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57-649: Orbignya phalerata Orbignya martiana Barb.Rodr. Orbygnia speciosa (Mart. ex Spreng.) Barb.Rodr. Attalea speciosa , the babassu , babassu palm , babaçu , or cusi , is a palm native to the Amazon Rainforest region in South America . The babassu palm is the predominant species in the Maranhão Babaçu forests of Maranhão and Piauí states. This plant has commercial value because its seeds produce an edible oil called babassu oil , which

114-499: A break in rainfall during mid-season when the Intertropical Convergence Zone or monsoon trough moves poleward of their location during the middle of the warm season; Typical vegetation in these areas ranges from moist seasonal tropical forests to savannahs . When the wet season occurs during the warm season, or summer , precipitation falls mainly during the late afternoon and early evening hours. The wet season

171-407: A clustering habit results. Exclusively sympodial genera include many of the rattans , Guihaia , and Rhapis . Several palm genera have both solitary and clustering members. Palms which are usually solitary may grow in clusters and vice versa. Palms have large, evergreen leaves that are either palmately ('fan-leaved') or pinnately ('feather-leaved') compound and spirally arranged at the top of

228-458: A large amount of the human diet and several other human uses, both by absolute amount produced and by number of species domesticated . This is far higher than almost any other plant family, sixth out of domesticated crops in the human diet, and first in total economic value produced – sharing the top spot with the Poaceae and Fabaceae . These human uses have also spread many Arecaceae species around

285-408: A lubricant, and in soap and cosmetics. The shell of the nut can be used to make smokeless charcoal, and flesh of the babassu nut is used to produce flour. Palm hearts are extracted from the tree to make a juice. The leaf and stalk of the babassu palm are used in building materials. Baskets and other handicrafts can be made from the fibre of the leaves. A high-protein cake with about 20% protein content

342-416: A valid taxon but Sidney F Glassman considered it a dubious taxon in his 1999 Taxonomic Treatment of Palm Subtribe Attaleinae . Attalea vitrivir was recognised as a distinct species by Michael Balick and coauthors; Glassman and Govaerts and Dransfield concurred, but Henderson considered it part of A. speciosa . Glassman also described a fourth member of this group, A. brejinhoensis , and it

399-617: A variety of different habitats. Their diversity is highest in wet, lowland forests. South America , the Caribbean , and areas of the South Pacific and southern Asia are regions of concentration. Colombia may have the highest number of palm species in one country. There are some palms that are also native to desert areas such as the Arabian Peninsula and parts of northwestern Mexico. Only about 130 palm species naturally grow entirely beyond

456-609: A water source, and in desert oases such as the date palm . A few palms are adapted to extremely basic lime soils, while others are similarly adapted to extreme potassium deficiency and toxicity of heavy metals in serpentine soils . Palms are a monophyletic group of plants, meaning the group consists of a common ancestor and all its descendants. Extensive taxonomic research on palms began with botanist H.E. Moore , who organized palms into 15 major groups based mostly on general morphological characteristics. The following classification, proposed by N.W. Uhl and J. Dransfield in 1987,

513-508: Is a revision of Moore's classification that organizes palms into 6 subfamilies. A few general traits of each subfamily are listed below. The Phytelephantoideae is the sixth subfamily of Arecaceae in N.W. Uhl and J. Dransfield's 1987 classification. Members of this group have distinct monopodial flower clusters. Other distinct features include a gynoecium with five to 10 joined carpels, and flowers with more than three parts per whorl. Fruits are multiple-seeded and have multiple parts. From

570-500: Is a time when air quality improves, freshwater quality improves and vegetation grows significantly due to the wet season supplementing flora, leading to crop yields late in the season. Floods and rains cause rivers to overflow their banks, and some animals to retreat to higher ground. Soil nutrients are washed away and erosion increases. The incidence of malaria increases in areas where the rainy season coincides with high temperatures. Animals have adaptation and survival strategies for

627-475: Is accepted by Govaerts and Dransfield. A. speciosa is monoecious —male and female flowers are separate, but are borne by the same plant. Pollination has been attributed both to insects and wind. In February 2008, babassu palm oil was used in a blend with coconut oil and jet fuel to power one engine of a Virgin Atlantic Boeing 747 during a flight test. Babassu oil can also be used in food cooking, as

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684-501: Is also used in cleaners and skin-care products. The fruit is used to produce products such as medicines , beauty aids, and beverages . Traditional communities of the Maranhão region also produce a flour from the fruit, and this is commercialized as a nutritional supplement. The leaves are also used to provide thatch for houses and can be woven into mats for constructing house walls . The stems are used for timbers. The babassu palm

741-556: Is considered a weed in pasture areas of Cerrado vegetation in Brazil. In his 1995 The Palms of the Amazon , Andrew Henderson recognised A. speciosa and A. spectabilis as valid species, but considered the latter to either be an acaulescent form of A. speciosa or a hybrid between A. microcarpa and it. In their 2005 World Checklist of Palms Rafäel Govaerts and John Dransfield accepted A. spectabilis as

798-692: Is formed after oil is extracted from the fruits. This cake is typically used as animal feed. Arecaceae The Arecaceae ( / ˌ ær ə ˈ k eɪ s i . iː , - ˌ aɪ / ) is a family of perennial , flowering plants in the monocot order Arecales . Their growth form can be climbers , shrubs , tree -like and stemless plants, all commonly known as palms . Those having a tree-like form are colloquially called palm trees . Currently, 181 genera with around 2,600 species are known, most of which are restricted to tropical and subtropical climates. Most palms are distinguished by their large, compound, evergreen leaves, known as fronds , arranged at

855-595: Is from Dravidian *aṭ-ay-kkāy ("areca nut"). The suffix -aceae is the feminine plural of the Latin -āceus ("resembling"). Palm originates from Latin palma semantically overlapping with sense of " hand front " (due to similar splayed shape) ultimately from Proto-Indo-European * pl̥h₂meh₂ , a direct descendant folm once existed in Old English. Whether as shrubs, tree-like, or vines, palms have two methods of growth: solitary or clustered. The common representation

912-496: Is often called "anomalous secondary growth". The Arecaceae are notable among monocots for their height and for the size of their seeds, leaves, and inflorescences. Ceroxylon quindiuense , Colombia's national "tree", is the tallest monocot in the world, reaching up to 60 metres (197 ft) tall. The coco de mer ( Lodoicea maldivica ) has the largest seeds of any plant, 40–50 centimetres (16–20 in) in diameter and weighing 15–30 kilograms (33–66 lb) each ( coconuts are

969-662: Is possible north of subtropical climates, and some higher latitude locales such as Ireland, Scotland, England, and the Pacific Northwest feature a few palms in protected locations and microclimates . In the United States , there are at least 12 native palm species, mostly occurring in the states of the Deep South and Florida . Palms inhabit a variety of ecosystems. More than two-thirds of palm species live in humid moist forests, where some species grow tall enough to form part of

1026-460: Is that of a solitary shoot ending in a crown of leaves. This monopodial character may be exhibited by prostrate, trunkless, and trunk-forming members. Some common palms restricted to solitary growth include Washingtonia and Roystonea . Palms may instead grow in sparse though dense clusters. The trunk develops an axillary bud at a leaf node, usually near the base, from which a new shoot emerges. The new shoot, in turn, produces an axillary bud and

1083-585: Is that the tropics represent a ' Garden of Eden ', a heaven on Earth, a land of rich biodiversity or a tropical paradise. The alternative is that the tropics consist of wild, unconquerable nature. The latter view was often discussed in old Western literature more so than the first. Evidence suggests over time that the view of the tropics as such in popular literature has been supplanted by more well-rounded and sophisticated interpretations. Western scholars tried to theorise why tropical areas were relatively more inhospitable to human civilisations than colder regions of

1140-589: The Andes as far south as the northernmost parts of Chile and Perú . The climate is changing in the tropics, as it is in the rest of the world. The effects of steadily rising concentrations of greenhouse gases on the climate may be less obvious to tropical residents, however, because they are overlain by considerable natural variability. Much of this variability is driven by the El Niño-Southern Oscillation (ENSO) . The Tropics has warmed by 0.7–0.8 °C over

1197-559: The Dasypogonaceae . For a phylogenetic tree of the family, see the list of Arecaceae genera . The Arecaceae were the first modern family of monocots to appear in the fossil record around 80 million years ago (Mya), during the late Cretaceous period. The first modern species, such as Nypa fruticans and Acrocomia aculeata , appeared 69 Mya, as evidenced by fossil Nypa pollen. Palms appear to have undergone an early period of adaptive radiation . By 60 Mya, many of

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1254-703: The Pantropic . The system of biogeographic realms differs somewhat; the Neotropical realm includes both the Neotropics and temperate South America, and the Paleotropics correspond to the Afrotropical , Indomalayan , Oceanian , and tropical Australasian realms . Flora are plants found in a specific region at a specific time. Some well-known plants that are exclusively found in, originate from, or are often associated with

1311-570: The Waterberg Biosphere of South Africa , and eastern Madagascar rainforests. Often the soils of tropical forests are low in nutrient content, making them quite vulnerable to slash-and-burn deforestation techniques, which are sometimes an element of shifting cultivation agricultural systems. In biogeography , the tropics are divided into Paleotropics (Africa, Asia and Australia) and Neotropics (Caribbean, Central America, and South America). Together, they are sometimes referred to as

1368-696: The axial tilt of the Earth . The Tropic of Cancer is the Northernmost latitude from which the Sun can ever be seen directly overhead , and the Tropic of Capricorn is the Southernmost. This means that the tropical zone includes everywhere on Earth which is a subsolar point at least once during the solar year . Thus the maximum latitudes of the tropics have equal distances from the equator on either side. Likewise, they approximate

1425-522: The canopy and shorter ones form part of the understory . Some species form pure stands in areas with poor drainage or regular flooding, including Raphia hookeri which is common in coastal freshwater swamps in West Africa. Other palms live in tropical mountain habitats above 1 thousand metres (3 thousand feet), such as those in the genus Ceroxylon native to the Andes . Palms may also live in grasslands and scrublands, usually associated with

1482-640: The Ceroxyloideae and Phytelephantoideae. Studies have suggested the lack of a fully resolved hypothesis for the relationships within the family is due to a variety of factors, including difficulties in selecting appropriate outgroups, homoplasy in morphological character states, slow rates of molecular evolution important for the use of standard DNA markers , and character polarization. However, hybridization has been observed among Orbignya and Phoenix species, and using chloroplast DNA in cladistic studies may produce inaccurate results due to maternal inheritance of

1539-558: The IUCN, and was published in 1996. The rarest palm known is Hyophorbe amaricaulis . The only living individual remains at the Botanic Gardens of Curepipe in Mauritius . Some pests are specialists to particular taxa . Pests that attack a variety of species of palms include: The palm branch was a symbol of triumph and victory in classical antiquity . The Romans rewarded champions of

1596-520: The Middle East and North Africa. A wide range of common products and foods are derived from palms. In contemporary times, palms are also widely used in landscaping. In many historical cultures, because of their importance as food, palms were symbols for such ideas as victory, peace, and fertility. The word Arecaceae is derived from the word areca with the suffix "-aceae". Areca is derived from Portuguese , via Malayalam അടയ്ക്ക ( aṭaykka ), which

1653-565: The Rathayatra carts which carry the deities of Krishna and his family members in the cart festival of Jagganath Puri in India are marked with the emblem of a palm tree. Specifically it is the symbol of Krishna's brother, Baladeva. In 1840, the American geologist Edward Hitchcock (1793–1864) published the first tree-like paleontology chart in his Elementary Geology , with two separate trees of life for

1710-399: The angle of the Earth's axial tilt. This angle is not perfectly fixed, mainly due to the influence of the moon, but the limits of the tropics are a geographic convention, and their variance from the true latitudes is very small. Many tropical areas have both a dry and a wet season. The wet season , rainy season or green season is the time of year, ranging from one or more months when most of

1767-530: The area. The sale of seeds to nurseries and collectors is another threat, as the seeds of popular palms are sometimes harvested directly from the wild. In 2006, at least 100 palm species were considered endangered, and nine species have been reported as recently extinct. However, several factors make palm conservation more difficult. Palms live in almost every type of warm habitat and have tremendous morphological diversity. Most palm seeds lose viability quickly, and they cannot be preserved in low temperatures because

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1824-455: The average annual rainfall in a region falls. Areas with wet seasons are disseminated across portions of the tropics and subtropics , some even in temperate regions. Under the Köppen climate classification , for tropical climates , a wet-season month is defined as one or more months where average precipitation is 60 mm (2.4 in) or more. Some areas with pronounced rainy seasons see

1881-407: The base. The stamens generally number six, with filaments that may be separate, attached to each other, or attached to the pistil at the base. The fruit is usually a single- seeded drupe (sometimes berry-like) but some genera (e.g., Salacca ) may contain two or more seeds in each fruit. Like all monocots , palms do not have the ability to increase the width of a stem ( secondary growth ) via

1938-466: The chloroplast DNA. Chemical and molecular data from non-organelle DNA, for example, could be more effective for studying palm phylogeny. Recently, nuclear genomes and transcriptomes have been used to reconstruct the phylogeny of palms. This has revealed, for example, that a whole-genome duplication event occurred early in the evolution of the Arecaceae lineage, that was not experienced by its sister clade,

1995-493: The cold kills the embryo. Using botanical gardens for conservation also presents problems, since they can rarely house more than a few plants of any species or truly imitate the natural setting. There is also the risk that cross-pollination can lead to hybrid species. The Palm Specialist Group of the World Conservation Union (IUCN) began in 1984, and has performed a series of three studies to find basic information on

2052-721: The deserts, it also created a more amenable habitat for the people to live in by providing shade and protection from the desert winds. An indication of the importance of palms in ancient times is that they are mentioned more than 30 times in the Bible , and at least 22 times in the Quran . The Torah also references the "70 date palm trees", which symbolize the 70 aspects of Torah that are revealed to those who "eat of its fruit." Arecaceae have great economic importance, including coconut products, oils, dates, palm syrup , ivory nuts, carnauba wax, rattan cane, raffia, and palm wood . This family supplies

2109-652: The games and celebrated military successes with palm branches. Early Christians used the palm branch to symbolize the victory of the faithful over enemies of the soul, as in the Palm Sunday festival celebrating the triumphal entry of Jesus Christ into Jerusalem . In Judaism , the palm represents peace and plenty, and is one of the Four Species of Sukkot ; the palm may also symbolize the Tree of Life in Kabbalah . The canopies of

2166-456: The geographic region; these usages ought not be confused. The Earth's axial tilt is currently around 23.4° , and therefore so are the latitudes of the tropical circles , marking the boundary of the tropics: specifically, ±23°26′09.8″ (or 23.43605°). The northern one is called the Tropic of Cancer , and the southern is the Tropic of Capricorn . As the Earth's axial tilt changes, so too do

2223-500: The habitat, and those with small habitat ranges are most vulnerable to them. The harvesting of heart of palm, a delicacy in salads, also poses a threat because it is derived from the palm's apical meristem , a vital part of the palm that cannot be regrown (except in domesticated varieties, e.g. of peach palm ). The use of rattan palms in furniture has caused a major population decrease in these species that has negatively affected local and international markets, as well as biodiversity in

2280-432: The history of the Middle East and North Africa. In the text "Date Palm Products" (1993), W.H. Barreveld wrote: One could go as far as to say that, had the date palm not existed, the expansion of the human race into the hot and barren parts of the "old" world would have been much more restricted. The date palm not only provided a concentrated energy food, which could be easily stored and carried along on long journeys across

2337-955: The last century—only slightly less than the global average—but a strong El Niño made 1998 the warmest year in most areas, with no significant warming since. Climate models predict a further 1–2 °C warming by 2050 and 1–4 °C by 2100. Tropical plants and animals are those species native to the tropics. Tropical ecosystems may consist of tropical rainforests , seasonal tropical forests , dry (often deciduous) forests , spiny forests, deserts , savannahs, grasslands and other habitat types. There are often wide areas of biodiversity , and species endemism present, particularly in rainforests and seasonal forests. Some examples of important biodiversity and high-endemism ecosystems are El Yunque National Forest in Puerto Rico , Costa Rican and Nicaraguan rainforests, Amazon Rainforest territories of several South American countries, Madagascar dry deciduous forests ,

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2394-673: The modern phylogenomic data, the Phytelephantoideae are tribe in the Ceroxyloideae subfamily. Currently, few extensive phylogenetic studies of the Arecaceae exist. In 1997, Baker et al. explored subfamily and tribe relationships using chloroplast DNA from 60 genera from all subfamilies and tribes. The results strongly showed the Calamoideae are monophyletic, and Ceroxyloideae and Coryphoideae are paraphyletic. The relationships of Arecoideae are uncertain, but they are possibly related to

2451-660: The modern, specialized genera of palms appeared and became widespread and common, much more widespread than their range today. Because palms separated from the monocots earlier than other families, they developed more intrafamilial specialization and diversity. By tracing back these diverse characteristics of palms to the basic structures of monocots, palms may be valuable in studying monocot evolution. Several species of palms have been identified from flowers preserved in amber, including Palaeoraphe dominicana and Roystonea palaea . Fossil evidence of them can also be found in samples of petrified palmwood . The relationship between

2508-719: The overhead sun, the tropics receive the most solar energy over the course of the year, and consequently have the highest temperatures on the planet. Even when not directly overhead, the sun is still close to overhead throughout the year, therefore the tropics also have the lowest seasonal variation on the planet; "winter" and "summer" lose their contrast. Instead, seasons are more commonly divided by precipitation variations than by temperature variations. The tropics maintain wide diversity of local climates, such as rain forests , monsoons , savannahs , deserts , and high altitude snow-capped mountains . The word "tropical" can specifically refer to certain kinds of weather , rather than to

2565-505: The plants and the animals. These are crowned (graphically) with the Palms and with Man. Today, the palm, especially the coconut palm, remains a symbol of the tropical island paradise . Palms appear on the flags and seals of several places where they are native, including those of Haiti , Guam , Saudi Arabia , Florida , and South Carolina . Some species commonly called palms, though they are not true palms, include: Tropics Due to

2622-399: The same kind of vascular cambium found in non-monocot woody plants . This explains the cylindrical shape of the trunk (almost constant diameter) that is often seen in palms, unlike in ring -forming trees. However, many palms, like some other monocots, do have secondary growth, although because it does not arise from a single vascular cambium producing xylem inwards and phloem outwards, it

2679-486: The second largest). Raffia palms ( Raphia spp.) have the largest leaves of any plant, up to 25 metres (82 ft) long and 3 metres (10 ft) wide. The Corypha species have the largest inflorescence of any plant, up to 7.5 metres (25 ft) tall and containing millions of small flowers. Calamus stems can reach 200 metres (656 ft) in length. Most palms are native to tropical and subtropical climates. Palms thrive in moist and hot climates but can be found in

2736-476: The status of palms in the wild, use of wild palms, and palms under cultivation. Two projects on palm conservation and use supported by the World Wildlife Fund took place from 1985 to 1990 and 1986–1991, in the American tropics and southeast Asia, respectively. Both studies produced copious new data and publications on palms. Preparation of a global action plan for palm conservation began in 1991, supported by

2793-411: The stem. The leaves have a tubular sheath at the base that usually splits open on one side at maturity. The inflorescence is a spadix or spike surrounded by one or more bracts or spathes that become woody at maturity. The flowers are generally small and white, radially symmetric, and can be either uni- or bisexual. The sepals and petals usually number three each and may be distinct or joined at

2850-476: The subfamilies is shown in the following cladogram: Calamoideae Nypoideae Coryphoideae Ceroxyloideae Arecoideae Evidence for cultivation of the date palm by Mesopotamians and other Middle Eastern peoples exists from more than 5,000 years ago, in the form of date wood, pits for storing dates, and other remains of the date palm in Mesopotamian sites. The date palm had a significant effect on

2907-479: The top of an unbranched stem, except for the Hyphaene genus, who has branched palms. However, palms exhibit an enormous diversity in physical characteristics and inhabit nearly every type of habitat within their range, from rainforests to deserts . Palms are among the best known and most extensively cultivated plant families. They have been important to humans throughout much of history, especially in regions like

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2964-516: The tropical and polar circles . The tropics constitute 39.8% of Earth's surface area and contain 36% of Earth's landmass . As of 2014 , the region was home also to 40% of the world's population , and this figure was then projected to reach 50% by 2050. Because of global warming , the weather conditions of the tropics are expanding with areas in the subtropics , having more extreme weather events such as heatwaves and more intense storms. These changes in weather conditions may make certain parts of

3021-464: The tropics include: Tropicality refers to the image of the tropics that people from outside the tropics have of the region, ranging from critical to verging on fetishism. Tropicality gained renewed interest in geographical discourse when French geographer Pierre Gourou published Les pays tropicaux ( The Tropical World in English), in the late 1940s. Tropicality encompassed two major images. One,

3078-550: The tropics uninhabitable. The word "tropic" comes via Latin from Ancient Greek τροπή ( tropē ), meaning "to turn" or "change direction". The tropics are defined as the region between the Tropic of Cancer in the Northern Hemisphere at 23°26′09.8″ (or 23.43605°) N and the Tropic of Capricorn in the Southern Hemisphere at 23°26′09.8″ (or 23.43605°) S; these latitudes correspond to

3135-625: The tropics, mostly in humid lowland subtropical climates, in highlands in southern Asia, and along the rim lands of the Mediterranean Sea . The northernmost native palm is Chamaerops humilis , which reaches 44°N latitude along the coast of Liguria , Italy . In the southern hemisphere, the southernmost palm is the Rhopalostylis sapida , which reaches 44°S on the Chatham Islands where an oceanic climate prevails. Cultivation of palms

3192-708: The wetter regime. The previous dry season leads to food shortages into the wet season, as the crops have yet to mature. However, regions within the tropics may well not have a tropical climate. Under the Köppen climate classification, much of the area within the geographical tropics is classed not as "tropical" but as "dry" ( arid or semi-arid ), including the Sahara Desert , the Atacama Desert and Australian Outback . Also, there are alpine tundra and snow-capped peaks, including Mauna Kea , Mount Kilimanjaro , Puncak Jaya and

3249-426: The world. Along with dates mentioned above, members of the palm family with human uses are numerous: Like many other plants, palms have been threatened by human intervention and exploitation . The greatest risk to palms is destruction of habitat , especially in the tropical forests , due to urbanization , wood-chipping , mining , and conversion to farmland . Palms rarely reproduce after such great changes in

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