54-770: Albulidae is a family of fish , commonly known as the bonefishes , that are popular as game fish in Florida , select locations in the South Pacific and the Bahamas (where two bonefish are featured on the 10-cent coin) and elsewhere. The family is small, with 11 species in 3 genera. Presently, the bonefishes are in their own order : Albuliformes / ˈ æ l b j ə l ɪ f ɔːr m iː z / . The families Halosauridae and Notacanthidae were previously classified in this order, but are now, according to FishBase , given their own order Notacanthiformes . The largest bonefish caught in
108-407: A clustering habit results. Exclusively sympodial genera include many of the rattans , Guihaia , and Rhapis . Several palm genera have both solitary and clustering members. Palms which are usually solitary may grow in clusters and vice versa. Palms have large, evergreen leaves that are either palmately ('fan-leaved') or pinnately ('feather-leaved') compound and spirally arranged at the top of
162-447: A consensus over time. The naming of families is codified by various international bodies using the following suffixes: Name changes at the family level are regulated by the codes of nomenclature. For botanical families, some traditional names like Palmae ( Arecaceae ), Cruciferae ( Brassicaceae ), and Leguminosae ( Fabaceae ) are conserved alongside their standardized -aceae forms due to their historical significance and widespread use in
216-446: A family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community for extended periods. The continual publication of new data and diverse opinions plays a crucial role in facilitating adjustments and ultimately reaching
270-471: A hundred taxonomic publications. Such descriptions typically result from either the discovery of organisms with unique combinations of characters that do not fit existing families, or from phylogenetic analyses that reveal the need for reclassification. The taxonomic term familia was first used by French botanist Pierre Magnol in his Prodromus historiae generalis plantarum, in quo familiae plantarum per tabulas disponuntur (1689) where he called
324-458: A large amount of the human diet and several other human uses, both by absolute amount produced and by number of species domesticated . This is far higher than almost any other plant family, sixth out of domesticated crops in the human diet, and first in total economic value produced – sharing the top spot with the Poaceae and Fabaceae . These human uses have also spread many Arecaceae species around
378-509: A significant practical role in biological education and research. They provide an efficient framework for teaching taxonomy, as they group organisms with general similarities while remaining specific enough to be useful for identification purposes. For example, in botany, learning the characteristics of major plant families helps students identify related species across different geographic regions, since families often have worldwide distribution patterns. In many groups of organisms, families serve as
432-617: A variety of different habitats. Their diversity is highest in wet, lowland forests. South America , the Caribbean , and areas of the South Pacific and southern Asia are regions of concentration. Colombia may have the highest number of palm species in one country. There are some palms that are also native to desert areas such as the Arabian Peninsula and parts of northwestern Mexico. Only about 130 palm species naturally grow entirely beyond
486-609: A water source, and in desert oases such as the date palm . A few palms are adapted to extremely basic lime soils, while others are similarly adapted to extreme potassium deficiency and toxicity of heavy metals in serpentine soils . Palms are a monophyletic group of plants, meaning the group consists of a common ancestor and all its descendants. Extensive taxonomic research on palms began with botanist H.E. Moore , who organized palms into 15 major groups based mostly on general morphological characteristics. The following classification, proposed by N.W. Uhl and J. Dransfield in 1987,
540-505: Is a family of perennial , flowering plants in the monocot order Arecales . Their growth form can be climbers , shrubs , tree -like and stemless plants, all commonly known as palms . Those having a tree-like form are colloquially called palm trees . Currently, 181 genera with around 2,600 species are known, most of which are restricted to tropical and subtropical climates. Most palms are distinguished by their large, compound, evergreen leaves, known as fronds , arranged at
594-508: Is a revision of Moore's classification that organizes palms into 6 subfamilies. A few general traits of each subfamily are listed below. The Phytelephantoideae is the sixth subfamily of Arecaceae in N.W. Uhl and J. Dransfield's 1987 classification. Members of this group have distinct monopodial flower clusters. Other distinct features include a gynoecium with five to 10 joined carpels, and flowers with more than three parts per whorl. Fruits are multiple-seeded and have multiple parts. From
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#1733106700425648-595: Is from Dravidian *aṭ-ay-kkāy ("areca nut"). The suffix -aceae is the feminine plural of the Latin -āceus ("resembling"). Palm originates from Latin palma semantically overlapping with sense of " hand front " (due to similar splayed shape) ultimately from Proto-Indo-European * pl̥h₂meh₂ , a direct descendant folm once existed in Old English. Whether as shrubs, tree-like, or vines, palms have two methods of growth: solitary or clustered. The common representation
702-496: Is often called "anomalous secondary growth". The Arecaceae are notable among monocots for their height and for the size of their seeds, leaves, and inflorescences. Ceroxylon quindiuense , Colombia's national "tree", is the tallest monocot in the world, reaching up to 60 metres (197 ft) tall. The coco de mer ( Lodoicea maldivica ) has the largest seeds of any plant, 40–50 centimetres (16–20 in) in diameter and weighing 15–30 kilograms (33–66 lb) each ( coconuts are
756-662: Is possible north of subtropical climates, and some higher latitude locales such as Ireland, Scotland, England, and the Pacific Northwest feature a few palms in protected locations and microclimates . In the United States , there are at least 12 native palm species, mostly occurring in the states of the Deep South and Florida . Palms inhabit a variety of ecosystems. More than two-thirds of palm species live in humid moist forests, where some species grow tall enough to form part of
810-460: Is that of a solitary shoot ending in a crown of leaves. This monopodial character may be exhibited by prostrate, trunkless, and trunk-forming members. Some common palms restricted to solitary growth include Washingtonia and Roystonea . Palms may instead grow in sparse though dense clusters. The trunk develops an axillary bud at a leaf node, usually near the base, from which a new shoot emerges. The new shoot, in turn, produces an axillary bud and
864-569: The Prodromus of Augustin Pyramus de Candolle and the Genera Plantarum of George Bentham and Joseph Dalton Hooker this word ordo was used for what now is given the rank of family. Families serve as valuable units for evolutionary, paleontological, and genetic studies due to their relatively greater stability compared to lower taxonomic levels like genera and species. Families play
918-559: The Dasypogonaceae . For a phylogenetic tree of the family, see the list of Arecaceae genera . The Arecaceae were the first modern family of monocots to appear in the fossil record around 80 million years ago (Mya), during the late Cretaceous period. The first modern species, such as Nypa fruticans and Acrocomia aculeata , appeared 69 Mya, as evidenced by fossil Nypa pollen. Palms appear to have undergone an early period of adaptive radiation . By 60 Mya, many of
972-569: The canopy and shorter ones form part of the understory . Some species form pure stands in areas with poor drainage or regular flooding, including Raphia hookeri which is common in coastal freshwater swamps in West Africa. Other palms live in tropical mountain habitats above 1 thousand metres (3 thousand feet), such as those in the genus Ceroxylon native to the Andes . Palms may also live in grasslands and scrublands, usually associated with
1026-447: The -idae suffix for animal family names, derived from the Greek 'eidos' meaning 'resemblance' or 'like'. The adoption of this naming convention helped establish families as an important taxonomic rank. By the mid-1800s, many of Linnaeus's broad genera were being elevated to family status to accommodate the rapidly growing number of newly discovered species. In nineteenth-century works such as
1080-640: The Ceroxyloideae and Phytelephantoideae. Studies have suggested the lack of a fully resolved hypothesis for the relationships within the family is due to a variety of factors, including difficulties in selecting appropriate outgroups, homoplasy in morphological character states, slow rates of molecular evolution important for the use of standard DNA markers , and character polarization. However, hybridization has been observed among Orbignya and Phoenix species, and using chloroplast DNA in cladistic studies may produce inaccurate results due to maternal inheritance of
1134-466: The Florida record being 16 pounds 3 oz (7.34 kg). The bonefishes are brackish or saltwater fish typically living in estuaries and travelling out to sea to spawn on a lunar cycle. They feed in the shallow sand and mud flats, on benthic organisms , such as worms , mollusks , shrimp and crabs . They use their conical-shaped snouts to root out their prey and can often be seen with their tails out of
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#17331067004251188-558: The IUCN, and was published in 1996. The rarest palm known is Hyophorbe amaricaulis . The only living individual remains at the Botanic Gardens of Curepipe in Mauritius . Some pests are specialists to particular taxa . Pests that attack a variety of species of palms include: The palm branch was a symbol of triumph and victory in classical antiquity . The Romans rewarded champions of
1242-520: The Middle East and North Africa. A wide range of common products and foods are derived from palms. In contemporary times, palms are also widely used in landscaping. In many historical cultures, because of their importance as food, palms were symbols for such ideas as victory, peace, and fertility. The word Arecaceae is derived from the word areca with the suffix "-aceae". Areca is derived from Portuguese , via Malayalam അടയ്ക്ക ( aṭaykka ), which
1296-565: The Rathayatra carts which carry the deities of Krishna and his family members in the cart festival of Jagganath Puri in India are marked with the emblem of a palm tree. Specifically it is the symbol of Krishna's brother, Baladeva. In 1840, the American geologist Edward Hitchcock (1793–1864) published the first tree-like paleontology chart in his Elementary Geology , with two separate trees of life for
1350-547: The Western Hemisphere is a 16-pound, 3 ounce example caught off Islamorada, Florida , on March 19, 2007. The bonefishes' closest relatives are the tarpons and ladyfishes in the order Elopiformes . Bonefishes are unlike tarpons in that their mouths are under the snout rather than the end of it. Like tarpons and ladyfishes, bonefishes can breathe air via a modified swim bladder and are found in brackish waters . Bonefish larvae are leptocephalic . The slender body of
1404-530: The area. The sale of seeds to nurseries and collectors is another threat, as the seeds of popular palms are sometimes harvested directly from the wild. In 2006, at least 100 palm species were considered endangered, and nine species have been reported as recently extinct. However, several factors make palm conservation more difficult. Palms live in almost every type of warm habitat and have tremendous morphological diversity. Most palm seeds lose viability quickly, and they cannot be preserved in low temperatures because
1458-407: The base. The stamens generally number six, with filaments that may be separate, attached to each other, or attached to the pistil at the base. The fruit is usually a single- seeded drupe (sometimes berry-like) but some genera (e.g., Salacca ) may contain two or more seeds in each fruit. Like all monocots , palms do not have the ability to increase the width of a stem ( secondary growth ) via
1512-407: The bonefish is silver, with a blue to green tinted back. On the upper half of the body there are dark streaks with cross bands connecting to the lateral line . The body is rounded with a long, slightly downturned snout. The dorsal and caudal fins are black. Bonefish vary in adult length from 40–100 cm depending on species. The average size of a bonefish is from 3 to 5 pounds (1–2 kg) with
1566-466: The chloroplast DNA. Chemical and molecular data from non-organelle DNA, for example, could be more effective for studying palm phylogeny. Recently, nuclear genomes and transcriptomes have been used to reconstruct the phylogeny of palms. This has revealed, for example, that a whole-genome duplication event occurred early in the evolution of the Arecaceae lineage, that was not experienced by its sister clade,
1620-493: The cold kills the embryo. Using botanical gardens for conservation also presents problems, since they can rarely house more than a few plants of any species or truly imitate the natural setting. There is also the risk that cross-pollination can lead to hybrid species. The Palm Specialist Group of the World Conservation Union (IUCN) began in 1984, and has performed a series of three studies to find basic information on
1674-721: The deserts, it also created a more amenable habitat for the people to live in by providing shade and protection from the desert winds. An indication of the importance of palms in ancient times is that they are mentioned more than 30 times in the Bible , and at least 22 times in the Quran . The Torah also references the "70 date palm trees", which symbolize the 70 aspects of Torah that are revealed to those who "eat of its fruit." Arecaceae have great economic importance, including coconut products, oils, dates, palm syrup , ivory nuts, carnauba wax, rattan cane, raffia, and palm wood . This family supplies
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1728-652: The games and celebrated military successes with palm branches. Early Christians used the palm branch to symbolize the victory of the faithful over enemies of the soul, as in the Palm Sunday festival celebrating the triumphal entry of Jesus Christ into Jerusalem . In Judaism , the palm represents peace and plenty, and is one of the Four Species of Sukkot ; the palm may also symbolize the Tree of Life in Kabbalah . The canopies of
1782-500: The habitat, and those with small habitat ranges are most vulnerable to them. The harvesting of heart of palm, a delicacy in salads, also poses a threat because it is derived from the palm's apical meristem , a vital part of the palm that cannot be regrown (except in domesticated varieties, e.g. of peach palm ). The use of rattan palms in furniture has caused a major population decrease in these species that has negatively affected local and international markets, as well as biodiversity in
1836-432: The history of the Middle East and North Africa. In the text "Date Palm Products" (1993), W.H. Barreveld wrote: One could go as far as to say that, had the date palm not existed, the expansion of the human race into the hot and barren parts of the "old" world would have been much more restricted. The date palm not only provided a concentrated energy food, which could be easily stored and carried along on long journeys across
1890-406: The literature. Family names are typically formed from the stem of a type genus within the family. In zoology, when a valid family name is based on a genus that is later found to be a junior synonym , the family name may be maintained for stability if it was established before 1960. In botany, some family names that were found to be junior synonyms have been conserved due to their widespread use in
1944-673: The modern phylogenomic data, the Phytelephantoideae are tribe in the Ceroxyloideae subfamily. Currently, few extensive phylogenetic studies of the Arecaceae exist. In 1997, Baker et al. explored subfamily and tribe relationships using chloroplast DNA from 60 genera from all subfamilies and tribes. The results strongly showed the Calamoideae are monophyletic, and Ceroxyloideae and Coryphoideae are paraphyletic. The relationships of Arecoideae are uncertain, but they are possibly related to
1998-660: The modern, specialized genera of palms appeared and became widespread and common, much more widespread than their range today. Because palms separated from the monocots earlier than other families, they developed more intrafamilial specialization and diversity. By tracing back these diverse characteristics of palms to the basic structures of monocots, palms may be valuable in studying monocot evolution. Several species of palms have been identified from flowers preserved in amber, including Palaeoraphe dominicana and Roystonea palaea . Fossil evidence of them can also be found in samples of petrified palmwood . The relationship between
2052-907: The primary level for taxonomic identification keys, making them particularly valuable for field guides and systematic work as they often represent readily recognizable groups of related organisms with shared characteristics. In ecological and biodiversity research, families frequently serve as the foundational level for identification in survey work and environmental studies. This is particularly useful because families often share life history traits or occupy similar ecological niches . Some families show strong correlations between their taxonomic grouping and ecological functions, though this relationship varies among different groups of organisms. The stability of family names has practical importance for applied biological work, though this stability faces ongoing challenges from new scientific findings. Modern molecular studies and phylogenetic analyses continue to refine
2106-488: The ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae , but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists . There are not strict regulations for outlining or acknowledging
2160-399: The same kind of vascular cambium found in non-monocot woody plants . This explains the cylindrical shape of the trunk (almost constant diameter) that is often seen in palms, unlike in ring -forming trees. However, many palms, like some other monocots, do have secondary growth, although because it does not arise from a single vascular cambium producing xylem inwards and phloem outwards, it
2214-486: The scientific literature. The family-group in zoological nomenclature includes several ranks: superfamily (-oidea), family (-idae), subfamily (-inae), and tribe (-ini). Under the principle of coordination, a name established at any of these ranks can be moved to another rank while retaining its original authorship and date, requiring only a change in suffix to reflect its new rank. New family descriptions are relatively rare in taxonomy, occurring in fewer than one in
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2268-486: The second largest). Raffia palms ( Raphia spp.) have the largest leaves of any plant, up to 25 metres (82 ft) long and 3 metres (10 ft) wide. The Corypha species have the largest inflorescence of any plant, up to 7.5 metres (25 ft) tall and containing millions of small flowers. Calamus stems can reach 200 metres (656 ft) in length. Most palms are native to tropical and subtropical climates. Palms thrive in moist and hot climates but can be found in
2322-567: The seventy-six groups of plants he recognised in his tables families ( familiae ). The concept of rank at that time was not yet settled, and in the preface to the Prodromus Magnol spoke of uniting his families into larger genera , which is far from how the term is used today. In his work Philosophia Botanica published in 1751, Carl Linnaeus employed the term familia to categorize significant plant groups such as trees , herbs , ferns , palms , and so on. Notably, he restricted
2376-476: The status of palms in the wild, use of wild palms, and palms under cultivation. Two projects on palm conservation and use supported by the World Wildlife Fund took place from 1985 to 1990 and 1986–1991, in the American tropics and southeast Asia, respectively. Both studies produced copious new data and publications on palms. Preparation of a global action plan for palm conservation began in 1991, supported by
2430-411: The stem. The leaves have a tubular sheath at the base that usually splits open on one side at maturity. The inflorescence is a spadix or spike surrounded by one or more bracts or spathes that become woody at maturity. The flowers are generally small and white, radially symmetric, and can be either uni- or bisexual. The sepals and petals usually number three each and may be distinct or joined at
2484-476: The subfamilies is shown in the following cladogram: Calamoideae Nypoideae Coryphoideae Ceroxyloideae Arecoideae Evidence for cultivation of the date palm by Mesopotamians and other Middle Eastern peoples exists from more than 5,000 years ago, in the form of date wood, pits for storing dates, and other remains of the date palm in Mesopotamian sites. The date palm had a significant effect on
2538-479: The top of an unbranched stem, except for the Hyphaene genus, who has branched palms. However, palms exhibit an enormous diversity in physical characteristics and inhabit nearly every type of habitat within their range, from rainforests to deserts . Palms are among the best known and most extensively cultivated plant families. They have been important to humans throughout much of history, especially in regions like
2592-625: The tropics, mostly in humid lowland subtropical climates, in highlands in southern Asia, and along the rim lands of the Mediterranean Sea . The northernmost native palm is Chamaerops humilis , which reaches 44°N latitude along the coast of Liguria , Italy . In the southern hemisphere, the southernmost palm is the Rhopalostylis sapida , which reaches 44°S on the Chatham Islands where an oceanic climate prevails. Cultivation of palms
2646-542: The understanding of family relationships, sometimes leading to reclassification. The impact of these changes varies among different groups of organisms – while some families remain well-defined and easily recognizable, others require revision as new evidence emerges about evolutionary relationships. This balance between maintaining nomenclatural stability and incorporating new scientific discoveries remains an active area of taxonomic practice. Arecaceae The Arecaceae ( / ˌ ær ə ˈ k eɪ s i . iː , - ˌ aɪ / )
2700-508: The use of this term solely within the book's morphological section, where he delved into discussions regarding the vegetative and generative aspects of plants. Subsequently, in French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille was used as a French equivalent of the Latin ordo (or ordo naturalis ). The family concept in botany
2754-515: The water. Bonefishes possess crushing teeth in their palates. These genera are similar to Albula , except they can be found in deeper waters . [REDACTED] Family (biology) Family ( Latin : familia , pl. : familiae ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy . It is classified between order and genus . A family may be divided into subfamilies , which are intermediate ranks between
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#17331067004252808-426: The world. Along with dates mentioned above, members of the palm family with human uses are numerous: Like many other plants, palms have been threatened by human intervention and exploitation . The greatest risk to palms is destruction of habitat , especially in the tropical forests , due to urbanization , wood-chipping , mining , and conversion to farmland . Palms rarely reproduce after such great changes in
2862-412: Was further developed by the French botanists Antoine Laurent de Jussieu and Michel Adanson . Jussieu's 1789 Genera Plantarum divided plants into 100 'natural orders,' many of which correspond to modern plant families. However, the term 'family' did not become standardized in botanical usage until after the mid-nineteenth century. In zoology , the family as a rank intermediate between order and genus
2916-435: Was introduced by Pierre André Latreille in his Précis des caractères génériques des insectes, disposés dans un ordre naturel (1796). He used families (some of them were not named) in some but not in all his orders of "insects" (which then included all arthropods ). The standardization of zoological family names began in the early nineteenth century. A significant development came in 1813 when William Kirby introduced
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