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120-639: The region is administered by Waterton Lakes National Park and the Waterton Biosphere Reserve Association. The steep environmental gradients from the Continental Divide to the prairies have created an unusually rich mosaic of habitats with their associated flora and fauna . The biosphere region covers native prairie grasslands , aspen grove forests, subalpine forests , alpine tundra and meadows , cliffs, lakes and freshwater wetlands as well as tame pasture and cropland in

240-409: A cell wall . Newly dead animals may be covered by an exoskeleton . Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material. The chemical alteration of the dead organic matter

360-495: A food chain . Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey that is part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains which present

480-605: A habitat . Ecosystem ecology is the "study of the interactions between organisms and their environment as an integrated system". The size of ecosystems can range up to ten orders of magnitude , from the surface layers of rocks to the surface of the planet. The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire . It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as

600-449: A temperate climate and a varied view. According to Theodore Roosevelt : We have taken into our language the word prairie , because when our backwoodsmen first reached the land [in the Midwest ] and saw the great natural meadows of long grass—sights unknown to the gloomy forests wherein they had always dwelt—they knew not what to call them, and borrowed the term already in use among

720-609: A central role over a wide range, for example, in the slow development of soil from bare rock and the faster recovery of a community from disturbance . Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time that removes plant biomass". This can range from herbivore outbreaks, treefalls, fires, hurricanes, floods, glacial advances , to volcanic eruptions . Such disturbances can cause large changes in plant, animal and microbe populations, as well as soil organic matter content. Disturbance

840-407: A cliff (called a buffalo jump ), to kill or injure the bison en masse . The introduction of the horse and the gun greatly expanded the killing power of the plains Natives. That was followed by the policy of indiscriminate killing by European Americans and Canadians for both commercial reasons and to weaken the independence of plains Natives, and caused a dramatic drop in bison numbers from millions to

960-507: A critical role in global nutrient cycling and ecosystem function. Phosphorus enters ecosystems through weathering . As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for

1080-517: A faster recovery. More severe and more frequent disturbance result in longer recovery times. From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought , a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. Longer-term changes also shape ecosystem processes. For example,

1200-402: A few hundred in a century's time, and almost caused their extinction. The very dense soil plagued the first European settlers who were using wooden plows , which were more suitable for loose forest soil. On the prairie, the plows bounced around, and the soil stuck to them. This problem was solved in 1837 by an Illinois blacksmith named John Deere who developed a steel moldboard plow that

1320-454: A form that can be readily used by plants and microbes. Ecosystems provide a variety of goods and services upon which people depend, and may be part of. Ecosystem goods include the "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

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1440-427: A function-based typology has been proposed to leverage the strengths of these different approaches into a unified system. Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate. Ecosystems provide a variety of goods and services upon which people depend. Ecosystem goods include

1560-698: A general level, for example, tropical forests , temperate grasslands , and arctic tundra . There can be any degree of subcategories among ecosystem types that comprise a biome, e.g., needle-leafed boreal forests or wet tropical forests. Although ecosystems are most commonly categorized by their structure and geography, there are also other ways to categorize and classify ecosystems such as by their level of human impact (see anthropogenic biome ), or by their integration with social processes or technological processes or their novelty (e.g. novel ecosystem ). Each of these taxonomies of ecosystems tends to emphasize different structural or functional properties. None of these

1680-598: A large channel in which the Minnesota River evident today. The Red River of the North flows northward through a plain formerly covered by Lake Agassiz. Certain extraordinary features were produced when the retreat of the ice sheet had progressed so far as to open an eastward outlet for the marginal lakes. This outlet occurred along the depression between the northward slope of the Appalachian plateau in west-central New York and

1800-665: A means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem. Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades. Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Studies can be carried out at

1920-528: A more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients that become available. Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands ), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at

2040-406: A number of common, non random properties in the topology of their network. The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition,

2160-421: A process known as denitrification . Mycorrhizal fungi which are symbiotic with plant roots, use carbohydrates supplied by the plants and in return transfer phosphorus and nitrogen compounds back to the plant roots. This is an important pathway of organic nitrogen transfer from dead organic matter to plants. This mechanism may contribute to more than 70 Tg of annually assimilated plant nitrogen, thereby playing

2280-485: A rich growth of natural grass and annual flowering plants, but today, they are covered with farms. Ecosystem An ecosystem (or ecological system ) is a system that environments and their organisms form through their interaction. The biotic and abiotic components are linked together through nutrient cycles and energy flows. Ecosystems are controlled by external and internal factors . External factors such as climate , parent material which forms

2400-696: A significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter. Biodiversity plays an important role in ecosystem functioning. Ecosystem processes are driven by the species in an ecosystem, the nature of the individual species, and the relative abundance of organisms among these species. Ecosystem processes are the net effect of the actions of individual organisms as they interact with their environment. Ecological theory suggests that in order to coexist, species must have some level of limiting similarity —they must be different from one another in some fundamental way, otherwise, one species would competitively exclude

2520-408: A slower rate) even after soils become too dry to support plant growth. Ecosystems are dynamic entities. They are subject to periodic disturbances and are always in the process of recovering from past disturbances. When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance,

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2640-464: A small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. An ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys

2760-667: A substantial portion of global energy needs, and leave fertile land for food production." Unlike corn and soybeans, which are both directly and indirectly major food crops, including livestock feed, prairie grasses are not used for human consumption. Prairie grasses can be grown in infertile soil, eliminating the cost of adding nutrients to the soil. Tilman and his colleagues estimate that prairie grass biofuels would yield 51 percent more energy per acre than ethanol from corn grown on fertile land. Some plants commonly used are lupine, big bluestem (turkey foot), blazing star, switchgrass , and prairie clover . Because rich and thick topsoil made

2880-526: A system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation. Biomes are general classes or categories of ecosystems. However, there

3000-522: A variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. In such cases, microcosm experiments may fail to accurately predict ecosystem-level dynamics. Biomes are general classes or categories of ecosystems. However, there

3120-543: Is 1,037,070 hectares (4,004.1 sq mi). The core area is 49,936 hectares (192.80 sq mi), surrounded by buffer zone(s) of 336,476 hectares (1,299.14 sq mi) and transition area(s) of 445,213 hectares (1,718.98 sq mi). [REDACTED]  This article incorporates text from a free content work ( license statement/permission ). Text taken from UNESCO - MAB Biosphere Reserves Directory​ , UNESCO, UNESCO. Prairie Prairies are ecosystems considered part of

3240-574: Is a garden consisting primarily of plants from a prairie. The originally treeless prairies of the upper Mississippi basin began in Indiana and extended westward and north-westward until they merged with the drier region known as the Great Plains . An eastward extension of the same area, originally tree-covered, extended to central Ohio . Thus, the prairies generally lie between the Ohio and Missouri rivers on

3360-454: Is an international synthesis by over 1000 of the world's leading biological scientists that analyzes the state of the Earth's ecosystems and provides summaries and guidelines for decision-makers. The report identified four major categories of ecosystem services: provisioning, regulating, cultural and supporting services. It concludes that human activity is having a significant and escalating impact on

3480-491: Is below the soil surface and will re-grow from its deep (upwards of 20 feet ) roots . Without disturbance , trees will encroach on a grassland and cast shade, which suppresses the understory . Prairie and widely spaced oak trees evolved to coexist in the oak savanna ecosystem. Prairie ecosystems in the United States and Canada are divided into the easternmost tallgrass prairie , the westernmost shortgrass prairie , and

3600-680: Is constituted by most or all of the states, from north to south, of North Dakota , South Dakota , Nebraska , Kansas , and Oklahoma , and sizable parts of the states of Montana , Wyoming , Colorado , New Mexico , Texas in the west, and to the east, Minnesota , Wisconsin , Iowa , Missouri , Illinois , and Indiana . The Palouse of Washington and the Central Valley of California are also prairies. The Canadian Prairies occupy vast areas of Manitoba , Saskatchewan , and Alberta . Prairies may contain various lush flora and fauna , often contain rich soil maintained by biodiversity , with

3720-442: Is consumed by animals while still alive and enters the plant-based trophic system. After plants and animals die, the organic matter contained in them enters the detritus-based trophic system. Ecosystem respiration is the sum of respiration by all living organisms (plants, animals, and decomposers) in the ecosystem. Net ecosystem production is the difference between gross primary production (GPP) and ecosystem respiration. In

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3840-423: Is controlled by internal factors. Therefore, internal factors not only control ecosystem processes but are also controlled by them. Ecosystems are dynamic entities—they are subject to periodic disturbances and are always in the process of recovering from some past disturbance. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance . The capacity of

3960-460: Is estimated at over 6,000, and in southern Wisconsin, where it is placed at 5,000. They completely dominate the topography of their districts. A curious deposit of an impalpably fine and unstratified silt, known by the German name bess (or loess ), lies on the older drift sheets near the larger river courses of the upper Mississippi basin. It attains a thickness of 20 ft (6.1 m) or more near

4080-463: Is evidenced by the angularity of its grains (a bank of it will stand without slumping for years), whereas, if it had been transported significantly by water, the grains would have been rounded and polished. Loess is parent material for an extremely fertile, but droughty soil. Southwestern Wisconsin and parts of the adjacent states of Illinois, Iowa, and Minnesota are known as the driftless zone , because, although bordered by drift sheets and moraines, it

4200-569: Is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resource supply." The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession . A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and

4320-400: Is free from glacial deposits. It must therefore have been a sort of oasis, when the ice sheets from the north advanced past it on the east and west, and joined around its southern border. The reason for this exemption from glaciation is the converse of that for the southward convexity of the morainic loops. While they mark the paths of greatest glacial advance along lowland troughs (lake basins),

4440-554: Is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster the microbial decomposition occurs. Temperature also affects soil moisture, which affects decomposition. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play

4560-598: Is in cemetery prairies , railroad rights-of-way, or rocky/sandy/hilly places unsuitable for agriculture. States formerly with landcover in native tallgrass prairie including Iowa, Illinois, Minnesota, Wisconsin, Nebraska, and Missouri have become valued for their highly productive soils and are included in the Corn Belt . As an example of this land use intensity, Illinois and Iowa rank 49th and 50th, out of 50 US states, in total uncultivated land remaining. Drier shortgrass prairies were once used mostly for open-range ranching. With

4680-405: Is no clear distinction between biomes and ecosystems. Biomes are always defined at a very general level. Ecosystems can be described at levels that range from very general (in which case the names are sometimes the same as those of biomes) to very specific, such as "wet coastal needle-leafed forests". Biomes vary due to global variations in climate . Biomes are often defined by their structure: at

4800-516: Is no clear distinction between biomes and ecosystems. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Biotic factors of the ecosystem are living things; such as plants, animals, and bacteria, while abiotic are non-living components; such as water, soil and atmosphere. Plants allow energy to enter

4920-416: Is not known if that is because of inherent differences in the species or because farmed cattle tend to be confined in smaller areas. Bison dung is a vital source of nutrients for prairie soil, spreads seeds, and supports over 1,000 insect species, including specialist dung beetles which cannot subsist on the feces of any other animal. In spite of long recurrent droughts and occasional torrential rains ,

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5040-430: Is now buried under the drift. The most significant area of the prairies, from Indiana to North Dakota , consists of till plains, that is, sheets of unstratified drift. The plains are 30, 50 or even 100 ft (up to 30 m) thick covering the underlying rock surface for thousands of square miles except where postglacial stream erosion has locally laid it bare. The plains have an extraordinarily even surface. The till

5160-519: Is presumably made in part of preglacial soils, but it is largely composed of rock waste mechanically transported by the creeping ice sheets. Although the crystalline rocks from Canada and some of the more resistant stratified rocks south of the Great Lakes occur as boulders and stones, a great part of the till has been crushed and ground to a clayey texture. The till plains, although sweeping in broad swells of slowly changing altitude, often appear level to

5280-531: Is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes that can break through the tough outer structures surrounding dead plant material. They also produce enzymes that break down lignin , which allows them access to both cell contents and the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources. Decomposition rates vary among ecosystems. The rate of decomposition

5400-400: Is similar to the Appalachian piedmont which though not exhausted by the primeval forest cover, are by no means so rich as the till sheets of the prairies. Moreover, whatever the rocky understructure, the till soil has been averaged by a thorough mechanical mixture of rock grindings. Hence, the prairies are continuously fertile for scores of miles together. The true prairies were once covered with

5520-485: Is termed its resistance . The capacity of a system to absorb disturbance and reorganize while undergoing change so as to retain essentially the same function, structure, identity, and feedbacks is termed its ecological resilience . Resilience thinking also includes humanity as an integral part of the biosphere where we are dependent on ecosystem services for our survival and must build and maintain their natural capacities to withstand shocks and disturbances. Time plays

5640-422: Is the "best" classification. Ecosystem classifications are specific kinds of ecological classifications that consider all four elements of the definition of ecosystems : a biotic component, an abiotic complex, the interactions between and within them, and the physical space they occupy. Different approaches to ecological classifications have been developed in terrestrial, freshwater and marine disciplines, and

5760-814: Is the dominant habitat type in the Southern Canadian agricultural and climatic region which is known as Palliser's Triangle . It was once thought to be completely unarable, but is now one of the most important agricultural regions in Canada thanks to advances in irrigation technology. The dominant plant life in prairies consists of grasses , which may include 40 to 60 different grass species. In addition to grasses, prairies can include over 300 species of flowering plants. The Konza Tallgrass Prairie in Kansas hosts 250 species of native plants and provides habitat for 208 birds, 27 mammals, 25 reptiles, and over 3,000 insects. Some of

5880-599: The Great Plains ecoregion. The Dust Bowl was a major reason for the Great Depression . Nomadic hunting has been the main human activity on the prairies for the majority of the archaeological record. This once included many now-extinct species of megafauna . After the other extinctions, the main hunted animal on the prairies was the plains bison . Using loud noises and waving large signals, Native peoples would drive bison into fenced pens called buffalo pounds to be killed with bows and arrows or spears, or drive them off

6000-555: The resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors. Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis . The energy incorporated through this process supports life on earth, while

6120-578: The temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type . Temperate grassland regions include the Pampas of Argentina , Brazil and Uruguay , and the steppe of Ukraine , Russia , and Kazakhstan . Lands typically referred to as "prairie" tend to be in North America . The term encompasses

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6240-435: The "tangible, material products" of ecosystem processes such as water, food, fuel, construction material, and medicinal plants . They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species. Ecosystem services , on the other hand, are generally "improvements in the condition or location of things of value". These include things like

6360-709: The French inhabitants. Prairie ( pronounced [pʁɛʁi] ) is the French word for "meadow", formed ultimately from the Latin root word pratum (which has the same meaning). The formation of the North American Prairies started with the uplift of the Rocky Mountains near Alberta . The mountains created a rain shadow which resulted in lower precipitation rates downwind. The parent material of most prairie soil

6480-481: The absence of disturbance, net ecosystem production is equivalent to the net carbon accumulation in the ecosystem. Energy can also be released from an ecosystem through disturbances such as wildfire or transferred to other ecosystems (e.g., from a forest to a stream to a lake) by erosion . In aquatic systems , the proportion of plant biomass that gets consumed by herbivores is much higher than in terrestrial systems. In trophic systems, photosynthetic organisms are

6600-402: The amount of energy available to the ecosystem. Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate , soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on

6720-504: The amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), the rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis. Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on

6840-462: The area is still very prone to extended periods of drought, which can be disastrous for the industry if it is significantly prolonged. Research by David Tilman , ecologist at the University of Minnesota , suggests " Biofuels made from high-diversity mixtures of prairie plants can reduce global warming by removing carbon dioxide from the atmosphere. Even when grown on infertile soils, they can provide

6960-408: The carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels . It also drives the carbon cycle , which influences global climate via the greenhouse effect . Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen . The photosynthesis carried out by all

7080-462: The central mixed-grass prairie. Tallgrass prairies receive over 30 inches of rainfall per year, whereas shortgrass prairies are much more arid, receiving only 12 inches or so, and mixed-grass prairies receive intermediate rainfall. Wet, mesic, and dry prairie ecosystems can also form more locally due to soil and terrain characteristics. Wet prairies may form in low-lying areas with poor drainage; dry prairie can be found on uplands or slopes. Dry prairie

7200-432: The combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems. When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in

7320-418: The concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined

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7440-403: The dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted. Decomposition processes can be separated into three categories— leaching , fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in

7560-541: The development of barbed wire in the 1870s and improved irrigation techniques, this region has mostly been converted to cropland and small fenced pastures. In southern Canada, Palliser's Triangle has been changed into one of the most important sources of wheat in the world as a result of improved methods of watering wheat fields (along with the rest of the Southern prairie provinces which also grow wheat, canola and many other grains). Despite those advances in farming technology,

7680-535: The disturbances of grazing and fire. Native ungulates such as bison , elk , and white-tailed deer roamed the expansive, diverse grasslands before European colonization of the Americas . For 10,000-20,000 years, native people used fire annually as a tool to assist in hunting, transportation, and safety. Evidence of ignition sources of fire in the tall grass prairie are overwhelmingly human as opposed to lightning. Humans, and grazing animals, were active participants in

7800-472: The diverse grasses and herbaceous plants. Prairies also depend on the presence of large grazing animals, particularly bison . Bison are important to the prairie ecosystem because they shape and alter the environment by grazing, trampling areas with their hooves, wallowing, and depositing manure. Bison eat more grass than flowering plants, increasing the diversity of plants in the prairie. Cattle are thought to prefer to eat flowering plants over grasses, but it

7920-431: The dominant grasses of prairies are Indian grass , big bluestem , side-oats grama , Canada wildrye , and switchgrass . Prairies are considered to be fire-dependent ecosystems . Regular controlled burning by Native Americans, practices developed through observation of non-anthropogenic fire and its effects, maintained the biodiversity of the prairie, clearing away dead vegetation and preventing trees from shading out

8040-563: The drift on the under rock. Further south, as far as the entrance of the Ohio River, the Mississippi follows a rock-walled valley 300 to 400 ft (91 to 122 m) deep, with a flood-plain 2 to 4 mi (3.2 to 6.4 km) wide. This valley seems to represent the path of an enlarged early-glacial Mississippi, when much precipitation that is today discharged to Hudson Bay and the Gulf of St. Lawrence

8160-452: The driftless zone is a district protected from ice invasion by reason of the obstruction which the highlands of northern Wisconsin and Michigan (part of the Superior upland) offered to glacial advance. The course of the upper Mississippi River is largely consequent upon glacial deposits. Its sources are in the morainic lakes in northern Minnesota. The drift deposits thereabouts are so heavy that

8280-508: The eastern tallgrass prairies were plowed and turned into agricultural lands, the prairie grasses with their strong root systems were destroyed. In combination with severe droughts that resulted in the Dust Bowl , a major ecological disaster in which winds picked up the dry, unprotected prairie soil and formed it into "black blizzards" of airborne dirt that blackened the skies for days at a time across 19 states and forced 400,000 people to abandon

8400-413: The ecologically diverse core area. The region is also home to many vibrant communities – Pincher Creek , Cardston , Crowsnest Pass , Piikani and Kainai Reserves, and others – each with a rich and distinct cultural history, and reliance on a wide range of economic activities. In Waterton Biosphere Region, people are actively involved with ranching, farming, tourism and recreation, wind energy, mining,

8520-643: The ecosystem or to gradual disruption of biotic processes and degradation of abiotic conditions of the ecosystem. Once the original ecosystem has lost its defining features, it is considered "collapsed ". Ecosystem restoration can contribute to achieving the Sustainable Development Goals . An ecosystem (or ecological system) consists of all the organisms and the abiotic pools (or physical environment) with which they interact. The biotic and abiotic components are linked together through nutrient cycles and energy flows. "Ecosystem processes" are

8640-670: The escarpment to the lower, Lake Ontario . That gave rise to Niagara Falls . Lake Ontario's outlet for a time ran down the Mohawk Valley to the Hudson River. At the higher elevation, it was known as Lake Iroquois . When ice melted from the northeastern end of the lake, it dropped to a lower level, and drained through the St. Lawrence area creating a lower base level for the Niagara River and increasing its erosive capacity. In certain districts,

8760-435: The eye with a view stretching to the horizon. Here and there, faint depressions occur, occupied by marshy sloughs or floored with a rich black soil of postglacial origin. Thus, by sub-glacial aggradation, the prairies have been leveled up to a smooth surface, in contrast to the higher and non-glaciated hilly country just to the south. The great ice sheets formed terminal moraines around their border at various stages. However,

8880-474: The father of waters, like so many other rivers in the Northern states, owes many of its features more or less directly to glacial action. The fertility of the prairies is a natural consequence of their origin. During the mechanical transportation of the till, no vegetation was present to remove the minerals essential to plant growth, as is the case in the soils of normally weathered and dissected peneplains. The soil

9000-421: The flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum , further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems. Ecosystems are controlled by both external and internal factors. External factors, also called state factors, control

9120-452: The forests of eastern North America still show legacies of cultivation which ceased in 1850 when large areas were reverted to forests. Another example is the methane production in eastern Siberian lakes that is controlled by organic matter which accumulated during the Pleistocene . Ecosystems continually exchange energy and carbon with the wider environment . Mineral nutrients, on

9240-461: The grasslands of the Great Plains were not subject to great soil erosion . The root systems of native prairie grasses firmly held the soil in place to prevent run-off of soil. When the plant died, the fungi and bacteria returned its nutrients to the soil. These deep roots also helped native prairie plants reach water in even the driest conditions. Native grasses suffer much less damage from dry conditions than many farm crops currently grown. When

9360-417: The head of the gorges are now occupied by little lakes. The most significant stage in this series of changes occurred when the glacio-marginal lake waters were lowered so that the long escarpment of Niagara limestone was laid bare in western New York. The previously confluent waters were then divided into two lakes. The higher one, Lake Erie , supplied the outflowing Niagara River , which poured its waters down

9480-416: The ice of the last glacial epoch had retreated so far that its front border lay on a northward slope, belonging to the drainage area of the Great Lakes, bodies of water accumulated in front of the ice margin, forming glacio-marginal lakes. The lakes were small at first, and each had its own outlet at the lowest depression of land to the south. As the ice melted further back, neighboring lakes became confluent at

9600-483: The ice sheets advanced in lobes along the lowlands of the Great Lakes . Neighboring morainic loops join each other in re-entrants (north-pointing cusps), where two adjacent glacial lobes came together and formed their moraines in largest volume. The moraines are of too small relief to be shown on any maps except of the largest scale. Small as they are, they are the chief relief of the prairie states, and, in association with

9720-471: The ice sheets extended to the land sloping southward to the Ohio River , Mississippi River and Missouri River , the drift-laden streams flowed freely away from the ice border. As the streams escaped from their subglacial channels, they spread into broader channels and deposited some of their load, and thus aggraded their courses. Local sheets or aprons of gravel and sand are spread more or less abundantly along

9840-516: The land well suited for agricultural use, only 1% of tallgrass prairie remains in the U.S. today. Shortgrass prairie is more abundant. Significant preserved areas of prairie include: Virgin prairie refers to prairie land which has never been plowed. Small virgin prairies exist in the American Midwestern states and in Canada. Restored prairie refers to a prairie that has been reseeded after plowing or other disturbance. A prairie garden

9960-691: The landscape, versus one present on an adjacent steep hillside. Other external factors that play an important role in ecosystem functioning include time and potential biota , the organisms that are present in a region and could potentially occupy a particular site. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function. Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. While

10080-446: The level of the lowest outlet of the group. The outflowing streams grew in the same proportion and eroded a broad channel across the height of land and far down stream, while the lake waters built sand reefs or carved shore cliffs along their margin, and laid down sheets of clay on their floors. All of these features are easily recognized in the prairie region. The present site of Chicago was determined by an Indian portage or carry across

10200-423: The living and dead plant matter, and eventually released through respiration. The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus . In terrestrial ecosystems , the vast majority of the net primary production ends up being broken down by decomposers . The remainder

10320-500: The low divide between Lake Michigan and the headwaters of the Illinois River . This divide lies on the floor of the former outlet channel of the glacial Lake Michigan. Corresponding outlets are known for Lake Erie , Lake Huron , and Lake Superior . A very large sheet of water, named Lake Agassiz , once overspread a broad till plain in northern Minnesota and North Dakota. The outlet of this glacial lake, called river Warren, eroded

10440-513: The lower and mid-latitude of the area referred to as the Interior Plains of Canada , the United States , and Mexico . It includes all of the Great Plains as well as the wetter, hillier land to the east. From west to east, generally the drier expanse of shortgrass prairie gives way to mixed grass prairie and ultimately the richer soils of the tallgrass prairie . In the U.S., the area

10560-471: The maintenance of hydrological cycles , cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. Many ecosystems become degraded through human impacts, such as soil loss , air and water pollution , habitat fragmentation , water diversion , fire suppression , and introduced species and invasive species . These threats can lead to abrupt transformation of

10680-408: The maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted. The Millennium Ecosystem Assessment

10800-419: The morainic belts are of slight relief in comparison to the great area of the ice. They rise gently from the till plains to 50, 100 or more feet. They may be one, two or three miles (5 km) wide and their hilly surface, dotted over with boulders, contains many small lakes in basins or hollows, instead of streams in valleys. The morainic belts are arranged in groups of concentric loops, convex southward, because

10920-406: The nearly imperceptible slopes of the till plains, they determine the course of many streams and rivers, which as a whole are consequent upon the surface form of the glacial deposits. The complexity of the glacial period and its subdivision into several glacial epochs, separated by interglacial epochs of considerable length (certainly longer than the postglacial epoch) has a structural consequence in

11040-419: The oil and gas industry, and other natural resource-related activities. The Waterton Biosphere Reserve Association strives to achieve a balance between conserving biological diversity, promoting sustainable use of resources, and building the capacity of local residents, First Nations, and organizations to positively affect their communities and the environment. The region's surface area (terrestrial and marine)

11160-537: The other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation , is deposited through precipitation, dust, gases or is applied as fertilizer . Most terrestrial ecosystems are nitrogen-limited in the short term making nitrogen cycling an important control on ecosystem production. Over the long term, phosphorus availability can also be critical. Macronutrients which are required by all plants in large quantities include

11280-509: The other. Despite this, the cumulative effect of additional species in an ecosystem is not linear: additional species may enhance nitrogen retention, for example. However, beyond some level of species richness, additional species may have little additive effect unless they differ substantially from species already present. This is the case for example for exotic species . The addition (or loss) of species that are ecologically similar to those already present in an ecosystem tends to only have

11400-417: The outer side of the morainic belts. Long trains of gravel and sands clog the valleys that lead southward from the glaciated to the non-glaciated area. Later, when the ice retreated further and the unloaded streams returned to their earlier degrading habit, they more or less completely scoured out the valley deposits, the remains of which are now seen in terraces on either side of the present flood plains. When

11520-403: The overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. On broad geographic scales, climate is the factor that "most strongly determines ecosystem processes and structure". Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine

11640-405: The plants in an ecosystem is called the gross primary production (GPP). About half of the gross GPP is respired by plants in order to provide the energy that supports their growth and maintenance. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include

11760-466: The prairies. Prairie grasslands including Danthonia spp., Festuca scabrella and prairie Junegrass ; aspen grove forests with quaking aspen ( Populus tremuloides ), Amelanchier alnifolia and cow parsnip { Heracleum latanum ); alpine tundra/high meadows characterized by Douglas fir ( Pseudotsuga menziesii ), lodgepole pine ( Pinus contorta ), limber pine ( P. flexilis ) and white barkpine ( P. albicaulis ); arctic-alpine communities above

11880-486: The present divides between the drainage basins of Hudson Bay , Lake Superior , and the Gulf of Mexico evidently stand in no very definite relation to the preglacial divides. The course of the Mississippi through Minnesota is largely guided by the form of the drift cover. Several rapids and the Saint Anthony Falls (determining the site of Minneapolis ) are signs of immaturity, resulting from superposition through

12000-585: The primary nutrients (which are most limiting as they are used in largest amounts): Nitrogen, phosphorus, potassium. Secondary major nutrients (less often limiting) include: Calcium, magnesium, sulfur. Micronutrients required by all plants in small quantities include boron, chloride, copper, iron, manganese, molybdenum, zinc. Finally, there are also beneficial nutrients which may be required by certain plants or by plants under specific environmental conditions: aluminum, cobalt, iodine, nickel, selenium, silicon, sodium, vanadium. Until modern times, nitrogen fixation

12120-413: The primary producers. The organisms that consume their tissues are called primary consumers or secondary producers — herbivores . Organisms which feed on microbes ( bacteria and fungi ) are termed microbivores . Animals that feed on primary consumers— carnivores —are secondary consumers. Each of these constitutes a trophic level. The sequence of consumption—from plant to herbivore, to carnivore—forms

12240-410: The process of prairie formation and the establishment of the diversity of graminoid and forbs species. Fire has the effect on prairies of removing trees , clearing dead plant matter, and changing the availability of certain nutrients in the soil from the ash produced. Fire kills the vascular tissue of trees, but not prairie species, as up to 75% (depending on the species) of the total plant biomass

12360-460: The rivers and gradually fades away at a distance of ten or more miles (16 or more km) on either side. It contains land shells, and hence cannot be attributed to marine or lacustrine submergence. The best explanation is that, during certain phases of the glacial period, it was carried as dust by the winds from the flood plains of aggrading rivers, and slowly deposited on the neighboring grass-covered plains. The glacial and eolian origin of this sediment

12480-409: The soil and topography , control the overall structure of an ecosystem but are not themselves influenced by the ecosystem. Internal factors are controlled, for example, by decomposition , root competition, shading, disturbance, succession, and the types of species present. While the resource inputs are generally controlled by external processes, the availability of these resources within the ecosystem

12600-611: The soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars , amino acids and mineral nutrients. Leaching is more important in wet environments and less important in dry ones. Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark , and cell contents are protected by

12720-511: The soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization . Others convert ammonium to nitrite and nitrate ions, a process known as nitrification . Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas ,

12840-523: The south and the Great Lakes on the north. The prairies are a contribution of the glacial period. They consist of glacial drift deposited unconformably on an underlying rock surface of moderate or small relief. Here, the rocks are an extension of the same stratified Palaeozoic formations already described as occurring in the Appalachian region and around the Great Lakes . They are usually fine-textured limestones and shales lying horizontal. The moderate or small relief they were given by mature preglacial erosion

12960-505: The southward slope of the melting ice sheet. When this eastward outlet came to be lower than the south-westward outlet across the height of land to the Ohio or Mississippi river, the discharge of the marginal lakes was changed from the Mississippi system to the Hudson system. Many well-defined channels, cutting across the north-sloping spurs of the plateau in the neighborhood of Syracuse, New York mark

13080-483: The spatial extent of ecosystems using the term " ecotope ". G. Evelyn Hutchinson , a limnologist who was a contemporary of Tansley's, combined Charles Elton 's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky . As a result, he suggested that mineral nutrient availability in a lake limited algal production . This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that

13200-509: The subglacial till was not spread out in a smooth plain, but accumulated in elliptical mounds, 100–200 feet. high and 0.5 to 1 mile (0.80 to 1.61 kilometres) long with axes parallel to the direction of the ice motion as indicated by striae on the underlying rock floor. These hills are known by the Irish name, drumlins , used for similar hills in north-western Ireland . The most remarkable groups of drumlins occur in western New York , where their number

13320-425: The superposition of successive till sheets, alternating with non-glacial deposits. It also has a physiographic consequence in the very different amount of normal postglacial erosion suffered by the different parts of the glacial deposits. The southernmost drift sheets, as in southern Iowa and northern Missouri , have lost their initially plain surface and are now maturely dissected into gracefully rolling forms. Here,

13440-454: The system through photosynthesis , building up plant tissue. Animals play an important role in the movement of matter and energy through the system, by feeding on plants and on one another. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter , decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to

13560-413: The temporary paths of the ice-bordered outlet river. Successive channels are found at lower and lower levels on the plateau slope, indicating the successive courses taken by the lake outlet as the ice melted further and further back. On some of the channels, deep gorges were eroded heading in temporary cataracts which exceeded Niagara in height but not in breadth. The pools excavated by the plunging waters at

13680-421: The transfers of energy and materials from one pool to another. Ecosystem processes are known to "take place at a wide range of scales". Therefore, the correct scale of study depends on the question asked. The term "ecosystem" was first used in 1935 in a publication by British ecologist Arthur Tansley . The term was coined by Arthur Roy Clapham , who came up with the word at Tansley's request. Tansley devised

13800-939: The tree line dominated by Dryas octopetala and Polemonium viscosum ; upper subalpine forests with alpine larch ( Larix lyallii ), Luzula hitchcockii , Engelmann spruce ( Picea engelmannii ) and subalpine fir ( Abies lasiocarpa ); lower subalpine forests dominated by Engelmann spruce and subalpine fir; deciduous forest , coniferous forests dominated by Douglas fir and limber pine; cliffs, lakes and freshwater wetlands; disturbed, heavily grazed land with quaking aspen, Urtica dioica , Bromus inermis and Phleum pratense . Several rare or threatened plant and animal species are found in Waterton Biosphere Region with more than 1000 vascular plant species, 182 bryophytes and 218 lichen species, 60 species of mammals , over 260 species of birds , 24 species of fish , and 10 reptiles and amphibians being recorded within

13920-456: The valleys of even the small streams are well opened and graded, and marshes and lakes are rare. These sheets are of early Pleistocene origin. Nearer the Great Lakes, the till sheets are trenched only by the narrow valleys of the large streams. Marshy sloughs still occupy the faint depressions in the till plains and the associated moraines have abundant small lakes in their undrained hollows. These drift sheets are of late Pleistocene origin. When

14040-521: Was delivered to the Gulf of Mexico, for the curves of the present river are of distinctly smaller radii than the curves of the valley. Lake Pepin (30 mi [48 km] below St. Paul ), a picturesque expansion of the river across its flood-plain, is due to the aggradation of the valley floor where the Chippewa River , coming from the northeast, brought an overload of fluvio-glacial drift. Hence, even

14160-450: Was distributed during the last glacial advance that began about 110,000 years ago. The glaciers expanding southward scraped the landscape, picking up geologic material and leveling the terrain. As the glaciers retreated about 10,000 years ago, they deposited this material in the form of till . Wind-based loess deposits also form an important parent material for prairie soils. Tallgrass prairie evolved over tens of thousands of years with

14280-409: Was stronger and cut the roots, making the fertile soils ready for farming. Former grasslands are now among the most productive agricultural lands on Earth. The tallgrass prairie has been converted into one of the most intensive crop producing areas in North America. Less than one tenth of one percent (<0.09%) of the original landcover of the tallgrass prairie biome remains. Much of what persists

14400-679: Was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants that support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs , which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through

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