171-418: In botany , a tree is a perennial plant with an elongated stem , or trunk , usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth , plants that are usable as lumber or plants above a specified height. In wider definitions, the taller palms , tree ferns , bananas , and bamboos are also trees. Trees are not
342-429: A monophyletic taxonomic group but consist of a wide variety of plant species that have independently evolved a trunk and branches as a way to tower above other plants to compete for sunlight. The majority of tree species are angiosperms or hardwoods; of the rest, many are gymnosperms or softwoods. Trees tend to be long-lived, some reaching several thousand years old. Trees evolved around 370 million years ago, and it
513-408: A symbiotic relationship with Frankia species, a filamentous bacterium that can fix nitrogen from the air, converting it into ammonia . They have actinorhizal root nodules on their roots in which the bacteria live. This process enables the tree to live in low nitrogen habitats where they would otherwise be unable to thrive. The plant hormones called cytokinins initiate root nodule formation, in
684-539: A beard. However, no direct or contemporary attestation of this origin exists, and the name Joshua tree is not recorded until after Mormon contact; moreover, the physical appearance of the Joshua tree more closely resembles a similar story told of Moses . Ranchers and miners who were contemporaneous with the Mormon immigrants used the trunks and branches as fencing and for fuel for ore-processing steam engines. In addition to
855-483: A botanically and pharmacologically important herbal Historia Plantarum in 1544 and a pharmacopoeia of lasting importance, the Dispensatorium in 1546. Naturalist Conrad von Gesner (1516–1565) and herbalist John Gerard (1545– c. 1611 ) published herbals covering the supposed medicinal uses of plants. Naturalist Ulisse Aldrovandi (1522–1605) was considered the father of natural history , which included
1026-413: A classic example of parallel evolution . With an estimated 60,000-100,000 species, the number of trees worldwide might total twenty-five per cent of all living plant species. The greatest number of these grow in tropical regions; many of these areas have not yet been fully surveyed by botanists , making tree diversity and ranges poorly known. The majority of tree species are angiosperms or hardwoods. Of
1197-414: A considerable problem in agriculture, and the biology and control of plant pathogens in agriculture and natural ecosystems . Ethnobotany is the study of the relationships between plants and people. When applied to the investigation of historical plant–people relationships ethnobotany may be referred to as archaeobotany or palaeoethnobotany . Some of the earliest plant-people relationships arose between
1368-452: A continuum between the major morphological categories of root, stem (caulome), leaf (phyllome) and trichome . Furthermore, it emphasises structural dynamics. Modern systematics aims to reflect and discover phylogenetic relationships between plants. Modern Molecular phylogenetics largely ignores morphological characters, relying on DNA sequences as data. Molecular analysis of DNA sequences from most families of flowering plants enabled
1539-427: A desert species; new seedlings may grow at an average rate of 7.6 cm (3.0 in) per year in their first 10 years, then only about 3.8 cm (1.5 in) per year. The trunk consists of thousands of small fibers and lacks annual growth rings , making determining the tree's age difficult. This tree has a top-heavy branch system, and a broad root system, with roots in one case found 11 m (36 ft) from
1710-502: A drier savanna climate and insufficient rainfall to support dense forests, the canopy is not closed, and plenty of sunshine reaches the ground which is covered with grass and scrub. Acacia and baobab are well adapted to living in such areas. The roots of a tree serve to anchor it to the ground and gather water and nutrients to transfer to all parts of the tree. They are also used for reproduction, defence, survival, energy storage and many other purposes. The radicle or embryonic root
1881-523: A group of Mormon settlers crossing the Mojave Desert in the mid-19th century: The tree's role in guiding them through the desert combined with its unique shape reminded them of a biblical story in which Joshua keeps his hands reached out for an extended period of time to enable the Israelites in their conquest of Canaan ( Joshua 8:18–26 ). Further, the shaggy leaves may have provided the appearance of
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#17328374968082052-400: A hierarchical classification of plant species that remains the reference point for modern botanical nomenclature . This established a standardised binomial or two-part naming scheme where the first name represented the genus and the second identified the species within the genus. For the purposes of identification, Linnaeus's Systema Sexuale classified plants into 24 groups according to
2223-436: A high probability that their populations will be reduced by 90% of their current range by the end of the 21st century, thus fundamentally transforming the ecosystem of the park. Wildfires, invasive grasses and poor migration patterns for the trees' seeds are all additional factors in the species' imperilment. As an example, approximately 13%—or more than 1.3 million Joshua trees—in one of the densest Joshua tree populations in
2394-417: A hundred litres are required to make one litre of birch syrup. Various parts of trees are used as spices. These include cinnamon , made from the bark of the cinnamon tree ( Cinnamomum zeylanicum ) and allspice , the dried small fruits of the pimento tree ( Pimenta dioica ). Nutmeg is a seed found in the fleshy fruit of the nutmeg tree ( Myristica fragrans ) and cloves are the unopened flower buds of
2565-404: A plant sucks water through them under water stress. Lignin is also used in other cell types like sclerenchyma fibres that provide structural support for a plant and is a major constituent of wood. Sporopollenin is a chemically resistant polymer found in the outer cell walls of spores and pollen of land plants responsible for the survival of early land plant spores and the pollen of seed plants in
2736-552: A polymer of fructose is used for the same purpose in the sunflower family Asteraceae . Some of the glucose is converted to sucrose (common table sugar) for export to the rest of the plant. Unlike in animals (which lack chloroplasts), plants and their eukaryote relatives have delegated many biochemical roles to their chloroplasts , including synthesising all their fatty acids , and most amino acids . The fatty acids that chloroplasts make are used for many things, such as providing material to build cell membranes out of and making
2907-507: A process closely related to mycorrhizal association. It has been demonstrated that some trees are interconnected through their root system, forming a colony. The interconnections are made by the inosculation process, a kind of natural grafting or welding of vegetal tissues. The tests to demonstrate this networking are performed by injecting chemicals, sometimes radioactive , into a tree, and then checking for its presence in neighbouring trees. The roots are, generally, an underground part of
3078-400: A process known as cladoptosis . The crown is the spreading top of a tree including the branches and leaves, while the uppermost layer in a forest, formed by the crowns of the trees, is known as the canopy . A sapling is a young tree. Many tall palms are herbaceous monocots, which do not undergo secondary growth and never produce wood. In many tall palms, the terminal bud on the main stem is
3249-460: A process that generates molecular oxygen (O 2 ) as a by-product. The light energy captured by chlorophyll a is initially in the form of electrons (and later a proton gradient ) that's used to make molecules of ATP and NADPH which temporarily store and transport energy. Their energy is used in the light-independent reactions of the Calvin cycle by the enzyme rubisco to produce molecules of
3420-505: A pure form of carbon made by pyrolysis of wood, has a long history as a metal- smelting fuel, as a filter material and adsorbent and as an artist's material and is one of the three ingredients of gunpowder . Cellulose , the world's most abundant organic polymer, can be converted into energy, fuels, materials and chemical feedstock. Products made from cellulose include rayon and cellophane , wallpaper paste , biobutanol and gun cotton . Sugarcane , rapeseed and soy are some of
3591-421: A second spurt of growth, a Lammas growth may occur which is believed to be a strategy to compensate for loss of early foliage to insect predators. Primary growth is the elongation of the stems and roots. Secondary growth consists of a progressive thickening and strengthening of the tissues as the outer layer of the epidermis is converted into bark and the cambium layer creates new phloem and xylem cells. The bark
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#17328374968083762-489: A significant role in reducing erosion and moderating the climate . They remove carbon dioxide from the atmosphere and store large quantities of carbon in their tissues. Trees and forests provide a habitat for many species of animals and plants. Tropical rainforests are among the most biodiverse habitats in the world. Trees provide shade and shelter , timber for construction, fuel for cooking and heating, and fruit for food as well as having many other uses. In much of
3933-422: A similar growth form, by usually growing larger and having a single main stem; but there is no consistent distinction between a tree and a shrub, made more confusing by the fact that trees may be reduced in size under harsher environmental conditions such as on mountains and subarctic areas. The tree form has evolved separately in unrelated classes of plants in response to similar environmental challenges, making it
4104-465: A single tree species, which will not flourish in the absence of its mycorrhizal associate. Others are generalists and associate with many species. The tree acquires minerals such as phosphorus from the fungus, while the fungus obtains the carbohydrate products of photosynthesis from the tree. The hyphae of the fungus can link different trees and a network is formed, transferring nutrients and signals from one place to another. The fungus promotes growth of
4275-630: A slightly looser definition; while the Joshua tree , bamboos and palms do not have secondary growth and never produce true wood with growth rings, they may produce "pseudo-wood" by lignifying cells formed by primary growth. Tree species in the genus Dracaena , despite also being monocots, do have secondary growth caused by meristem in their trunk, but it is different from the thickening meristem found in dicotyledonous trees. Aside from structural definitions, trees are commonly defined by use; for instance, as those plants which yield lumber. The tree growth habit
4446-526: A student of Aristotle who invented and described many of its principles and is widely regarded in the scientific community as the "Father of Botany". His major works, Enquiry into Plants and On the Causes of Plants , constitute the most important contributions to botanical science until the Middle Ages , almost seventeen centuries later. Another work from Ancient Greece that made an early impact on botany
4617-625: A threat to population density of prone areas but not to the limits of the range itself, that several population studies showed Yucca brevifolia was abundant, and that although the southern region of the species' range has been reduced, the trend is that the northern region has been expanding over the last 11,700 years as the North American ice cap melted, allowing the species to occupy its current range. The studies showing reduced population after fires used aerial photography to document populations which would underreport smaller and thus younger trees, as
4788-478: A tree is any plant with the general form of an elongated stem, or trunk, which supports the photosynthetic leaves or branches at some distance above the ground. Trees are also typically defined by height, with smaller plants from 0.5 to 10 m (1.6 to 32.8 ft) being called shrubs , so the minimum height of a tree is only loosely defined. Large herbaceous plants such as papaya and bananas are trees in this broad sense. A commonly applied narrower definition
4959-454: A useful proxy for temperature in historical climatology , and the biological impact of climate change and global warming . Palynology , the analysis of fossil pollen deposits in sediments from thousands or millions of years ago allows the reconstruction of past climates. Estimates of atmospheric CO 2 concentrations since the Palaeozoic have been obtained from stomatal densities and
5130-1010: A variety of spatial scales in groups, populations and communities that collectively constitute vegetation. Regions with characteristic vegetation types and dominant plants as well as similar abiotic and biotic factors, climate , and geography make up biomes like tundra or tropical rainforest . Herbivores eat plants, but plants can defend themselves and some species are parasitic or even carnivorous . Other organisms form mutually beneficial relationships with plants. For example, mycorrhizal fungi and rhizobia provide plants with nutrients in exchange for food, ants are recruited by ant plants to provide protection, honey bees , bats and other animals pollinate flowers and humans and other animals act as dispersal vectors to spread spores and seeds . Plant responses to climate and other environmental changes can inform our understanding of how these changes affect ecosystem function and productivity. For example, plant phenology can be
5301-628: A way of drug discovery . Plants can synthesise coloured dyes and pigments such as the anthocyanins responsible for the red colour of red wine , yellow weld and blue woad used together to produce Lincoln green , indoxyl , source of the blue dye indigo traditionally used to dye denim and the artist's pigments gamboge and rose madder . Sugar, starch , cotton, linen , hemp , some types of rope , wood and particle boards , papyrus and paper, vegetable oils , wax , and natural rubber are examples of commercially important materials made from plant tissues or their secondary products. Charcoal ,
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5472-436: A wide range of edible fruits are found in the tropics. Other commercially important fruit include dates, figs and olives. Palm oil is obtained from the fruits of the oil palm ( Elaeis guineensis ). The fruits of the cocoa tree ( Theobroma cacao ) are used to make cocoa and chocolate and the berries of coffee trees, Coffea arabica and Coffea canephora , are processed to extract the coffee beans. In many rural areas of
5643-416: A wide range of shapes and sizes, in response to environmental pressures including climate and predation. They can be broad or needle-like, simple or compound, lobed or entire, smooth or hairy, delicate or tough, deciduous or evergreen. The needles of coniferous trees are compact but are structurally similar to those of broad-leaved trees. They are adapted for life in environments where resources are low or water
5814-452: A wider range of shared characters and were widely followed. The Candollean system reflected his ideas of the progression of morphological complexity and the later Bentham & Hooker system , which was influential until the mid-19th century, was influenced by Candolle's approach. Darwin 's publication of the Origin of Species in 1859 and his concept of common descent required modifications to
5985-516: A winter freeze before they bloom. Once they bloom, the flowers are pollinated by the yucca moth ( Tegeticula synthetica ), which spreads pollen while laying eggs inside the flower. The larvae feed on the seeds, but enough seeds remain to reproduce. The Joshua tree is also able to actively abort ovaries in which too many eggs have been produced. The Joshua tree is native to the southwestern United States (Arizona, California, Nevada, and Utah) and northwestern Mexico. This range mostly coincides with
6156-406: Is De materia medica , a five-volume encyclopedia about preliminary herbal medicine written in the middle of the first century by Greek physician and pharmacologist Pedanius Dioscorides . De materia medica was widely read for more than 1,500 years. Important contributions from the medieval Muslim world include Ibn Wahshiyya 's Nabatean Agriculture , Abū Ḥanīfa Dīnawarī 's (828–896)
6327-405: Is peppermint , Mentha × piperita , a sterile hybrid between Mentha aquatica and spearmint, Mentha spicata . The many cultivated varieties of wheat are the result of multiple inter- and intra- specific crosses between wild species and their hybrids. Angiosperms with monoecious flowers often have self-incompatibility mechanisms that operate between the pollen and stigma so that
6498-476: Is a broad, multidisciplinary subject with contributions and insights from most other areas of science and technology. Research topics include the study of plant structure , growth and differentiation, reproduction , biochemistry and primary metabolism , chemical products, development , diseases , evolutionary relationships , systematics , and plant taxonomy . Dominant themes in 21st-century plant science are molecular genetics and epigenetics , which study
6669-548: Is a flavouring obtained from distilling bark from the roots of the sassafras tree ( Sassafras albidum ). The leaves of trees are widely gathered as fodder for livestock and some can be eaten by humans but they tend to be high in tannins which makes them bitter. Leaves of the curry tree ( Murraya koenigii ) are eaten, those of kaffir lime ( Citrus × hystrix ) (in Thai food ) and Ailanthus (in Korean dishes such as bugak ) and those of
6840-475: Is a plant species belonging to the genus Yucca . It is tree-like in habit, which is reflected in its common names. This monocotyledonous tree is native to the arid Southwestern United States , specifically California , Arizona , Utah , and Nevada , and northwestern Mexico . It is confined mostly to the Mojave Desert between 400 and 1,800 m (1,300 and 5,900 ft) elevation. It thrives in
7011-511: Is a rudimentary stalk and neatly folded miniature leaves, ready to expand when the next growing season arrives. Buds also form in the axils of the leaves ready to produce new side shoots. A few trees, such as the eucalyptus , have "naked buds" with no protective scales and some conifers, such as the Lawson's cypress , have no buds but instead have little pockets of meristem concealed among the scale-like leaves. When growing conditions improve, such as
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7182-416: Is a sudden movement of sap at the end of the winter as trees prepare to burst into growth. In North America, the sap of the sugar maple ( Acer saccharum ) is used in the production of maple syrup . About 90% of the sap is water, the remaining 10% being a mixture of various sugars and certain minerals. The sap is harvested by drilling holes in the trunks of the trees and collecting the liquid that flows out of
7353-524: Is an evolutionary adaptation found in different groups of plants: by growing taller, trees are able to compete better for sunlight. Trees tend to be tall and long-lived, some reaching several thousand years old. Several trees are among the oldest organisms now living. Trees have modified structures such as thicker stems composed of specialised cells that add structural strength and durability, allowing them to grow taller than many other plants and to spread out their foliage. They differ from shrubs , which have
7524-468: Is called "hunuvat chiy'a" or "humwichawa" by the indigenous Cahuilla . It is also called izote de desierto (Spanish, "desert dagger"). It was first formally described in the botanical literature as Yucca brevifolia by George Engelmann in 1871 as part of the Geological Exploration of the 100th meridian (or " Wheeler Survey "). The name "Joshua tree" is commonly said to have been given by
7695-417: Is continually replaced by a living layer of cells called the cork cambium or phellogen. The London plane ( Platanus × hispanica ) periodically sheds its bark in large flakes. Similarly, the bark of the silver birch ( Betula pendula ) peels off in strips. As the tree's girth expands, newer layers of bark are larger in circumference, and the older layers develop fissures in many species. In some trees such as
7866-497: Is developed by the Indian banyan . Many large trees have buttress roots which flare out from the lower part of the trunk. These brace the tree rather like angle brackets and provide stability, reducing sway in high winds. They are particularly prevalent in tropical rainforests where the soil is poor and the roots are close to the surface. Some tree species have developed root extensions that pop out of soil, in order to get oxygen, when it
8037-414: Is estimated that there are around three trillion mature trees in the world currently. A tree typically has many secondary branches supported clear of the ground by the trunk, which typically contains woody tissue for strength, and vascular tissue to carry materials from one part of the tree to another. For most trees the trunk is surrounded by a layer of bark which serves as a protective barrier. Below
8208-762: Is gathered by ethnobotanists. This information can relay a great deal of information on how the land once was thousands of years ago and how it has changed over that time. The goals of plant ecology are to understand the causes of their distribution patterns, productivity, environmental impact, evolution, and responses to environmental change. Plants depend on certain edaphic (soil) and climatic factors in their environment but can modify these factors too. For example, they can change their environment's albedo , increase runoff interception, stabilise mineral soils and develop their organic content, and affect local temperature. Plants compete with other organisms in their ecosystem for resources. They interact with their neighbours at
8379-483: Is inelastic. Eventually the growth of a tree slows down and stops and it gets no taller. If damage occurs the tree may in time become hollow. Leaves are structures specialised for photosynthesis and are arranged on the tree in such a way as to maximise their exposure to light without shading each other. They are an important investment by the tree and may be thorny or contain phytoliths , lignins , tannins or poisons to discourage herbivory. Trees have evolved leaves in
8550-422: Is not available in the soil because of excess water. These root extensions are called pneumatophores , and are present, among others, in black mangrove and pond cypress. The main purpose of the trunk is to raise the leaves above the ground, enabling the tree to overtop other plants and outcompete them for light. It also transports water and nutrients from the roots to the aerial parts of the tree, and distributes
8721-656: Is not likely to become an endangered species in the foreseeable future in the absence of special protection and management efforts required by CESA. In February 2023, California governor Gavin Newsom 's administration proposed a budget trailer bill The Western Joshua Tree Conservation Act to focus on protecting the climate-threatened species and permitting development in the Southern California desert . The legislation requires conservation plan for this and other species that may be threatened by climate change and would authorize
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#17328374968088892-457: Is ready to eat when the process is complete, having been tenderised and flavoured by the smoke it has absorbed. In the cold process, the temperature is not allowed to rise above 100 °F (38 °C). The flavour of the food is enhanced but raw food requires further cooking. If it is to be preserved, meat should be cured before cold smoking. Wood has traditionally been used for fuel, especially in rural areas. In less developed nations it may be
9063-428: Is related to climatic conditions; growth normally ceases when conditions are either too cold or too dry. In readiness for the inactive period, trees form buds to protect the meristem , the zone of active growth. Before the period of dormancy, the last few leaves produced at the tip of a twig form scales. These are thick, small and closely wrapped and enclose the growing point in a waterproof sheath. Inside this bud there
9234-407: Is scarce. Frozen ground may limit water availability and conifers are often found in colder places at higher altitudes and higher latitudes than broad leaved trees. In conifers such as fir trees, the branches hang down at an angle to the trunk, enabling them to shed snow. In contrast, broad leaved trees in temperate regions deal with winter weather by shedding their leaves. When the days get shorter and
9405-451: Is still a major foundation of modern botany. Her books Plant Anatomy and Anatomy of Seed Plants have been key plant structural biology texts for more than half a century. The discipline of plant ecology was pioneered in the late 19th century by botanists such as Eugenius Warming , who produced the hypothesis that plants form communities , and his mentor and successor Christen C. Raunkiær whose system for describing plant life forms
9576-400: Is still given to these groups by botanists, and fungi (including lichens) and photosynthetic protists are usually covered in introductory botany courses. Palaeobotanists study ancient plants in the fossil record to provide information about the evolutionary history of plants . Cyanobacteria , the first oxygen-releasing photosynthetic organisms on Earth, are thought to have given rise to
9747-444: Is still in use today. The concept that the composition of plant communities such as temperate broadleaf forest changes by a process of ecological succession was developed by Henry Chandler Cowles , Arthur Tansley and Frederic Clements . Clements is credited with the idea of climax vegetation as the most complex vegetation that an environment can support and Tansley introduced the concept of ecosystems to biology. Building on
9918-400: Is that a tree has a woody trunk formed by secondary growth , meaning that the trunk thickens each year by growing outwards, in addition to the primary upwards growth from the growing tip . Under such a definition, herbaceous plants such as palms , bananas and papayas are not considered trees regardless of their height, growth form or stem girth. Certain monocots may be considered trees under
10089-495: Is the science of plant life and a branch of biology . A botanist , plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek word botanē ( βοτάνη ) meaning " pasture ", " herbs " " grass ", or " fodder "; Botanē is in turn derived from boskein ( Greek : βόσκειν ), "to feed" or "to graze ". Traditionally, botany has also included
10260-471: Is the first part of a seedling to emerge from the seed during the process of germination . This develops into a taproot which goes straight downwards. Within a few weeks lateral roots branch out of the side of this and grow horizontally through the upper layers of the soil. In most trees, the taproot eventually withers away and the wide-spreading laterals remain. Near the tip of the finer roots are single cell root hairs . These are in immediate contact with
10431-498: Is the study of ferns and allied plants. A number of other taxa of ranks varying from family to subgenus have terms for their study, including agrostology (or graminology) for the study of grasses, synantherology for the study of composites, and batology for the study of brambles. Study can also be divided by guild rather than clade or grade . Dendrology is the study of woody plants. Many divisions of biology have botanical subfields. These are commonly denoted by prefixing
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#173283749680810602-553: The Angiosperm Phylogeny Group to publish in 1998 a phylogeny of flowering plants, answering many of the questions about relationships among angiosperm families and species. The theoretical possibility of a practical method for identification of plant species and commercial varieties by DNA barcoding is the subject of active current research. Botany is divided along several axes. Some subfields of botany relate to particular groups of organisms. Divisions related to
10773-495: The Book of Plants , and Ibn Bassal 's The Classification of Soils . In the early 13th century, Abu al-Abbas al-Nabati , and Ibn al-Baitar (d. 1248) wrote on botany in a systematic and scientific manner. In the mid-16th century, botanical gardens were founded in a number of Italian universities. The Padua botanical garden in 1545 is usually considered to be the first which is still in its original location. These gardens continued
10944-417: The C 4 carbon fixation pathway for photosynthesis which avoid the losses resulting from photorespiration in the more common C 3 carbon fixation pathway. These biochemical strategies are unique to land plants. Phytochemistry is a branch of plant biochemistry primarily concerned with the chemical substances produced by plants during secondary metabolism . Some of these compounds are toxins such as
11115-570: The California Department of Fish and Wildlife to permit taking a western Joshua tree only under certain conditions. The legislation requires a fee of up to $ 2500 for the removal, relocation or trimming of limbs of a Western Joshua tree, including dead trees and limbs. This bill was passed by California lawmakers in June 2023 and went into effect on July 10, 2023. Different forms of the species are cultivated, including smaller plants native from
11286-503: The Cretaceous period. These began to displace the conifers during the Tertiary era (66 to 2 million years ago) when forests covered the globe. When the climate cooled 1.5 million years ago and the first of four glacial periods occurred, the forests retreated as the ice advanced. In the interglacials , trees recolonised the land that had been covered by ice, only to be driven back again in
11457-504: The alkaloid coniine from hemlock . Others, such as the essential oils peppermint oil and lemon oil are useful for their aroma, as flavourings and spices (e.g., capsaicin ), and in medicine as pharmaceuticals as in opium from opium poppies . Many medicinal and recreational drugs , such as tetrahydrocannabinol (active ingredient in cannabis ), caffeine , morphine and nicotine come directly from plants. Others are simple derivatives of botanical natural products. For example,
11628-410: The annual growth rings There may also be rays running at right angles to growth rings. These are vascular rays which are thin sheets of living tissue permeating the wood. Many older trees may become hollow but may still stand upright for many years. Trees do not usually grow continuously throughout the year but mostly have spurts of active expansion followed by periods of rest. This pattern of growth
11799-490: The autonymic subspecies Y. b. subsp. brevifolia , two other subspecies have been described: Y. b. subsp. herbertii (Webber's yucca or Herbert Joshua tree) and Y. b. subsp. jaegeriana (the Jaeger Joshua tree or Jaeger's Joshua tree or pygmae yucca), though both are sometimes treated as varieties or forms . Y.b. subsp. jaegeriana has also been treated as its own species. Joshua trees grow quickly for
11970-565: The auxin plant hormones by Kenneth V. Thimann in 1948 enabled regulation of plant growth by externally applied chemicals. Frederick Campion Steward pioneered techniques of micropropagation and plant tissue culture controlled by plant hormones . The synthetic auxin 2,4-dichlorophenoxyacetic acid or 2,4-D was one of the first commercial synthetic herbicides . 20th century developments in plant biochemistry have been driven by modern techniques of organic chemical analysis , such as spectroscopy , chromatography and electrophoresis . With
12141-626: The cell theory with Theodor Schwann and Rudolf Virchow and was among the first to grasp the significance of the cell nucleus that had been described by Robert Brown in 1831. In 1855, Adolf Fick formulated Fick's laws that enabled the calculation of the rates of molecular diffusion in biological systems. Building upon the gene-chromosome theory of heredity that originated with Gregor Mendel (1822–1884), August Weismann (1834–1914) proved that inheritance only takes place through gametes . No other cells can pass on inherited characters. The work of Katherine Esau (1898–1997) on plant anatomy
12312-444: The cellulose and lignin used to build their bodies, and secondary products like resins and aroma compounds . Plants and various other groups of photosynthetic eukaryotes collectively known as " algae " have unique organelles known as chloroplasts . Chloroplasts are thought to be descended from cyanobacteria that formed endosymbiotic relationships with ancient plant and algal ancestors. Chloroplasts and cyanobacteria contain
12483-427: The indigenous people of Canada in identifying edible plants from inedible plants. This relationship the indigenous people had with plants was recorded by ethnobotanists. Plant biochemistry is the study of the chemical processes used by plants. Some of these processes are used in their primary metabolism like the photosynthetic Calvin cycle and crassulacean acid metabolism . Others make specialised materials like
12654-544: The pine ( Pinus species) the bark exudes sticky resin which deters attackers whereas in rubber trees ( Hevea brasiliensis ) it is a milky latex that oozes out. The quinine bark tree ( Cinchona officinalis ) contains bitter substances to make the bark unpalatable. Large tree-like plants with lignified trunks in the Pteridophyta , Arecales , Cycadophyta and Poales such as the tree ferns, palms, cycads and bamboos have different structures and outer coverings. Although
12825-426: The pines , and flowering plants ) and the free-sporing cryptogams including ferns , clubmosses , liverworts , hornworts and mosses . Embryophytes are multicellular eukaryotes descended from an ancestor that obtained its energy from sunlight by photosynthesis . They have life cycles with alternating haploid and diploid phases. The sexual haploid phase of embryophytes, known as the gametophyte , nurtures
12996-520: The 12,000 years since the start of human agriculture, the number of trees worldwide has decreased by 46%. There are approximately 64,100 known tree species in the world. With 43% of all tree species, South America has the highest biodiversity, followed by Eurasia (22%), Africa (16%), North America (15%), and Oceania (11%). In suitable environments, such as the Daintree Rainforest in Queensland , or
13167-408: The 3-carbon sugar glyceraldehyde 3-phosphate (G3P). Glyceraldehyde 3-phosphate is the first product of photosynthesis and the raw material from which glucose and almost all other organic molecules of biological origin are synthesised. Some of the glucose is converted to starch which is stored in the chloroplast. Starch is the characteristic energy store of most land plants and algae, while inulin ,
13338-506: The Candollean system to reflect evolutionary relationships as distinct from mere morphological similarity. Botany was greatly stimulated by the appearance of the first "modern" textbook, Matthias Schleiden 's Grundzüge der Wissenschaftlichen Botanik , published in English in 1849 as Principles of Scientific Botany . Schleiden was a microscopist and an early plant anatomist who co-founded
13509-550: The English oak ( Quercus robur ) and 306 species of invertebrate on the Tasmanian oak ( Eucalyptus obliqua ). Non-native tree species provide a less biodiverse community, for example in the United Kingdom the sycamore ( Acer pseudoplatanus ), which originates from southern Europe, has few associated invertebrate species, though its bark supports a wide range of lichens, bryophytes and other epiphytes. Trees differ ecologically in
13680-533: The European bay tree ( Laurus nobilis ) and the California bay tree ( Umbellularia californica ) are used for flavouring food. Camellia sinensis , the source of tea, is a small tree but seldom reaches its full height, being heavily pruned to make picking the leaves easier. Wood smoke can be used to preserve food. In the hot smoking process the food is exposed to smoke and heat in a controlled environment. The food
13851-474: The Vegetable Kingdom at the start of chapter XII noted "The first and most important of the conclusions which may be drawn from the observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented." An important adaptive benefit of outcrossing is that it allows the masking of deleterious mutations in
14022-412: The above pair of categories gives rise to fields such as bryogeography (the study of the distribution of mosses). Different parts of plants also give rise to their own subfields, including xylology , carpology (or fructology) and palynology , these been the study of wood, fruit and pollen/spores respectively. Botany also overlaps on the one hand with agriculture, horticulture and silviculture, and on
14193-439: The ancestor of plants by entering into an endosymbiotic relationship with an early eukaryote, ultimately becoming the chloroplasts in plant cells. The new photosynthetic plants (along with their algal relatives) accelerated the rise in atmospheric oxygen started by the cyanobacteria , changing the ancient oxygen-free, reducing , atmosphere to one in which free oxygen has been abundant for more than 2 billion years. Among
14364-495: The arrival of warmer weather and the longer days associated with spring in temperate regions, growth starts again. The expanding shoot pushes its way out, shedding the scales in the process. These leave behind scars on the surface of the twig. The whole year's growth may take place in just a few weeks. The new stem is unlignified at first and may be green and downy. The Arecaceae (palms) have their leaves spirally arranged on an unbranched trunk. In some tree species in temperate climates,
14535-437: The bark functions as a protective barrier, it is itself attacked by boring insects such as beetles. These lay their eggs in crevices and the larvae chew their way through the cellulose tissues leaving a gallery of tunnels. This may allow fungal spores to gain admittance and attack the tree. Dutch elm disease is caused by a fungus ( Ophiostoma species) carried from one elm tree to another by various beetles. The tree reacts to
14706-447: The base of most food chains because they use the energy from the sun and nutrients from the soil and atmosphere, converting them into a form that can be used by animals. This is what ecologists call the first trophic level . The modern forms of the major staple foods , such as hemp , teff , maize, rice, wheat and other cereal grasses, pulses , bananas and plantains, as well as hemp , flax and cotton grown for their fibres, are
14877-444: The base, tapering to a sharp point; they are borne in a dense spiral arrangement at the apex of the stems. The leaf margins are white and serrated . Flowers typically appear from February to late April, in panicles 30 to 55 cm tall and 30 to 38 cm broad, the individual flowers erect, 4 to 7 cm tall, with six creamy white to green tepals . The tepals are lanceolate and are fused to the middle. The fused pistils are 3 cm tall and
15048-500: The blue-green pigment chlorophyll a . Chlorophyll a (as well as its plant and green algal-specific cousin chlorophyll b ) absorbs light in the blue-violet and orange/red parts of the spectrum while reflecting and transmitting the green light that we see as the characteristic colour of these organisms. The energy in the red and blue light that these pigments absorb is used by chloroplasts to make energy-rich carbon compounds from carbon dioxide and water by oxygenic photosynthesis ,
15219-446: The broader historical sense of botany include bacteriology , mycology (or fungology) and phycology - the study of bacteria, fungi and algae respectively - with lichenology as a subfield of mycology. The narrower sense of botany in the sense of the study of embryophytes (land plants) is disambiguated as phytology. Bryology is the study of mosses (and in the broader sense also liverworts and hornworts). Pteridology (or filicology)
15390-453: The characters may be artificial in keys designed purely for identification ( diagnostic keys ) or more closely related to the natural or phyletic order of the taxa in synoptic keys. By the 18th century, new plants for study were arriving in Europe in increasing numbers from newly discovered countries and the European colonies worldwide. In 1753, Carl Linnaeus published his Species Plantarum ,
15561-437: The clove tree ( Syzygium aromaticum ). Many trees have flowers rich in nectar which are attractive to bees. The production of forest honey is an important industry in rural areas of the developing world where it is undertaken by small-scale beekeepers using traditional methods. The flowers of the elder ( Sambucus ) are used to make elderflower cordial and petals of the plum ( Prunus spp. ) can be candied. Sassafras oil
15732-463: The cone for years waiting for a trigger event to liberate it. Fire stimulates release and germination of seeds of the jack pine , and also enriches the forest floor with wood ash and removes competing vegetation. Similarly, a number of angiosperms including Acacia cyclops and Acacia mangium have seeds that germinate better after exposure to high temperatures. The single extant species of Ginkgophyta ( Ginkgo biloba ) has fleshy seeds produced at
15903-461: The developing diploid embryo sporophyte within its tissues for at least part of its life, even in the seed plants, where the gametophyte itself is nurtured by its parent sporophyte. Other groups of organisms that were previously studied by botanists include bacteria (now studied in bacteriology ), fungi ( mycology ) – including lichen -forming fungi ( lichenology ), non- chlorophyte algae ( phycology ), and viruses ( virology ). However, attention
16074-459: The earliest was the Padua botanical garden . These gardens facilitated the academic study of plants. Efforts to catalogue and describe their collections were the beginnings of plant taxonomy and led in 1753 to the binomial system of nomenclature of Carl Linnaeus that remains in use to this day for the naming of all biological species. In the 19th and 20th centuries, new techniques were developed for
16245-423: The ease with which they can be found by herbivores. Tree apparency varies with a tree's size and semiochemical content, and with the extent to which it is concealed by nonhost neighbours from its insect pests . In ecosystems such as mangrove swamps, trees play a role in developing the habitat, since the roots of the mangrove trees reduce the speed of flow of tidal currents and trap water-borne sediment, reducing
16416-758: The eastern part of the species range. These smaller plants grow 2.5 m tall and branch when about 1 m tall. Red-shafted flickers make nests in the branches, which are later used by other birds. Prior to the twentieth century, Native Americans of the Mojave and western Sonoran Desert routinely used several parts of the Joshua tree as food and fiber (Cornett, J.W., 2018, Indian Uses of Desert Plants, Nature Trails Press, Palm Springs, CA). Leaf fibers were occasionally used to bind and manufacture sandals. Root sheaths were woven into baskets to add reddish-brown designs. Fruits were baked or boiled then eaten. Seeds were ground into flour and mixed with flour from other plant species. The flour
16587-419: The efforts of early humans to identify – and later cultivate – plants that were edible, poisonous, and possibly medicinal, making it one of the first endeavours of human investigation. Medieval physic gardens , often attached to monasteries , contained plants possibly having medicinal benefit. They were forerunners of the first botanical gardens attached to universities , founded from the 1540s onwards. One of
16758-643: The ends of short branches on female trees, and Gnetum , a tropical and subtropical group of gymnosperms produce seeds at the tip of a shoot axis. The earliest trees were tree ferns , horsetails and lycophytes , which grew in forests in the Carboniferous period. The first tree may have been Wattieza , fossils of which were found in New York state in 2007 dating back to the Middle Devonian (about 385 million years ago). Prior to this discovery, Archaeopteris
16929-453: The extensive earlier work of Alphonse de Candolle , Nikolai Vavilov (1887–1943) produced accounts of the biogeography , centres of origin , and evolutionary history of economic plants. Particularly since the mid-1960s there have been advances in understanding of the physics of plant physiological processes such as transpiration (the transport of water within plant tissues), the temperature dependence of rates of water evaporation from
17100-458: The far north of the northern hemisphere is moist taiga or northern coniferous forest (also called boreal forest). Taiga is the world's largest land biome , forming 29% of the world's forest cover. The long cold winter of the far north is unsuitable for plant growth and trees must grow rapidly in the short summer season when the temperature rises and the days are long. Light is very limited under their dense cover and there may be little plant life on
17271-478: The food produced by the leaves to all other parts, including the roots. In the case of angiosperms and gymnosperms, the outermost layer of the trunk is the bark , mostly composed of dead cells of phellem (cork). It provides a thick, waterproof covering to the living inner tissue. It protects the trunk against the elements, disease, animal attack and fire. It is perforated by a large number of fine breathing pores called lenticels , through which oxygen diffuses. Bark
17442-415: The forest floor, although fungi may abound. Similar woodland is found on mountains where the altitude causes the average temperature to be lower thus reducing the length of the growing season. Where rainfall is relatively evenly spread across the seasons in temperate regions, temperate broadleaf and mixed forest typified by species like oak, beech, birch and maple is found. Temperate forest is also found in
17613-563: The fossil record. It is widely regarded as a marker for the start of land plant evolution during the Ordovician period. The concentration of carbon dioxide in the atmosphere today is much lower than it was when plants emerged onto land during the Ordovician and Silurian periods. Many monocots like maize and the pineapple and some dicots like the Asteraceae have since independently evolved pathways like Crassulacean acid metabolism and
17784-423: The fruits and either discard the seeds, or swallow them so they pass through the gut to be deposited in the animal's droppings well away from the parent tree. The germination of some seeds is improved when they are processed in this way. Nuts may be gathered by animals such as squirrels that cache any not immediately consumed. Many of these caches are never revisited; the nut-casing softens with rain and frost, and
17955-451: The functional relationships between plants and their habitats – the environments where they complete their life cycles . Plant ecologists study the composition of local and regional floras , their biodiversity , genetic diversity and fitness , the adaptation of plants to their environment, and their competitive or mutualistic interactions with other species. Some ecologists even rely on empirical data from indigenous people that
18126-447: The genome of progeny. This beneficial effect is also known as hybrid vigor or heterosis. Once outcrossing is established, subsequent switching to inbreeding becomes disadvantageous since it allows expression of the previously masked deleterious recessive mutations, commonly referred to as inbreeding depression. Yucca brevifolia Yucca brevifolia (also known as the Joshua tree , yucca palm , tree yucca , and palm tree yucca )
18297-425: The geographical reach of the Mojave Desert, where it is considered one of the major indicator species for the desert. It occurs at elevations between 400 and 1,800 m (1,300 and 5,900 ft). Joshua trees are one of the species predicted to have their range reduced and shifted by climate change . Concern remains that they will be eliminated from Joshua Tree National Park, with ecological research suggesting
18468-444: The global carbon and water cycles and plant roots bind and stabilise soils, preventing soil erosion . Plants are crucial to the future of human society as they provide food, oxygen, biochemicals , and products for people, as well as creating and preserving soil. Historically, all living things were classified as either animals or plants and botany covered the study of all organisms not considered animals. Botanists examine both
18639-477: The ground underneath trees there is shade, and often there is undergrowth, leaf litter , and decaying wood that provide other habitat. Trees stabilise the soil, prevent rapid run-off of rain water, help prevent desertification, have a role in climate control and help in the maintenance of biodiversity and ecosystem balance. Many species of tree support their own specialised invertebrates . In their natural habitats, 284 different species of insect have been found on
18810-557: The ground, the roots branch and spread out widely; they serve to anchor the tree and extract moisture and nutrients from the soil . Above ground, the branches divide into smaller branches and shoots. The shoots typically bear leaves, which capture light energy and convert it into sugars by photosynthesis , providing the food for the tree's growth and development. Trees usually reproduce using seeds . Flowering plants have their seeds inside fruits, while conifers carry their seeds in cones, and tree ferns produce spores instead. Trees play
18981-419: The ground. Trees use a variety of methods of seed dispersal . Some rely on wind, with winged or plumed seeds. Others rely on animals, for example with edible fruits. Others again eject their seeds (ballistic dispersal), or use gravity so that seeds fall and sometimes roll. Seeds are the primary way that trees reproduce and their seeds vary greatly in size and shape. Some of the largest seeds come from trees, but
19152-412: The growth of the fungus by blocking off the xylem tissue carrying sap upwards and the branch above, and eventually the whole tree, is deprived of nourishment and dies. In Britain in the 1990s, 25 million elm trees were killed by this disease. The innermost layer of bark is known as the phloem and this is involved in the transport of the sap containing the sugars made by photosynthesis to other parts of
19323-446: The heartwood are blocked in some species. Heartwood is usually darker in colour than the sapwood. It is the dense central core of the trunk giving it rigidity. Three quarters of the dry mass of the xylem is cellulose , a polysaccharide , and most of the remainder is lignin, a complex polymer . A transverse section through a tree trunk or a horizontal core will show concentric circles of lighter or darker wood – tree rings. These rings are
19494-621: The important botanical questions of the 21st century are the role of plants as primary producers in the global cycling of life's basic ingredients: energy, carbon, oxygen, nitrogen and water, and ways that our plant stewardship can help address the global environmental issues of resource management , conservation , human food security , biologically invasive organisms , carbon sequestration , climate change , and sustainability . Virtually all staple foods come either directly from primary production by plants, or indirectly from animals that eat them. Plants and other photosynthetic organisms are at
19665-411: The inserted spigots; the sap is then heated to concentrate the flavour. Similarly in northern Europe the spring rise in the sap of the silver birch ( Betula pendula ) is tapped and collected, either to be drunk fresh or fermented into an alcoholic drink. In Alaska, the sap of the sweet birch ( Betula lenta ) is made into a syrup with a sugar content of 67%. Sweet birch sap is more dilute than maple sap;
19836-399: The inside. The newly created xylem is the sapwood . It is composed of water-conducting cells and associated cells which are often living, and is usually pale in colour. It transports water and minerals from the roots to the upper parts of the tree. The oldest, inner part of the sapwood is progressively converted into heartwood as new sapwood is formed at the cambium. The conductive cells of
20007-478: The internal functions and processes within plant organelles , cells, tissues, whole plants, plant populations and plant communities. At each of these levels, a botanist may be concerned with the classification ( taxonomy ), phylogeny and evolution , structure ( anatomy and morphology ), or function ( physiology ) of plant life. The strictest definition of "plant" includes only the "land plants" or embryophytes , which include seed plants (gymnosperms, including
20178-527: The largest tree, Sequoiadendron giganteum , produces one of the smallest tree seeds. The great diversity in tree fruits and seeds reflects the many different ways that tree species have evolved to disperse their offspring. For a tree seedling to grow into an adult tree it needs light. If seeds only fell straight to the ground, competition among the concentrated saplings and the shade of the parent would likely prevent it from flourishing. Many seeds such as birch are small and have papery wings to aid dispersal by
20349-403: The leaf shapes and sizes of ancient land plants . Ozone depletion can expose plants to higher levels of ultraviolet radiation-B (UV-B), resulting in lower growth rates. Moreover, information from studies of community ecology , plant systematics , and taxonomy is essential to understanding vegetation change , habitat destruction and species extinction . Inheritance in plants follows
20520-576: The leaf surface and the molecular diffusion of water vapour and carbon dioxide through stomatal apertures. These developments, coupled with new methods for measuring the size of stomatal apertures, and the rate of photosynthesis have enabled precise description of the rates of gas exchange between plants and the atmosphere. Innovations in statistical analysis by Ronald Fisher , Frank Yates and others at Rothamsted Experimental Station facilitated rational experimental design and data analysis in botanical research. The discovery and identification of
20691-576: The maintenance of biodiversity . Botany originated as herbalism , the study and use of plants for their possible medicinal properties . The early recorded history of botany includes many ancient writings and plant classifications. Examples of early botanical works have been found in ancient texts from India dating back to before 1100 BCE, Ancient Egypt , in archaic Avestan writings, and in works from China purportedly from before 221 BCE. Modern botany traces its roots back to Ancient Greece specifically to Theophrastus ( c. 371 –287 BCE),
20862-493: The major groups of organisms that carry out photosynthesis , a process that uses the energy of sunlight to convert water and carbon dioxide into sugars that can be used both as a source of chemical energy and of organic molecules that are used in the structural components of cells. As a by-product of photosynthesis, plants release oxygen into the atmosphere, a gas that is required by nearly all living things to carry out cellular respiration. In addition, they are influential in
21033-474: The mechanisms and control of gene expression during differentiation of plant cells and tissues . Botanical research has diverse applications in providing staple foods , materials such as timber , oil , rubber, fibre and drugs, in modern horticulture , agriculture and forestry , plant propagation , breeding and genetic modification , in the synthesis of chemicals and raw materials for construction and energy production, in environmental management , and
21204-407: The mixed podocarp and broadleaf forest of Ulva Island, New Zealand , forest is the more-or-less stable climatic climax community at the end of a plant succession, where open areas such as grassland are colonised by taller plants, which in turn give way to trees that eventually form a forest canopy. In cool temperate regions, conifers often predominate; a widely distributed climax community in
21375-409: The nearest Joshua tree. If it survives the rigors of the desert, it can live for several hundred years. The tallest trees reach about 15 m (49 ft). New plants can grow from seed , but in some populations, new stems grow from underground rhizomes that spread out around the parent tree. The evergreen leaves are dark green, linear, bayonet-shaped, 15 to 35 cm long, and 7 to 15 mm broad at
21546-516: The next glacial period. Trees are an important part of the terrestrial ecosystem , providing essential habitats including many kinds of forest for communities of organisms. Epiphytic plants such as ferns , some mosses, liverworts, orchids and some species of parasitic plants (e.g., mistletoe ) hang from branches; these along with arboreal lichens, algae, and fungi provide micro-habitats for themselves and for other organisms, including animals. Leaves, flowers and fruits are seasonally available. On
21717-596: The number of their male sexual organs. The 24th group, Cryptogamia , included all plants with concealed reproductive parts, mosses , liverworts , ferns , algae and fungi . Botany was originally a hobby for upper-class women. These women would collect and paint flowers and plants from around the world with scientific accuracy. The paintings were used to record many species that could not be transported or maintained in other environments. Marianne North illustrated over 900 species in extreme detail with watercolor and oil paintings. Her work and many other women's botany work
21888-419: The old lineages of flowering plants called basal angiosperms or paleodicots ; these include Amborella , Magnolia , nutmeg and avocado , while trees such as bamboo, palms and bananas are monocots . Wood gives structural strength to the trunk of most types of tree; this supports the plant as it grows larger. The vascular system of trees allows water, nutrients and other chemicals to be distributed around
22059-549: The only fuel available and collecting firewood is often a time-consuming task as it becomes necessary to travel further and further afield in the search for fuel. It is often burned inefficiently on an open fire. In more developed countries other fuels are available and burning wood is a choice rather than a necessity. Modern wood-burning stoves are very fuel efficient and new products such as wood pellets are available to burn. Botany Botany , also called plant science (or plant sciences ), plant biology or phytology ,
22230-448: The only one to develop, so they have unbranched trunks with large spirally arranged leaves. Some of the tree ferns, order Cyatheales , have tall straight trunks, growing up to 20 metres (66 ft), but these are composed not of wood but of rhizomes which grow vertically and are covered by numerous adventitious roots . The number of trees in the world, according to a 2015 estimate, is 3.04 trillion, of which 1.39 trillion (46%) are in
22401-504: The only survivor is the maidenhair tree Ginkgo biloba . This is considered to be a living fossil because it is virtually unchanged from the fossilised specimens found in Triassic deposits. During the Mesozoic (245 to 66 million years ago) the conifers flourished and became adapted to live in all the major terrestrial habitats. Subsequently, the tree forms of flowering plants evolved during
22572-911: The open grasslands of Queen Valley and Lost Horse Valley in Joshua Tree National Park . Other regions with large populations of the tree can be found northeast of Kingman, Arizona , in Mohave County ; and along U.S. 93 between the towns of Wickenburg and Wikieup , a route which has been designated the Joshua Tree Parkway of Arizona. The trees are abundant in Saddleback Butte State Park 135 kilometres (85 miles) north of Downtown Los Angeles in Los Angeles County's Antelope Valley . The common name Joshua tree apparently comes from Christian iconography. The Joshua tree
22743-447: The other hand with medicine and pharmacology, giving rise to fields such as agronomy , horticultural botany, phytopathology and phytopharmacology . The study of plants is vital because they underpin almost all animal life on Earth by generating a large proportion of the oxygen and food that provide humans and other organisms with aerobic respiration with the chemical energy they need to exist. Plants, algae and cyanobacteria are
22914-404: The outcome of prehistoric selection over thousands of years from among wild ancestral plants with the most desirable characteristics. Botanists study how plants produce food and how to increase yields, for example through plant breeding , making their work important to humanity's ability to feed the world and provide food security for future generations. Botanists also study weeds, which are
23085-791: The pain killer aspirin is the acetyl ester of salicylic acid , originally isolated from the bark of willow trees, and a wide range of opiate painkillers like heroin are obtained by chemical modification of morphine obtained from the opium poppy . Popular stimulants come from plants, such as caffeine from coffee, tea and chocolate, and nicotine from tobacco. Most alcoholic beverages come from fermentation of carbohydrate -rich plant products such as barley (beer), rice ( sake ) and grapes (wine). Native Americans have used various plants as ways of treating illness or disease for thousands of years. This knowledge Native Americans have on plants has been recorded by enthnobotanists and then in turn has been used by pharmaceutical companies as
23256-582: The plant and reduce water loss. Both the production of wood and the production of cork are forms of secondary growth. Trees are either evergreen , having foliage that persists and remains green throughout the year, or deciduous , shedding their leaves at the end of the growing season and then having a dormant period without foliage. Most conifers are evergreens, but larches ( Larix and Pseudolarix ) are deciduous, dropping their needles each autumn, and some species of cypress ( Glyptostrobus , Metasequoia and Taxodium ) shed small leafy shoots annually in
23427-408: The plant, and without it trees would not be able to grow as large as they do. Trees need to draw water high up the stem through the xylem from the roots by capillary action , as water continually evaporates from the leaves in the process of transpiration . If insufficient water is available the leaves will die. The three main parts of trees include the root, stem, and leaves; they are integral parts of
23598-479: The plants with a highly fermentable sugar or oil content that are used as sources of biofuels , important alternatives to fossil fuels , such as biodiesel . Sweetgrass was used by Native Americans to ward off bugs like mosquitoes . These bug repelling properties of sweetgrass were later found by the American Chemical Society in the molecules phytol and coumarin . Plant ecology is the science of
23769-486: The pollen either fails to reach the stigma or fails to germinate and produce male gametes . This is one of several methods used by plants to promote outcrossing . In many land plants the male and female gametes are produced by separate individuals. These species are said to be dioecious when referring to vascular plant sporophytes and dioicous when referring to bryophyte gametophytes . Charles Darwin in his 1878 book The Effects of Cross and Self-Fertilization in
23940-442: The polymer cutin which is found in the plant cuticle that protects land plants from drying out. Plants synthesise a number of unique polymers like the polysaccharide molecules cellulose , pectin and xyloglucan from which the land plant cell wall is constructed. Vascular land plants make lignin , a polymer used to strengthen the secondary cell walls of xylem tracheids and vessels to keep them from collapsing when
24111-545: The practical value of earlier "physic gardens", often associated with monasteries, in which plants were cultivated for suspected medicinal uses. They supported the growth of botany as an academic subject. Lectures were given about the plants grown in the gardens. Botanical gardens came much later to northern Europe; the first in England was the University of Oxford Botanic Garden in 1621. German physician Leonhart Fuchs (1501–1566)
24282-435: The rest, many are gymnosperms or softwood trees; these include conifers , cycads , ginkgophytes and gnetales , which produce seeds which are not enclosed in fruits, but in open structures such as pine cones , and many have tough waxy leaves, such as pine needles. Most angiosperm trees are eudicots , the "true dicotyledons", so named because the seeds contain two cotyledons or seed leaves. There are also some trees among
24453-461: The rise of the related molecular-scale biological approaches of molecular biology , genomics , proteomics and metabolomics , the relationship between the plant genome and most aspects of the biochemistry, physiology, morphology and behaviour of plants can be subjected to detailed experimental analysis. The concept originally stated by Gottlieb Haberlandt in 1902 that all plant cells are totipotent and can be grown in vitro ultimately enabled
24624-426: The roots and helps protect the trees against predators and pathogens. It can also limit damage done to a tree by pollution as the fungus accumulate heavy metals within its tissues. Fossil evidence shows that roots have been associated with mycorrhizal fungi since the early Paleozoic , four hundred million years ago, when the first vascular plants colonised dry land. Some trees such as Alder ( Alnus species) have
24795-676: The same fundamental principles of genetics as in other multicellular organisms. Gregor Mendel discovered the genetic laws of inheritance by studying inherited traits such as shape in Pisum sativum ( peas ). What Mendel learned from studying plants has had far-reaching benefits outside of botany. Similarly, " jumping genes " were discovered by Barbara McClintock while she was studying maize. Nevertheless, there are some distinctive genetic differences between plants and other organisms. Species boundaries in plants may be weaker than in animals, and cross species hybrids are often possible. A familiar example
24966-461: The seeds on the surface of water. Mangroves often grow in water and some species have buoyant fruits with seeds that start germinating before they detach from the parent tree. These float on the water and may become lodged on emerging mudbanks and successfully take root. Other seeds, such as apple pips and plum stones, have fleshy receptacles and smaller fruits like hawthorns have seeds enclosed in edible tissue; animals including mammals and birds eat
25137-433: The soil particles and can absorb water and nutrients such as potassium in solution. The roots require oxygen to respire and only a few species such as mangroves and the pond cypress ( Taxodium ascendens ) can live in permanently waterlogged soil. In the soil, the roots encounter the hyphae of fungi. Many of these are known as mycorrhiza and form a mutualistic relationship with the tree roots. Some are specific to
25308-526: The southern hemisphere, as for example in the Eastern Australia temperate forest, characterised by Eucalyptus forest and open acacia woodland. In tropical regions with a monsoon or monsoon-like climate, where a drier part of the year alternates with a wet period as in the Amazon rainforest , different species of broad-leaved trees dominate the forest, some of them being deciduous. In tropical regions with
25479-467: The start of the dry season. Many deciduous trees flower before the new leaves emerge. A few trees do not have true leaves but instead have structures with similar external appearance such as Phylloclades – modified stem structures – as seen in the genus Phyllocladus . Trees can be pollinated either by wind or by animals, mostly insects. Many angiosperm trees are insect pollinated. Wind pollination may take advantage of increased wind speeds high above
25650-421: The stigma cavity is surrounded by lobes. The semi-fleshy fruit that is produced is green-brown, elliptical, and contains many flat seeds. Joshua trees usually do not branch until after they bloom (though branching may also occur if the growing tip is destroyed by the yucca-boring weevil ), and they do not bloom every year. Like most desert plants, their blooming depends on rainfall at the proper time. They also need
25821-590: The study of fungi and algae by mycologists and phycologists respectively, with the study of these three groups of organisms remaining within the sphere of interest of the International Botanical Congress . Nowadays, botanists (in the strict sense) study approximately 410,000 species of land plants , including some 391,000 species of vascular plants (of which approximately 369,000 are flowering plants ) and approximately 20,000 bryophytes . Botany originated in prehistory as herbalism with
25992-450: The study of plants, including methods of optical microscopy and live cell imaging , electron microscopy , analysis of chromosome number , plant chemistry and the structure and function of enzymes and other proteins . In the last two decades of the 20th century, botanists exploited the techniques of molecular genetic analysis , including genomics and proteomics and DNA sequences to classify plants more accurately. Modern botany
26163-471: The study of plants. In 1665, using an early microscope, Polymath Robert Hooke discovered cells (a term he coined) in cork , and a short time later in living plant tissue. During the 18th century, systems of plant identification were developed comparable to dichotomous keys , where unidentified plants are placed into taxonomic groups (e.g. family, genus and species) by making a series of choices between pairs of characters . The choice and sequence of
26334-410: The surviving seeds germinate in the spring. Pine cones may similarly be hoarded by red squirrels , and grizzly bears may help to disperse the seed by raiding squirrel caches. The seeds of conifers, the largest group of gymnosperms, are enclosed in a cone and most species have seeds that are light and papery that can be blown considerable distances once free from the cone. Sometimes the seed remains in
26505-660: The temperature begins to decrease, the leaves no longer make new chlorophyll and the red and yellow pigments already present in the blades become apparent. Synthesis in the leaf of a plant hormone called auxin also ceases. This causes the cells at the junction of the petiole and the twig to weaken until the joint breaks and the leaf floats to the ground. In tropical and subtropical regions, many trees keep their leaves all year round. Individual leaves may fall intermittently and be replaced by new growth but most leaves remain intact for some time. Other tropical species and those in arid regions may shed all their leaves annually, such as at
26676-407: The tree, but some tree species have evolved roots that are aerial . The common purposes for aerial roots may be of two kinds, to contribute to the mechanical stability of the tree, and to obtain oxygen from air. An instance of mechanical stability enhancement is the red mangrove that develops prop roots that loop out of the trunk and branches and descend vertically into the mud. A similar structure
26847-431: The tree. It is a soft spongy layer of living cells, some of which are arranged end to end to form tubes. These are supported by parenchyma cells which provide padding and include fibres for strengthening the tissue. Inside the phloem is a layer of undifferentiated cells one cell thick called the vascular cambium layer. The cells are continually dividing, creating phloem cells on the outside and wood cells known as xylem on
27018-562: The trees' dispersal. However, ground squirrels are very effective at moving the seeds long distances. In March 2022, California Department of Fish and Wildlife conducted a status review of the Western Joshua Tree to determine whether to list the species as threatened under the California Endangered Species Act (CESA). The study showed that the largest threat to Yucca brevifolia was wildfires, that wildfires were
27189-492: The tropics or sub-tropics , 0.61 trillion (20%) in the temperate zones , and 0.74 trillion (24%) in the coniferous boreal forests . The estimate is about eight times higher than previous estimates, and is based on tree densities measured on over 400,000 plots. It remains subject to a wide margin of error, not least because the samples are mainly from Europe and North America. The estimate suggests that about 15 billion trees are cut down annually and about 5 billion are planted. In
27360-526: The use of genetic engineering experimentally to knock out a gene or genes responsible for a specific trait, or to add genes such as GFP that report when a gene of interest is being expressed. These technologies enable the biotechnological use of whole plants or plant cell cultures grown in bioreactors to synthesise pesticides , antibiotics or other pharmaceuticals , as well as the practical application of genetically modified crops designed for traits such as improved yield. Modern morphology recognises
27531-403: The vascular system which interconnects all the living cells. In trees and other plants that develop wood, the vascular cambium allows the expansion of vascular tissue that produces woody growth. Because this growth ruptures the epidermis of the stem, woody plants also have a cork cambium that develops among the phloem. The cork cambium gives rise to thickened cork cells to protect the surface of
27702-427: The water depth and creating suitable conditions for further mangrove colonisation. Thus mangrove swamps tend to extend seawards in suitable locations. Mangrove swamps also provide an effective buffer against the more damaging effects of cyclones and tsunamis. Trees are the source of many of the world's best known fleshy fruits. Apples, pears, plums, cherries and citrus are all grown commercially in temperate climates and
27873-435: The wind. Ash trees and maples have larger seeds with blade shaped wings which spiral down to the ground when released. The kapok tree has cottony threads to catch the breeze. The flame tree Delonix regia shoots its seeds through the air when the two sides of its long pods crack apart explosively on drying. The miniature cone-like catkins of alder trees produce seeds that contain small droplets of oil that help disperse
28044-485: The word plant (e.g. plant taxonomy, plant ecology, plant anatomy, plant morphology, plant systematics, plant ecology), or prefixing or substituting the prefix phyto- (e.g. phytochemistry , phytogeography ). The study of fossil plants is palaeobotany . Other fields are denoted by adding or substituting the word botany (e.g. systematic botany ). Phytosociology is a subfield of plant ecology that classifies and studies communities of plants. The intersection of fields from
28215-578: The world in Mojave National Preserve were killed in the Dome Fire in August 2020. Also, concern exists about the ability of the species to migrate to favorable climates due to the extinction of the giant Shasta ground sloth ( Nothrotheriops shastensis ) 13,000 years ago; ground sloth dung has been found to contain Joshua tree leaves, fruits, and seeds, suggesting that the sloths might have been key to
28386-463: The world, forests are shrinking as trees are cleared to increase the amount of land available for agriculture. Because of their longevity and usefulness, trees have always been revered, with sacred groves in various cultures, and they play a role in many of the world's mythologies . Although "tree" is a common word, there is no universally recognised precise definition of what a tree is, either botanically or in common language. In its broadest sense,
28557-766: The world, fruit is gathered from forest trees for consumption. Many trees bear edible nuts which can loosely be described as being large, oily kernels found inside a hard shell. These include coconuts ( Cocos nucifera ), Brazil nuts ( Bertholletia excelsa ), pecans ( Carya illinoinensis ), hazel nuts ( Corylus ), almonds ( Prunus dulcis ), walnuts ( Juglans regia ), pistachios ( Pistacia vera ) and many others. They are high in nutritive value and contain high-quality protein, vitamins and minerals as well as dietary fibre. A variety of nut oils are extracted by pressing for culinary use; some such as walnut, pistachio and hazelnut oils are prized for their distinctive flavours, but they tend to spoil quickly. In temperate climates there
28728-479: Was noted in the review. The review concluded: Based on the criteria described above, the best scientific information available to the Department at this time indicates that western Joshua tree is not in serious danger of becoming extinct throughout all, or a significant portion, of its range due to one or more causes, including loss of habitat, change in habitat, overexploitation, predation, competition, or disease, and
28899-399: Was one of "the three German fathers of botany", along with theologian Otto Brunfels (1489–1534) and physician Hieronymus Bock (1498–1554) (also called Hieronymus Tragus). Fuchs and Brunfels broke away from the tradition of copying earlier works to make original observations of their own. Bock created his own system of plant classification. Physician Valerius Cordus (1515–1544) authored
29070-412: Was the beginning of popularizing botany to a wider audience. Increasing knowledge of plant anatomy , morphology and life cycles led to the realisation that there were more natural affinities between plants than the artificial sexual system of Linnaeus. Adanson (1763), de Jussieu (1789), and Candolle (1819) all proposed various alternative natural systems of classification that grouped plants using
29241-484: Was the earliest known tree. Both of these reproduced by spores rather than seeds and are considered to be links between ferns and the gymnosperms which evolved in the Triassic period. The gymnosperms include conifers, cycads, gnetales and ginkgos and these may have appeared as a result of a whole genome duplication event which took place about 319 million years ago. Ginkgophyta was once a widespread diverse group of which
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