: The Orthodox Church . Catholics on the archipelago are pastorally served by the Territorial Prelature of Tromsø . The Svalbard Treaty of 1920 established full Norwegian sovereignty over the archipelago. The islands are, unlike the Norwegian Antarctic territories , a part of the Kingdom of Norway and not a dependency . The treaty came into effect in 1925, following the Svalbard Act. All forty-eight signatory countries of
12090-615: The Pleistocene have been excavated in China and the Himalayas. While Cervus and Dama appeared nearly 3 Mya, Axis emerged during the late Pliocene–Pleistocene. The tribes Capreolini and Rangiferini appeared around 4–7 Mya. Around 5 Mya, the rangiferina † Bretzia and † Eocoileus were the first cervids to reach North America. This implies the Bering Strait could be crossed during
12276-428: The Queen Charlotte Islands caribou ( R. t. dawsoni ) from western Canada, the Sakhalin reindeer ( R. t. setoni ) from Sakhalin and the East Greenland caribou from eastern Greenland, although some authorities believe that the latter, R. t. eogroenlandicus Degerbøl, 1957, is a junior synonym of the Peary caribou. Historically, the range of the sedentary boreal woodland caribou covered more than half of Canada and into
12462-411: The Riss glaciations , the second youngest of the Pleistocene Epoch, roughly 300,000–130,000 BP. By the 4-Würm period (110,000–70,000 to 12,000–10,000 BP), its European range was extensive, supplying a major food source for prehistoric Europeans. North American fossils outside of Beringia that predate the Last Glacial Maximum (LGM) are of Rancholabrean age (240,000–11,000 years BP) and occur along
12648-620: The Sami people of Finland and Scandinavia, the Tungusic peoples, Mongolians, and Turkic peoples of Southern Siberia, Northern Mongolia, and the Ussuri Region have also taken to raising semi-domesticated herds of Asian caribou. The highest concentration of large deer species in the tropics occurs in Southern Asia in India's Indo-Gangetic Plain Region and Nepal 's Terai Region. These fertile plains consist of tropical seasonal moist deciduous, dry deciduous forests, and both dry and wet savannas that are home to chital , hog deer , barasingha , Indian sambar , and Indian muntjac . Grazing species such as
12834-621: The Scandinavian mountains and R. t. sibiricus across Siberia) and east ( R. t. arcticus in the North American Barrenlands) when rising seas isolated them. Likewise in North America, DNA analysis shows that woodland caribou ( R. caribou ) diverged from primitive ancestors of tundra / barren-ground caribou not during the LGM, 26,000–19,000 years ago, as previously assumed, but in the Middle Pleistocene around 357,000 years ago. At that time, modern tundra caribou had not even evolved. Woodland caribou are likely more related to extinct North American forest caribou than to barren-ground caribou. For example,
13020-399: The Svalbard Act regulated the archipelago and the first governor , Johannes Gerckens Bassøe , took office. The archipelago has traditionally been known as Spitsbergen, and the main island as West Spitsbergen. During the 1920s, Norway renamed the archipelago Svalbard, and the main island became Spitsbergen. Kvitøya , Kong Karls Land , Hopen , and Bjørnøya were not regarded as part of
13206-421: The Sámi word raingo . Carl Linnaeus chose the word tarandus as the specific epithet, making reference to Ulisse Aldrovandi 's Quadrupedum omnium bisulcorum historia fol. 859–863, Cap. 30: De Tarando (1621). However, Aldrovandi and Conrad Gessner thought that rangifer and tarandus were two separate animals. In any case, the tarandos name goes back to Aristotle and Theophrastus . The use of
13392-526: The Tethys Ocean disappeared to give way to vast stretches of grassland; these provided the deer with abundant protein-rich vegetation that led to the development of ornamental antlers and allowed populations to flourish and colonise areas. As antlers had become pronounced, the canines were either lost or became poorly represented (as in elk), probably because diet was no longer browse -dominated and antlers were better display organs. In muntjac and tufted deer,
13578-494: The coat of arms of Åland . Their economic importance includes the use of their meat as venison , their skins as soft, strong buckskin , and their antlers as handles for knives. Deer hunting has been a popular activity since the Middle Ages and remains a resource for many families today. The word deer was originally broad in meaning, becoming more specific with time. Old English dēor and Middle English der meant
13764-641: The contiguous United States . The Committee on the Status of Endangered Wildlife in Canada (COSEWIC) classified both the Southern Mountain population DU9 ( R. t. montanus ) and the Central Mountain population DU8 ( R. t. fortidens ) as Endangered and the Northern Mountain population DU7 ( R. t. osborni ) as Threatened. Some species and subspecies are rare and three subspecies have already become extinct:
13950-493: The merycodontines eventually gave rise to the modern pronghorn. The Cervinae emerged as the first group of extant cervids around 7–9 Mya, during the late Miocene in central Asia. The tribe Muntiacini made its appearance as † Muntiacus leilaoensis around 7–8 Mya; The early muntjacs varied in size–as small as hares or as large as fallow deer. They had tusks for fighting and antlers for defence. Capreolinae followed soon after; Alceini appeared 6.4–8.4 Mya. Around this period,
14136-464: The photoperiod . Deer are also excellent jumpers and swimmers. Deer are ruminants , or cud-chewers, and have a four-chambered stomach. Some deer, such as those on the island of Rùm , do consume meat when it is available. Nearly all deer have a facial gland in front of each eye. The gland contains a strongly scented pheromone , used to mark its home range. Bucks of a wide range of species open these glands wide when angry or excited. All deer have
14322-830: The polar desert of the high Arctic Archipelago and Grant's caribou ( R. t. granti also called the Porcupine caribou ) lives in the western end of the Alaska Peninsula and the adjacent islands; the other four subspecies, Osborn's caribou ( R. t. osborni ), Stone's caribou ( R. t. stonei ), the Rocky Mountain caribou ( R. t. fortidens ) and the Selkirk Mountains caribou ( R. t. montanus ) are all montane . The extinct insular Queen Charlotte Islands caribou ( R. t. dawsoni ), lived on Graham Island in Haida Gwaii (formerly known as
14508-509: The tundra , taiga (boreal forest) and south through the Canadian Rocky Mountains . Of the eight subspecies classified by Harding (2022) into the Arctic caribou ( R. arcticus ), the migratory mainland barren-ground caribou of Arctic Alaska and Northern Canada ( R. t. arcticus ), summer in tundra and winter in taiga, a transitional forest zone between boreal forest and tundra; the nomadic Peary caribou ( R. t. pearyi ) lives in
14694-544: The 1820s, when the Dutch, British, and Danish whalers moved elsewhere in the Arctic. By the late 17th century, Russian hunters arrived; they overwintered to a greater extent and hunted land mammals such as the polar bear and fox. Norwegian hunting—mostly for walrus—started in the 1790s. The first Norwegian citizens to reach Spitsbergen proper were a number of Coast Sámi people from the Hammerfest region, who were hired as part of
14880-465: The 1920 Spitsbergen Treaty (now the Svalbard Treaty ). The name Spitsbergen originated with Dutch navigator and explorer Willem Barentsz , who in 1596 described the "pointed mountains" or, in Dutch, spitse bergen that he saw on the west coast of the main island. Barentsz did not recognize that he had discovered an archipelago, and consequently the name Spitsbergen long remained in use both for
15066-481: The Arctic caribou. Siberian tundra reindeer herds are also in decline, and Rangifer as a whole is considered to be Vulnerable by the International Union for Conservation of Nature (IUCN). Charles Hamilton Smith is credited with the name Rangifer for the reindeer genus, which Albertus Magnus used in his De animalibus , fol. Liber 22, Cap. 268: "Dicitur Rangyfer quasi ramifer". This word may go back to
15252-731: The Beverly herd was also indicated by eight alleles found in the Beverly herd which were absent from the Baffin Island samples. Jenkins et al. (2018) also reported genetic distinctiveness of Baffin Island caribou from all other barren-ground caribou; its genetic signature was not found on the mainland or on other islands; nor were Beverly herd (the nearest mainly barren-ground caribou) alleles present in Baffin Island caribou, evidence of reproductive isolation. These advances in Rangifer genetics were brought together with previous morphological-based descriptions, ecology, behavior and archaeology to propose
15438-536: The British Columbia side, and Banff National Park , Jasper National Park , and Glacier National Park (U.S.) on the Alberta and Montana sides. Mountain slope habitats vary from moist coniferous/mixed forested habitats to dry subalpine/pine forests with alpine meadows higher up. The foothills and river valleys between the mountain ranges provide a mosaic of cropland and deciduous parklands. The rare woodland caribou have
15624-502: The Cervidae, are believed to have evolved from Diacodexis , the earliest known artiodactyl (even-toed ungulate), 50–55 Mya in the Eocene. Diacodexis , nearly the size of a rabbit , featured the talus bone characteristic of all modern even-toed ungulates . This ancestor and its relatives occurred throughout North America and Eurasia, but were on the decline by at least 46 Mya. Analysis of
15810-460: The English zoologist Joshua Brookes in 1828), Cervinae (described by Goldfuss) and Hydropotinae (first described by French zoologist Édouard Louis Trouessart in 1898). Other attempts at the classification of deer have been based on morphological and genetic differences. The Anglo-Irish naturalist Victor Brooke suggested in 1878 that deer could be bifurcated into two classes on the according to
15996-676: The Eurasian reindeer radiation dates to the large Riss glaciation (347,000 to 128,000 years ago), based on the Norwegian-Svalbard split 225,000 years ago. Finnish forest reindeer ( R. t. fennicus ) likely evolved from Cervus [Rangifer] geuttardi Desmarest, 1822, a reindeer that adapted to forest habitats in Eastern Europe as forests expanded during an interglacial period before the LGM (the Würmian or Weichsel glaciation );. The fossil species geuttardi
16182-752: The European Dremotherium ; these sabre-toothed animals are believed to have been the direct ancestors of all modern antlered deer, though they themselves lacked antlers. Another contemporaneous form was the four-horned protoceratid Protoceras , that was replaced by Syndyoceras in the Miocene; these animals were unique in having a horn on the nose. Late Eocene fossils dated approximately 35 million years ago, which were found in North America, show that Syndyoceras had bony skull outgrowths that resembled non-deciduous antlers. Fossil evidence suggests that
16368-659: The George River herd, as reported by the Canadian Broadcasting Corporation . The New York Times reported in April 2018 of the disappearance of the only herd of southern mountain woodland caribou in the contiguous United States , with an expert calling it "functionally extinct" after the herd's size dwindled to a mere three animals. After the last individual, a female, was translocated to a wildlife rehabilitation center in Canada, caribou were considered extirpated from
16554-513: The Greenland caribou ( R. t. groenlandicus ) and the Svalbard reindeer ( R. t. platyrhynchus ), although not closely related to each other, were the most genetically divergent among Rangifer clades; that modern (see Evolution above) Eurasian tundra reindeer ( R. t. tarandus and R. t. sibiricus ) and North American barren-ground caribou ( R. t. arcticus ), although sharing ancestry, were separable at
16740-673: The Ice Ages, but afterwards became restricted primarily to the Anatolian Peninsula, in present-day Turkey. Present-day fallow deer populations in Europe are a result of historic man-made introductions of this species, first to the Mediterranean regions of Europe, then eventually to the rest of Europe. They were initially park animals that later escaped and reestablished themselves in the wild. Historically, Europe's deer species shared their deciduous forest habitat with other herbivores, such as
16926-623: The Indian Subcontinent) boasts the most species of deer in the world, with most species being found in Asia. Europe, in comparison, has lower diversity in plant and animal species. Many national parks and protected reserves in Europe have populations of red deer, roe deer , and fallow deer. These species have long been associated with the continent of Europe, but also inhabit Asia Minor , the Caucasus Mountains , and Northwestern Iran . "European" fallow deer historically lived over much of Europe during
17112-518: The Mammals of the World Vol. 2: Hoofed Mammals . Most Russian authors also recognized R. t. angustirostris , a forest reindeer from east of Lake Baikal . However, since 1991, many genetic studies have revealed deep divergence between modern tundra reindeer and woodland caribou. Geist (2007) and others continued arguing that the woodland caribou was incorrectly classified, noting that "true woodland caribou,
17298-557: The Norwegians returned and re-established their presence. In September 1944, the Germans set up their last weather station, Operation Haudegen in Nordaustlandet ; it functioned until after the German surrender. On 4 September 1945, the soldiers were picked up by a Norwegian seal hunting vessel and surrendered to its captain. This group of men were the last German troops to surrender after
17484-495: The Queen Charlotte Islands). The boreal woodland caribou ( R. t. caribou ), lives in the boreal forest of northeastern Canada: the Labrador or Ungava caribou of northern Quebec and northern Labrador ( R. t. caboti ), and the Newfoundland caribou of Newfoundland ( R. t. terranovae ) have been found to be genetically in the woodland caribou lineage. In Eurasia, both wild and domestic reindeer are distributed across
17670-572: The Reindeer and Caribou, Genus Rangifer (1961), eliminated R. t. caboti (the Labrador caribou), R. t. osborni (Osborn's caribou — from British Columbia ) and R. t. terranovae (the Newfoundland caribou) as invalid and included only barren-ground caribou , renamed as R. t. groenlandicus (formerly R. arcticus ) and woodland caribou as R. t. caribou . However, Banfield made multiple errors, eliciting
17856-488: The Second World War. After the war, the Soviet Union proposed common Norwegian and Soviet administration and military defence of Svalbard. This was rejected in 1947 by Norway, which two years later joined NATO . The Soviet Union retained high civilian activity on Svalbard, in part to ensure that the archipelago was not used by NATO. After the war, Norway re-established operations at Longyearbyen and Ny-Ålesund, while
18042-800: The Siberian forest reindeer ( R. t. valentinae , formerly called the Busk Mountains reindeer ( R. t. buskensis ) by American taxonomists) occupies the Altai and Ural Mountains . Male ("bull") and female ("cow") reindeer can grow antlers annually, although the proportion of females that grow antlers varies greatly between populations. Antlers are typically larger on males. Antler architecture varies by species and subspecies and, together with pelage differences, can often be used to distinguish between species and subspecies (see illustrations in Geist, 1991 and Geist, 1998). About 25,000 mountain reindeer ( R. t. tarandus ) still live in
18228-408: The Soviet Union established mining in Barentsburg, Pyramiden , and Grumant . The mine at Ny-Ålesund had several fatal accidents, killing 71 people while it was in operation from 1945 to 1954 and from 1960 to 1963. The Kings Bay Affair , caused by the 1962 accident killing 21 workers, forced Gerhardsen's Third Cabinet to resign. From 1964, Ny-Ålesund became a research outpost, and a facility for
18414-506: The Spitsbergen archipelago. Russians have traditionally called the archipelago Grumant ( Грумант ). The Soviet Union retained the name Spitsbergen ( Шпицберген ) to support undocumented claims that Russians were the first to discover the island. In 1928, Italian explorer Umberto Nobile and the crew of the airship Italia crashed on the icepack off the coast of Foyn Island . The subsequent rescue attempts were covered extensively in
18600-597: The Ussuri Region (Russia). These are among some of the richest deciduous and coniferous forests in the world where one can find Siberian roe deer , sika deer , elk, and moose. Asian caribou occupy the northern fringes of this region along the Sino-Russian border. Deer such as the sika deer, Thorold's deer , Central Asian red deer , and elk have historically been farmed for their antlers by Han Chinese , Turkic peoples , Tungusic peoples , Mongolians , and Koreans . Like
18786-556: The Yukon as invalid subspecies of woodland caribou, then R. t. caribou . This left the small, migratory barren-ground caribou of Alaska and the Yukon, including the Porcupine caribou herd, without a name, which Banfield rectified in his 1974 Mammals of Canada by extending to them the name " granti ". The late Valerius Geist (1998), in the only error in his whole illustrious career, re-analyzed Banfield's data with additional specimens found in an unpublished report he cites as "Skal, 1982", but
18972-407: The alimentary canal. The deer require a large amount of minerals such as calcium and phosphate in order to support antler growth, and this further necessitates a nutrient-rich diet. There are some reports of deer engaging in carnivorous activity, such as eating dead alewives along lakeshores or depredating the nests of northern bobwhites . Nearly all cervids are so-called uniparental species:
19158-608: The antlers as well as the canines are small. The tragulids have long canines to this day. With the onset of the Pliocene , the global climate became cooler. A fall in the sea-level led to massive glaciation; consequently, grasslands abounded in nutritious forage. Thus a new spurt in deer populations ensued. The oldest member of Cervini, † Cervocerus novorossiae , appeared around the transition from Miocene to Pliocene (4.2–6 Mya) in Eurasia; cervine fossils from early Pliocene to as late as
19344-864: The antlers create grooves that allow another male's antlers to lock into place. This allows the males to wrestle without risking injury to the face. Antlers are correlated to an individual's position in the social hierarchy and its behaviour. For instance, the heavier the antlers, the higher the individual's status in the social hierarchy, and the greater the delay in shedding the antlers; males with larger antlers tend to be more aggressive and dominant over others. Antlers can be an honest signal of genetic quality; males with larger antlers relative to body size tend to have increased resistance to pathogens and higher reproductive capacity. In elk in Yellowstone National Park , antlers also provide protection against predation by wolves . Homology of tines, that is,
19530-579: The archipelago, the Norwegian Coast Guard embarked on a significant modernization program. As of 2023, the Coast Guard is replacing its older Nordkapp -class offshore patrol vessels with significantly larger ice-capable ships, each displacing just under 10,000 tonnes. The three new Jan Mayen -class offshore patrol vessels are armed with a 57 mm (2.2 in) main gun and are capable of operating up to two medium-sized helicopters. The ships have
19716-519: The archipelago, the seat of the governor and the only incorporated town. The town features an airport , hospital, primary and secondary school , university , sports center with a swimming pool, library, culture center, cinema, bus transport, hotels, a bank, and several museums. The newspaper Svalbardposten is published weekly. Very little mining activity remains at Longyearbyen; coal mines at Sveagruva and Lunckefjellet suspended operations in 2017 and were closed permanently in 2020. Ny-Ålesund
19902-499: The basis of diploid number of chromosomes in the late 20th century has been flawed by several inconsistencies. In 1987, the zoologists Colin Groves and Peter Grubb identified three subfamilies: Cervinae, Hydropotinae and Odocoileinae; they noted that the hydropotines lack antlers, and the other two subfamilies differ in their skeletal morphology. They reverted from this classification in 2000. Molecular phylogenetic analyses since
20088-692: The branching structure of antlers among species, have been discussed before the 1900s. Recently, a new method to describe the branching structure of antlers and determining homology of tines was developed. Most deer bear 32 teeth; the corresponding dental formula is: 0.0.3.3 3.1.3.3 . The elk and the reindeer may be exceptions, as they may retain their upper canines and thus have 34 teeth (dental formula: 0.1.3.3 3.1.3.3 ). The Chinese water deer, tufted deer, and muntjac have enlarged upper canine teeth forming sharp tusks, while other species often lack upper canines altogether. The cheek teeth of deer have crescent ridges of enamel, which enable them to grind
20274-405: The chital, the fallow deer and the sika deer feature white spots on a brown coat. Coat of reindeer shows notable geographical variation. Deer undergo two moults in a year; for instance, in red deer the red, thin-haired summer coat is gradually replaced by the dense, greyish brown winter coat in autumn, which in turn gives way to the summer coat in the following spring. Moulting is affected by
20460-452: The combination of full moon and reflective snow can give additional light. Due to the Earth's tilt and the high latitude, Svalbard has extensive twilights. Longyearbyen sees the first and last day of polar night having seven and a half hours of twilight, whereas the perpetual light lasts for two weeks longer than the midnight sun. On the summer solstice, the sun bottoms out at 12° sun angle in
20646-417: The earliest members of the superfamily Cervoidea appeared in Eurasia in the Miocene. Dicrocerus , Euprox and Heteroprox were probably the first antlered cervids. Dicrocerus featured single-forked antlers that were shed regularly. Stephanocemas had more developed and diffuse ("crowned") antlers. Procervulus ( Palaeomerycidae ) also had antlers that were not shed. Contemporary forms such as
20832-643: The early Eocene , and gradually developed into the first antlered cervoids (the superfamily of cervids and related extinct families) in the Miocene . Eventually, with the development of antlers, the tusks as well as the upper incisors disappeared. Thus, evolution of deer took nearly 30 million years. Biologist Valerius Geist suggests evolution to have occurred in stages. There are not many prominent fossils to trace this evolution, but only fragments of skeletons and antlers that might be easily confused with false antlers of non-cervid species. The ruminants , ancestors of
21018-686: The early Pleistocene, probably as a result of abundant resources to drive evolution. The early Pleistocene cervid † Eucladoceros was comparable in size to the modern elk. † Megaloceros (Pliocene–Pleistocene) featured the Irish elk ( M. giganteus ), one of the largest known cervids . The Irish elk reached 2 metres ( 6 + 1 ⁄ 2 ft) at the shoulder and had heavy antlers that spanned 3.6 metres (11 ft 10 in) from tip to tip. These large animals were traditionally thought to have faced extinction due to conflict between sexual selection for large antlers and body and natural selection for
21204-413: The early taxonomists. Similarly, working on museum collections where skins were often faded and in poor states of preservation, early taxonomists could not readily perceive differences in coat patterns that are consistent within a subspecies, but variable among them. Geist calls these "nuptial" characteristics: sexually selected characters that are highly conserved and diagnostic among subspecies. Towards
21390-654: The ecological niches of the ibex and wild goat , with the fawns behaving more like goat kids. The highest concentration of large deer species in temperate North America lies in the Canadian Rocky Mountain and Columbia Mountain regions between Alberta and British Columbia where all five North American deer species ( white-tailed deer , mule deer , caribou , elk , and moose ) can be found. This region has several clusters of national parks including Mount Revelstoke National Park , Glacier National Park (Canada) , Yoho National Park , and Kootenay National Park on
21576-727: The emergence of cervids from the Oligocene to the early Pliocene . The latter half of the Oligocene (28–34 Mya) saw the appearance of the European Eumeryx and the North American Leptomeryx . The latter resembled modern-day bovids and cervids in dental morphology (for instance, it had brachyodont molars), while the former was more advanced . Other deer-like forms included the North American Blastomeryx and
21762-652: The end of the 19th century, national museums began sending out biological exploration expeditions and collections accumulated. Taxonomists, usually working for the museums, began naming subspecies more rigorously, based on statistical differences in detailed cranial, dental and skeletal measurements than antlers and pelage, supplemented by better knowledge of differences in ecology and behavior. From 1898 to 1937, mammalogists named 12 new species (other than barren-ground and woodland, which had been named earlier) of caribou in Canada and Alaska, and three new species and nine new subspecies in Eurasia, each properly described according to
21948-410: The end of their first winter. In the first twenty minutes of a fawn's life, the fawn begins to take its first steps. Its mother licks it clean until it is almost free of scent, so predators will not find it. Its mother leaves often to graze, and the fawn does not like to be left behind. Sometimes its mother must gently push it down with her foot. The fawn stays hidden in the grass for one week until it
22134-793: The endangered barasingha and very common chital are gregarious and live in large herds. Indian sambar can be gregarious but are usually solitary or live in smaller herds. Hog deer are solitary and have lower densities than Indian muntjac. Deer can be seen in several national parks in India, Nepal, and Sri Lanka of which Kanha National Park , Dudhwa National Park , and Chitwan National Park are most famous. Sri Lanka's Wilpattu National Park and Yala National Park have large herds of Indian sambar and chital. The Indian sambar are more gregarious in Sri Lanka than other parts of their range and tend to form larger herds than elsewhere. The Chao Praya River Valley of Thailand
22320-602: The evolution" of Rangifer species and other Arctic and sub-Arctic species. Isolation of tundra-adapted species Rangifer in Last Glacial Maximum refugia during the last glacial – the Wisconsin glaciation in North America and the Weichselian glaciation in Eurasia – shaped "intraspecific genetic variability " particularly between the North American and Eurasian parts of the Arctic . Reindeer / caribou ( Rangifer ) are in
22506-485: The evolving rules of zoological nomenclature, with type localities designated and type specimens deposited in museums (see table in Species and subspecies below). In the mid-20th century, as definitions of "species" evolved, mammalogists in Europe and North America made all Rangifer species conspecific with R. tarandus , and synonymized most of the subspecies. Alexander William Francis Banfield 's often-cited A Revision of
22692-950: The extinct tarpan (forest horse), extinct aurochs (forest ox), and the endangered wisent (European bison). Good places to see deer in Europe include the Scottish Highlands , the Austrian Alps , the wetlands between Austria , Hungary , and the Czech Republic , and some National Parks, including Doñana National Park in Spain , the Veluwe in the Netherlands , the Ardennes in Belgium , and Białowieża National Park in Poland . Spain , Eastern Europe , and
22878-458: The extinct caribou Torontoceros [Rangifer] hypogaeus , had features (robust and short pedicles, smooth antler surface, and high position of second tine) that relate it to forest caribou. Humans started hunting reindeer in both the Mesolithic and Neolithic Periods, and humans are today the main predator in many areas. Norway and Greenland have unbroken traditions of hunting wild reindeer from
23064-417: The features of the second and fifth metacarpal bones of their forelimbs: Plesiometacarpalia (most Old World deer) and Telemetacarpalia (most New World deer). He treated the musk deer as a cervid, placing it under Telemetacarpalia. While the telemetacarpal deer showed only those elements located far from the joint, the plesiometacarpal deer retained the elements closer to the joint as well. Differentiation on
23250-538: The females. Their bez tines are set low, just above the brow tine, which is vertically flattened to protect the eyes while the buck "threshes" low brush, a courtship display. The low bez tines help the wide flat brow tines dig craters in the hard-packed tundra snow for forage, for which reason brow tines are often called "shovels" in North America and "ice tines" in Europe. The differences in antler architecture reflect fundamental differences in ecology and behavior, and in turn deep divisions in ancestry that were not apparent to
23436-489: The first discovery of the archipelago in 1596, when he sighted the coast of the island of Spitsbergen while searching for the Northern Sea Route . The first recorded landing on the islands of Svalbard dates to 1604, when an English ship landed at Bjørnøya , or Bear Island, and started hunting walrus . Annual expeditions soon followed, and Spitsbergen became a base for hunting the bowhead whale from 1611. Because of
23622-523: The foothills and river valley bottoms of the Canadian Rockies owing to conversion of land to cropland and the clearing of coniferous forests allowing more deciduous vegetation to grow up the mountain slopes. They also live in the aspen parklands north of Calgary and Edmonton, where they share habitat with the moose. The adjacent Great Plains grassland habitats are left to herds of elk, American bison , and pronghorn . The Eurasian Continent (including
23808-612: The former R. t. groenlandicus (now R. t. arcticus ). R. t. granti was lost in the oblivion of invalid taxonomy until Alaskan researchers sampled some small, pale caribou from the western end of the Alaska Peninsula, their range enclosing the type locality designated by Allen (1902) and found them to be genetically distinct from all other caribou in Alaska. Thus, granti was rediscovered, its range restricted to that originally described. Deer A deer ( pl. : deer) or true deer
23994-758: The four western Canadian montane ecotypes are not woodland caribou: they share a common ancestor with modern barren-ground caribou / tundra reindeer, but distantly, having diverged > 60,000 years ago — before the modern ecotypes had evolved their cold- and darkness-adapted physiologies and mass-migration and aggregation behaviors (see Evolution above). Before Banfield (1961), taxonomists using cranial, dental and skeletal measurements had unequivocally allied these western montane ecotypes with barren-ground caribou, naming them (as in Osgood 1909 Murie, 1935 and Anderson 1946, among others) R. t. stonei , R. t. montanus , R. t. fortidens and R. t. osborni , respectively, and this phylogeny
24180-610: The fringes of the Rocky Mountain and Laurentide ice sheets as far south as northern Alabama ; and in Sangamonian deposits (~100,000 years BP) from western Canada. A R. t. pearyi -sized caribou occupied Greenland before and after the LGM and persisted in a relict enclave in northeastern Greenland until it went extinct about 1900 (see discussion of R. t. eogroenlandicus below). Archaeological excavations showed that larger barren-ground-sized caribou appeared in western Greenland about 4,000 years ago. The late Valerius Geist (1998) dates
24366-476: The genus Rangifer as R. grœnlandicus . It went back and forth as a full species or subspecies of the barren-ground caribou ( R. arcticus ) or a subspecies of the tundra reindeer ( R. tarandus ), but always as the Greenland reindeer / caribou. Taxonomists consistently documented morphological differences between Greenland and other caribou / reindeer in cranial measurements, dentition, antler architecture, etc. Then Banfield (1961) in his famously flawed revision, gave
24552-509: The institution of the Governor of Svalbard ( Norwegian : Sysselmester , formerly Sysselmannen ), who holds the responsibility as both county governor and chief of police , as well as holding other authority granted from the executive branch. Duties include environmental policy , family law , law enforcement , search and rescue , tourism management, information services, contact with foreign settlements, and judge in some areas of maritime inquiries and judicial examinations—albeit never in
24738-431: The island of Spitsbergen, which constitutes more than half the archipelago, followed by Nordaustlandet and Edgeøya. All settlements are on Spitsbergen, except the meteorological outposts on Bjørnøya and Hopen . The Norwegian state took possession of all unclaimed land, or 95.2% of the archipelago, at the time the Svalbard Treaty entered into force; Store Norske , a Norwegian coal mining company, owns 4%, Arktikugol ,
24924-641: The islands were being used as a base for Arctic exploration . The first mining was along Isfjorden by Norwegians in 1899; by 1904, British interests had established themselves in Adventfjorden and started the first year-round operations. Production in Longyearbyen, by US interests, started in 1908; and Store Norske established itself in 1916, as did other Norwegian interests during the First World War , in part by buying US interests. Discussions to establish
25110-476: The larger caribou that appeared in Greenland 4,000 years ago originated from Baffin Island (itself unique; see Taxonomy above), a reconstruction of LGM glacial retreat and caribou advance (Yannic et al. 2013) shows colonization by NAL lineage caribou more likely. Their PCA and tree diagrams show Greenland caribou clustering outside of the Beringian-Eurasian lineage. The scientific name R. t. granti has
25296-499: The late Miocene–Pliocene; this appears highly probable as the camelids migrated into Asia from North America around the same time. Deer invaded South America in the late Pliocene (2.5–3 Mya) as part of the Great American Interchange , thanks to the recently formed Isthmus of Panama , and emerged successful due to the small number of competing ruminants in the continent. Large deer with impressive antlers evolved during
25482-450: The latter half of the 2000s all show that hydropotes is a sister taxon of Capreolus , and “Hydropotinae” became outdated subfamily. Until 2003, it was understood that the family Moschidae (musk deer) was sister to Cervidae. Then a phylogenetic study by Alexandre Hassanin (of National Museum of Natural History, France ) and colleagues, based on mitochondrial and nuclear analyses, revealed that Moschidae and Bovidae form
25668-462: The lawless nature of the area, English , Danish , Dutch , and French companies and authorities tried to use force to keep out other countries' fleets. Smeerenburg was one of the first settlements, established by the Dutch in 1619. Smaller bases were also built by the English, Danish, and French. At first the outposts were merely summer camps, but from the early 1630s, a few individuals started to overwinter . Whaling at Spitsbergen lasted until
25854-591: The local economy. Apart from Longyearbyen, other settlements include the Russian mining-community of Barentsburg , the Norwegian research-station of Ny-Ålesund , and the Swedish-Norwegian mining outpost of Sveagruva (which closed in 2020). Other settlements lie farther north, but are populated only by rotating groups of researchers. No roads connect the settlements; instead snowmobiles , aircraft and boats provide inter-settlement transport. Svalbard Airport serves as
26040-419: The main gateway. Approximately 60% of the archipelago is covered with glaciers , and the islands feature many mountains and fjords . The archipelago has an Arctic climate , although with significantly higher temperatures than other areas at the same latitude. The flora has adapted to take advantage of the long period of midnight sun to compensate for the polar night . Svalbard is home to Rock Ptarmigan,
26226-432: The main island and for the archipelago as a whole. Later the main island was sometimes distinguished as West Spitsbergen . The spelling Spitzbergen , with z instead of s, derives from German. The Svalbard Treaty of 1920 defines Svalbard as all islands, islets, and skerries from 74° to 81° north latitude, and from 10° to 35° east longitude. The land area is 61,022 km (23,561 sq mi), and dominated by
26412-581: The main island. In 1827 Baltazar Keilhau first proposed that the Old Norse toponym Svalbarði , found in medieval Icelandic sources, referred to Spitsbergen. Keilhau's theory was revived by Gustav Storm in 1890 and Gunnar Isachsen in 1907, at a time when ancient Norse connection to the land would help modern Norway's contested claim to sovereignty. Svalbard is a modern Norwegian analogue of Svalbarði , which in turn derives from svalr ('cold') and barð ('edge', 'ridge', 'turf', 'beard'). The Icelandic Annals record that Svalbarði
26598-399: The male of any species is a hart , especially if over five years old, and the female is a hind , especially if three or more years old. The young of small species is a fawn and of large species a calf ; a very small young may be a kid . A castrated male is a havier . A group of any species is a herd . The adjective of relation is cervine ; like the family name Cervidae , this
26784-450: The middle of the night, being much higher during night than in mainland Norway's polar light areas. However, the daytime strength of the sun remains as low as 35°. Glacial ice covers 36,502 km (14,094 sq mi) or 60% of Svalbard; 30% is barren rock while 10% is vegetated. The largest glacier is Austfonna (8,412 km or 3,248 sq mi) on Nordaustlandet, followed by Olav V Land and Vestfonna . During summer, it
26970-434: The more general sense: for example, Dutch / Frisian dier , German Tier , and Norwegian dyr mean ' animal ' . For many types of deer in modern English usage, the male is a buck and the female a doe , but the terms vary with dialect, and according to the size of the species. The male red deer is a stag , while for other large species the male is a bull , the female a cow , as in cattle. In older usage,
27156-413: The most restricted range living at higher altitudes in the subalpine meadows and alpine tundra areas of some of the mountain ranges. Elk and mule deer both migrate between the alpine meadows and lower coniferous forests and tend to be most common in this region. Elk also inhabit river valley bottomlands, which they share with White-tailed deer. The White-tailed deer have recently expanded their range within
27342-578: The mountains of Norway, notably in Hardangervidda . In Sweden there are approximately 250,000 reindeer in herds managed by Sámi villages. Russia manages 19 herds of Siberian tundra reindeer ( R. t. sibiricus ) that total about 940,000. The Taimyr herd of Siberian tundra reindeer is the largest wild reindeer herd in the world, varying between 400,000 and 1,000,000; it is a metapopulation consisting of several subpopulations — some of which are phenotypically different — with different migration routes and calving areas. The Kamchatkan reindeer ( R. t. phylarchus ),
27528-546: The name groenlandicus to all the barren-ground caribou in North America, Greenland included, because groenlandicus pre-dates Richardson's R. arctus . However, because genetic data shows the Greenland caribou to be the most distantly related of any caribou to all the others (genetic distance, FST = 44%, whereas most cervid (deer family) species have a genetic distance of 2% to 5%)--as well as behavioral and morphological differences—a recent revision returned it to species status as R. groenlandicus . Although it has been assumed that
27714-568: The northern states of the contiguous United States from Maine to Washington . Boreal woodland caribou have disappeared from most of their original southern range and were designated as Threatened in 2002 by the Committee on the Status of Endangered Wildlife in Canada (COSEWIC). Environment and Climate Change Canada reported in 2011 that there were approximately 34,000 boreal woodland caribou in 51 ranges remaining in Canada (Environment Canada, 2011b), although those numbers included montane populations classified by Harding (2022) into subspecies of
27900-522: The population on the islands (Norwegians making up the remaining third) with the population of the archipelago slightly under 4,000. Russian activity has diminished considerably since then, falling from 2,500 to 450 people from 1990 to 2010. Grumant was closed after it was depleted in 1962. Pyramiden was closed in 1998. Coal exports from Barentsburg ceased in 2006 because of a fire, but resumed in 2010. The Russians experienced two air accidents: Vnukovo Airlines Flight 2801 , which killed 141 people, and
28086-544: The possible exception of the Indian muntjac . There are also several species of deer that are highly specialized and live almost exclusively in mountains, grasslands, swamps, and "wet" savannas, or riparian corridors surrounded by deserts . Some deer have a circumpolar distribution in both North America and Eurasia . Examples include the caribou that live in Arctic tundra and taiga (boreal forests) and moose that inhabit taiga and adjacent areas. Huemul deer ( taruca and Chilean huemul ) of South America 's Andes fill
28272-400: The press and Svalbard received short-lived fame as a result. Svalbard, known to both British and Germans as Spitsbergen, was little affected by the German invasion of Norway in April 1940. The settlements continued to operate as before, mining coal and monitoring the weather. In July 1941, following the German invasion of the Soviet Union, the Royal Navy reconnoitered the islands with
28458-414: The previously named subspecies distributions, without naming them as such, plus some ecotypes. Ecotypes are not phylogenetically based and cannot substitute for taxonomy. Meanwhile, genetic data continued to accumulate, revealing sufficiently deep divisions to easily separate Rangifer back into six previously named species and to resurrect several previously named subspecies. Molecular data showed that
28644-409: The recent revision (see Reindeer#Taxonomy below). Reindeer / caribou ( Rangifer ) vary in size from the smallest, the Svalbard reindeer ( R. ( t. ) platyrhynchus ), to the largest, Osborn's caribou ( R. t. osborni ). They also vary in coat color and antler architecture. The North American range of caribou extends from Alaska through the Yukon , the Northwest Territories and Nunavut throughout
28830-428: The recent revision): In North America, R. t. caboti , R. t. caribou , R. t. dawsoni , R. t. groenlandicus , R. t. osborni , R. t. pearyi , and R. t. terranovae ; and in Eurasia, R. t. tarandus , R. t. buskensis (called R. t. valentinae in Europe; see below), R. t. phylarchus , R. t. pearsoni , R. t. sibiricus and R. t. platyrhynchus . These subspecies were retained in the 2011 replacement work Handbook of
29016-402: The right spot and never really find it." Deer appear to be immune to this parasite; it passes through the digestive system and is excreted in the feces. The parasite is not screened by the moose intestine, and passes into the brain where damage is done that is externally apparent, both in behaviour and in gait. Deer, elk and moose in North America may suffer from chronic wasting disease , which
29202-610: The same cases as acting as police. Since 2021, Lars Fause has been governor. The institution is subordinate to the Ministry of Justice and the Police , but reports to other ministries in matters within their portfolio. Since 2002, Longyearbyen Community Council has had many of the same responsibilities of a municipality , including utilities, education, cultural facilities, fire department, roads, and ports. No care or nursing services are available, nor are welfare payments available. Norwegian residents retain pension and medical rights through their mainland municipalities. The hospital
29388-417: The second most diverse family of artiodactyla after bovids. Though of a similar build, deer are strongly distinguished from antelopes by their antlers , which are temporary and regularly regrown unlike the permanent horns of bovids. Characteristics typical of deer include long, powerful legs, a diminutive tail and long ears. Deer exhibit a broad variation in physical proportions. The largest extant deer
29574-501: The skull and are often used for combat between males. The musk deer ( Moschidae ) of Asia and chevrotains ( Tragulidae ) of tropical African and Asian forests are separate families that are also in the ruminant clade Ruminantia ; they are not especially closely related to Cervidae. Deer appear in art from Paleolithic cave paintings onwards, and they have played a role in mythology , religion, and literature throughout history, as well as in heraldry , such as red deer that appear in
29760-433: The smallest, the Svalbard reindeer ( R. ( t. ) platyrhynchus ), to the largest, Osborn's caribou ( R. t. osborni ). Although reindeer are quite numerous, some species and subspecies are in decline and considered vulnerable . They are unique among deer (Cervidae) in that females may have antlers , although the prevalence of antlered females varies by subspecies. Reindeer are the only successfully semi-domesticated deer on
29946-420: The sole maintenance host in the Michigan outbreak of bovine tuberculosis which remains a significant barrier to the US nationwide eradication of the disease in livestock. Moose and deer can carry rabies . Docile moose may suffer from brain worm , a helminth which drills holes through the brain in its search for a suitable place to lay its eggs. A government biologist states that "They move around looking for
30132-409: The sovereignty of the archipelago commenced in the 1910s, but were interrupted by World War I . On 9 February 1920, following the Paris Peace Conference , the Svalbard Treaty was signed, granting full sovereignty to Norway. However, all signatory countries were granted non-discriminatory rights to fishing, hunting, and mineral resources. The treaty took effect on 14 August 1925, at the same time as
30318-440: The subfamily Odocoileinae , along with roe deer ( Capreolus ), Eurasian elk / moose ( Alces ), and water deer ( Hydropotes ). These antlered cervids split from the horned ruminants Bos (cattle and yaks), Ovis (sheep) and Capra (goats) about 36 million years ago. The Eurasian clade of Odocoileinae (Capreolini, Hydropotini and Alcini) split from the New World tribes of Capreolinae ( Odocoileini and Rangiferini) in
30504-427: The subspecies level; that Finnish forest reindeer ( R. t. fennicus ) clustered well apart from both wild and domestic tundra reindeer and that boreal woodland caribou ( R. t. caribou ) were separable from all others. Meanwhile, archaeological evidence was accumulating that Eurasian forest reindeer descended from an extinct forest-adapted reindeer and not from tundra reindeer (see Evolution above); since they do not share
30690-461: The terms reindeer and caribou for essentially the same animal can cause confusion, but the ICUN clearly delineates the issue: "Reindeer is the European name for the species of Rangifer, while in North America, Rangifer species are known as Caribou." The word reindeer is an anglicized version of the Old Norse words hreinn ("reindeer") and dýr ("animal") and has nothing to do with reins. The word caribou comes through French, from
30876-574: The third year. This process of losing a set of antlers to develop a larger and more branched set continues for the rest of the life. The antlers emerge as soft tissues (known as velvet antlers ) and progressively harden into bony structures (known as hard antlers), following mineralisation and blockage of blood vessels in the tissue, from the tip to the base. Antlers might be one of the most exaggerated male secondary sexual characteristics , and are intended primarily for reproductive success through sexual selection and for combat. The tines (forks) on
31062-541: The time of parturition. The fallow deer and the various subspecies of the reindeer have the largest as well as the heaviest antlers, both in absolute terms as well as in proportion to body mass (an average of eight grams per kilogram of body mass); the tufted deer, on the other hand, has the smallest antlers of all deer, while the pudú has the lightest antlers with respect to body mass (0.6 g per kilogram of body mass). The structure of antlers show considerable variation; while fallow deer and elk antlers are palmate (with
31248-477: The treaty (including Spain, Iceland and particularly Russia) argue that the Treaty provides them with extensive rights beyond Svalbard's territorial sea. Norway claims an exclusive economic zone of more than three-quarters of a million square kilometers around Svalbard, though "Russia does not recognize Norwegian functional rights with respect to the Svalbard Fisheries Protection Area". In the 2020s, in order to strengthen Norway's ability to enforce its claims around
31434-431: The treaty have the right to conduct commercial activities on the archipelago without discrimination, although all activity is subject to Norwegian legislation. The treaty limits Norway's right to collect taxes to that of financing services on Svalbard. Therefore, Svalbard has a lower income tax than mainland Norway, and there is no value added tax . There is a separate budget for Svalbard to ensure compliance. Svalbard
31620-428: The tundra and into the taiga. Eurasian mountain reindeer ( R. t. tarandus ) are close to North American caribou genetically and visually, but with sufficient differences to warrant division into two species. The unique, insular Svalbard reindeer inhabits the Svalbard Archipelago . The Finnish forest reindeer ( R. t. fennicus ) is spottily distributed in the coniferous forest zones from Finland to east of Lake Baikal :
31806-944: The uniformly dark, small-maned type with the frontally emphasized, flat-beamed antlers", is "scattered thinly along the southern rim of North American caribou distribution". He affirms that the "true woodland caribou is very rare, in very great difficulties and requires the most urgent of attention." In 2011, noting that the former classifications of Rangifer tarandus , either with prevailing taxonomy on subspecies, designations based on ecotypes , or natural population groupings, failed to capture "the variability of caribou across their range in Canada" needed for effective subspecies conservation and management, COSEWIC developed Designatable Unit (DU) attribution, an adaptation of "evolutionary significant units". The 12 designatable units for caribou in Canada (that is, excluding Alaska and Greenland) based on ecology, behavior and, importantly, genetics (but excluding morphology and archaeology) essentially followed
31992-404: The various Asian rhinoceros species, various antelope species (such as nilgai , four-horned antelope , blackbuck , and Indian gazelle in India), and wild oxen (such as wild Asian water buffalo , gaur , banteng , and kouprey ). One way that different herbivores can survive together in a given area is for each species to have different food preferences, although there may be some overlap. As
32178-427: The various forms of the Woodland Caribou are so great in almost every respect that no detailed comparison is necessary. ...According to Mr. Stone, Rangifer granti inhabits the " barren land of Alaska Peninsula, ranging well up into the mountains in summer, but descending to the lower levels in winter, generally feeding on the low flat lands near the coast and in the foothills...As regards cranial characters no comparison
32364-438: The west coast of Spitsbergen. Whalers who sailed far north in the 17th and 18th centuries used the islands as a base; subsequently the archipelago was abandoned. Coal mining started at the beginning of the 20th century, and several permanent communities such as Pyramiden or Barentsburg were established. The Svalbard Treaty of 1920 recognizes Norwegian sovereignty, and the Norwegian Svalbard Act of 1925 made Svalbard
32550-406: The world's two most northerly Lenin statues and other socialist realist art. As of 2013 , a handful of workers are stationed in the largely abandoned Pyramiden to maintain local infrastructure and run its hotel, which has been re-opened to tourism. Most of the population is Christian. Most of the Norwegians are affiliated with the Church of Norway . Russian and Ukrainian population belongs to
32736-512: The world. Clearing open areas within forests to some extent may actually benefit deer populations by exposing the understory and allowing the types of grasses, weeds, and herbs to grow that deer like to eat. Access to adjacent croplands may also benefit deer. Adequate forest or brush cover must still be provided for populations to grow and thrive. Deer are widely distributed, with indigenous representatives in all continents except Antarctica and Australia , though Africa has only one native deer,
32922-491: The young, known in most species as fawns, are only cared for by the mother, most often called a doe. A doe generally has one or two fawns at a time (triplets, while not unknown, are uncommon). Mating season typically begins in later August and lasts until December. Some species mate until early March. The gestation period is anywhere up to ten months for the European roe deer. Most fawns are born with their fur covered with white spots, though in many species they lose these spots by
33108-399: Was "not able to find diagnostic features that could segregate this form from the western barren ground type." But Skal 1982 had included specimens from the eastern end of the Alaska Peninsula and the Kenai Peninsula , the range of the larger Stone's caribou. Later, geneticists comparing barren-ground caribou of Alaska with those of mainland Canada found little difference and they all became
33294-733: Was confirmed by genetic analysis. DNA also revealed three unnamed clades that, based on genetic distance, genetic divergence and shared vs. private haplotypes and alleles , together with ecological and behavioral differences, may justify separation at the subspecies level: the Atlantic- Gaspésie caribou (COSEWIC DU11), an eastern montane ecotype of the boreal woodland caribou, and the Baffin Island caribou. Neither one of these clades has yet been formally described or named. Jenkins et al. (2012) said that "[Baffin Island] caribou are unique compared to other Barrenground herds, as they do not overwinter in forested habitat, nor do all caribou undertake long seasonal migrations to calving areas." It also shares
33480-427: Was discovered in 1194, while the Landnámabók places it four days' sailing north of Langanes . The word dægr "day" might mean either 12 or 24 hours; Isachsen took the latter interpretation, thus discounting Jan Mayen as Svalbarði . Cultural studies academic Roald Berg says Svalbarði more likely referred to part of Greenland , but the 1925 renaming cemented Norwegian sovereignty as recognised by
33666-434: Was established at Ny-Ålesund in 1941, remaining until 1942. In May 1942, after the arrival of the Fritham force, the German unit at Banso was evacuated. In September 1943 in Operation Zitronella a German task force, which included the battleship Tirpitz , was sent to attack the garrison and destroy the settlements at Longyearbyen and Barentsburg. This was achieved, but had little long-term effect: after their departure
33852-513: Was identified at a Colorado laboratory in the 1960s and is believed to be a prion disease. Out of an abundance of caution hunters are advised to avoid contact with specified risk material (SRM) such as the brain, spinal column or lymph nodes. Deboning the meat when butchering and sanitizing the knives and other tools used to butcher are amongst other government recommendations. Deer are believed to have evolved from antlerless, tusked ancestors that resembled modern duikers and diminutive deer in
34038-407: Was incorporated on 1 January 2002, adopting a community council. In 2016, Svalbard had a population of 2,667, of which 423 were Russian and Ukrainian, 10 Polish, and 322 other non-Norwegians living in Norwegian settlements. The largest non-Norwegian groups in Longyearbyen in 2005 were from Russia, Ukraine, Poland, Germany, Sweden, Denmark, and Thailand. Longyearbyen is the largest settlement on
34224-404: Was later replaced by R. constantini , which was adapted for grasslands, in a second immigration 19,000–20,000 years ago when the LGM turned its forest habitats into tundra, while fennicus survived in isolation in southwestern Europe. R. constantini was then replaced by modern tundra / barren-ground caribou adapted to extreme cold, probably in Beringia, before dispersing west ( R. t. tarandus in
34410-474: Was once primarily tropical seasonal moist deciduous forest and wet savanna that hosted populations of hog deer, the now-extinct Schomburgk's deer , Eld's deer , Indian sambar, and Indian muntjac. Both the hog deer and Eld's deer are rare, whereas Indian sambar and Indian muntjac thrive in protected national parks, such as Khao Yai . Many of these South Asian and Southeast Asian deer species also share their habitat with other herbivores , such as Asian elephants ,
34596-454: Was selected as the senior synonym to R. t. valentinae Flerov, 1933, in Mammal Species of the World but Russian authors do not recognize Millais and Millais' articles in a hunting travelogue, The Gun at Home and Abroad , seem short of a taxonomic authority. The scientific name groenlandicus is fraught with problems. Edwards (1743) illustrated and claimed to have seen a male specimen ("head of perfect horns...") from Greenland and said that
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