Misplaced Pages

#774225

85-453: A walnut is the edible seed of any tree of the genus Juglans (family Juglandaceae ), particularly the Persian or English walnut, Juglans regia . They are accessory fruit because the outer covering of the fruit is technically an involucre and thus not morphologically part of the carpel; this means it cannot be a drupe but is instead a drupe-like nut. After full ripening, the shell

170-489: A coleoptile that forms the first leaf while the radicle is covered with a coleorhiza that connects to the primary root and adventitious roots form the sides. Here the hypocotyl is a rudimentary axis between radicle and plumule. The seeds of corn are constructed with these structures; pericarp, scutellum (single large cotyledon) that absorbs nutrients from the endosperm, plumule, radicle, coleoptile, and coleorhiza – these last two structures are sheath-like and enclose

255-538: A Germanic word in general use in the North Sea area since the Middle Ages (cf. Old English and Old Frisian tunne , Old High German and Medieval Latin tunna , German and French tonne ) to designate a large cask, or tun . A full tun, standing about a metre high, could easily weigh a tonne. See also the common German word de:Mülltonne (literal translation: garbage drum ). The spelling tonne pre-dates

340-500: A garnish on some foods. Nocino is a liqueur made from unripe green walnuts steeped in alcohol with syrup added. Walnut oil is available commercially and is chiefly used as a food ingredient, particularly in salad dressings . It has a low smoke point , which limits its use for frying . Walnut husks can be used to make durable ink for writing and drawing. It is thought to have been used by artists including Leonardo da Vinci and Rembrandt . Walnut husk pigments are used as

425-508: A zygote . The embryo within a seed develops from the zygote and grows within the mother plant to a certain size before growth is halted. The formation of the seed is the defining part of the process of reproduction in seed plants ( spermatophytes ). Other plants such as ferns , mosses and liverworts , do not have seeds and use water-dependent means to propagate themselves. Seed plants now dominate biological niches on land, from forests to grasslands both in hot and cold climates . In

510-480: A brown dye for fabric and were used in classical Rome and medieval Europe for dyeing hair . The US Army once used ground walnut shells for abrasive blasting to clean aviation parts because of low cost and low abrasive qualities. However, an investigation of a fatal Boeing CH-47 Chinook helicopter crash (11 September 1982, in Mannheim, Germany) revealed that walnut shell grit had clogged an oil port, leading to

595-422: A curved megagametophyte often giving the seed a tight "C" shape. The last ovule shape is called amphitropous , where the ovule is partly inverted and turned back 90 degrees on its stalk (the funicle or funiculus ). In the majority of flowering plants, the zygote's first division is transversely oriented in regards to the long axis, and this establishes the polarity of the embryo. The upper or chalazal pole becomes

680-411: A few will end in a favorable place for growth. Herbaceous perennials and woody plants often have larger seeds; they can produce seeds over many years, and larger seeds have more energy reserves for germination and seedling growth and produce larger, more established seedlings after germination. Seeds serve several functions for the plants that produce them. Key among these functions are nourishment of

765-422: A form of gambling known as du qing pi . Seed In botany , a seed is a plant embryo and nutrient reserve enclosed in a seed coat, a protective outer covering called a testa. More generally, the term "seed" means anything that can be sown , which may include seed and husk or tuber . Seeds are the product of the ripened ovule , after the embryo sac is fertilized by sperm from pollen , forming

850-483: A health claim that "Walnuts contribute to the improvement of endothelium -dependent vasodilation " at a daily intake of 30 grams (1.1 oz); it also found that a cause and effect relationship did not exist between consuming walnuts and reduction of blood LDL-cholesterol levels. The recommended health claim was later authorized by the European Commission . A 2020 systematic review assessing

935-415: A location and be there at a time favorable for germination and growth. When the fruits open and release their seeds in a regular way, it is called dehiscent , which is often distinctive for related groups of plants; these fruits include capsules , follicles , legumes , silicles and siliques . When fruits do not open and release their seeds in a regular fashion, they are called indehiscent, which include

SECTION 10

#1732844119775

1020-754: A mix such as muesli , or as an ingredient of a dish : e.g. walnut soup , walnut pie , walnut coffee cake , banana cake , brownie , fudge . Walnuts are often candied or pickled. Pickled walnuts that are the whole fruit can be savory or sweet depending on the preserving solution. Walnuts may be used as an ingredient in other foodstuffs. Walnut is an important ingredient in baklava , Circassian chicken , potica (a traditional festive pastry from Slovenia ), satsivi (chicken in walnut sauce), tarator (a summer soup in Bulgarian cuisine ), and poultry or meat ball stew from Iranian cuisine . Walnuts are also popular as an ice cream topping, and walnut pieces are used as

1105-435: A supply of nutrients for the embryo in most monocotyledons and the endospermic dicotyledons. Seeds have been considered to occur in many structurally different types (Martin 1946). These are based on a number of criteria, of which the dominant one is the embryo-to-seed size ratio. This reflects the degree to which the developing cotyledons absorb the nutrients of the endosperm, and thus obliterate it. Six types occur amongst

1190-550: A tegmen from the inner integument while unitegmic seeds have only one integument. Usually, parts of the testa or tegmen form a hard protective mechanical layer. The mechanical layer may prevent water penetration and germination. Amongst the barriers may be the presence of lignified sclereids . The outer integument has a number of layers, generally between four and eight organised into three layers: (a) outer epidermis, (b) outer pigmented zone of two to five layers containing tannin and starch, and (c) inner epidermis. The endotegmen

1275-415: A waste product, superior natural biodegradability, and odor control as good or better than clay litter. Disadvantages include the possibility of allergic reactions among humans and cats. Walnuts have been listed as one of the 38 substances used to prepare Bach flower remedies , a herbal remedy promoted in folk medicine practices for its supposed effect on health. According to Cancer Research UK , "there

1360-405: A way for the species to survive dry or cold seasons. Ephemeral plants are usually annuals that can go from seed to seed in as few as six weeks. Seed germination is a process by which a seed embryo develops into a seedling. It involves the reactivation of the metabolic pathways that lead to growth and the emergence of the radicle or seed root and plumule or shoot. The emergence of the seedling above

1445-509: Is at a suitable temperature with proper soil moisture. This true dormancy or innate dormancy is therefore caused by conditions within the seed that prevent germination. Thus dormancy is a state of the seed, not of the environment. Induced dormancy, enforced dormancy or seed quiescence occurs when a seed fails to germinate because the external environmental conditions are inappropriate for germination, mostly in response to conditions being too dark or light, too cold or hot, or too dry. Seed dormancy

1530-481: Is caused by conditions outside the embryo, including: Endogenous dormancy is caused by conditions within the embryo itself, including: The following types of seed dormancy do not involve seed dormancy, strictly speaking, as lack of germination is prevented by the environment, not by characteristics of the seed itself (see Germination ): Not all seeds undergo a period of dormancy. Seeds of some mangroves are viviparous; they begin to germinate while still attached to

1615-402: Is derived from the inner epidermis of the inner integument, the exotegmen from the outer surface of the inner integument. The endotesta is derived from the inner epidermis of the outer integument, and the outer layer of the testa from the outer surface of the outer integument is referred to as the exotesta . If the exotesta is also the mechanical layer, this is called an exotestal seed, but if

1700-478: Is discarded, and the kernel is eaten. Nuts of the eastern black walnut ( Juglans nigra ) and butternuts ( Juglans cinerea ) are less commonly consumed. Walnuts are the round, single-seed stone fruits of the walnut tree. They ripen between September and November in the northern hemisphere. The brown, wrinkly walnut shell is enclosed in a husk . Shells of walnuts available in commerce usually have two segments (but three or four-segment shells can also form). During

1785-553: Is no scientific evidence to prove that flower remedies can control, cure or prevent any type of disease, including cancer". Large, symmetrically shaped, and sometimes intricately carved walnut shells (mainly from J. hopeiensis ) are valued collectibles in China where they are rotated in hand as a plaything or as decoration. They are also an investment and status symbol , with some carvings having high monetary value if unique. Pairs of walnuts are sometimes sold in their green husks for

SECTION 20

#1732844119775

1870-444: Is not the same as seed persistence in the soil or on the plant, though even in scientific publications dormancy and persistence are often confused or used as synonyms. Often, seed dormancy is divided into four major categories: exogenous; endogenous; combinational; and secondary. A more recent system distinguishes five classes: morphological, physiological, morphophysiological, physical, and combinational dormancy. Exogenous dormancy

1955-535: Is protection against disease. Seeds protect and nourish the embryo or young plant. They usually give a seedling a faster start than a sporeling from a spore, because of the larger food reserves in the seed and the multicellularity of the enclosed embryo. Unlike animals, plants are limited in their ability to seek out favorable conditions for life and growth. As a result, plants have evolved many ways to disperse their offspring by dispersing their seeds (see also vegetative reproduction ). A seed must somehow "arrive" at

2040-400: Is the joule . One tonne of TNT is approximately equivalent to 4.2 gigajoules. In the petroleum industry the tonne of oil equivalent (toe) is a unit of energy : the amount of energy released by burning one tonne of crude oil , approximately 42 GJ. There are several slightly different definitions. This is ten times as much as a tonne of TNT because atmospheric oxygen is used. Like

2125-446: Is the ability of the embryo to germinate and is affected by a number of different conditions. Some plants do not produce seeds that have functional complete embryos, or the seed may have no embryo at all, often called empty seeds. Predators and pathogens can damage or kill the seed while it is still in the fruit or after it is dispersed. Environmental conditions like flooding or heat can kill the seed before or during germination. The age of

2210-411: Is then aborted or absorbed during early development. The seed is composed of the embryo (the result of fertilization) and tissue from the mother plant, which also form a cone around the seed in coniferous plants such as pine and spruce . Seeds are very diverse, and as such there are many terms are used to describe them. A typical seed includes two basic parts: In addition, the endosperm forms

2295-419: Is usually triploid , and is rich in oil or starch , and protein . In gymnosperms, such as conifers , the food storage tissue (also called endosperm) is part of the female gametophyte , a haploid tissue. The endosperm is surrounded by the aleurone layer (peripheral endosperm), filled with proteinaceous aleurone grains. Originally, by analogy with the animal ovum , the outer nucellus layer ( perisperm )

2380-570: The Byzantine era , the walnut was also known by the name "royal nut". An article on walnut tree cultivation in Spain is included in Ibn al-'Awwam 's 12th-century Book on Agriculture . The wal element in the name is Germanic and means foreign , especially in the sense of Latin or non-Germanic . Compare, for example, Wales , Walloons , Wallachia . The wal element is present in other Germanic-language words for

2465-510: The Daily Value , DV) of several dietary minerals , particularly manganese at 148% DV, along with significant amounts of B vitamins (table). Unlike most nuts, which are high in monounsaturated fatty acids , walnut oil is composed largely of polyunsaturated fatty acids (72% of total fats), particularly alpha-linolenic acid (14%) and linoleic acid (58%), although it does contain oleic acid as 13% of total fats (table source). In 2004,

2550-417: The cone scales as they develop in some species of conifer . Angiosperm (flowering plants) seeds consist of three genetically distinct constituents: (1) the embryo formed from the zygote, (2) the endosperm, which is normally triploid, (3) the seed coat from tissue derived from the maternal tissue of the ovule. In angiosperms, the process of seed development begins with double fertilization , which involves

2635-412: The embryo , dispersal to a new location, and dormancy during unfavorable conditions. Seeds fundamentally are means of reproduction, and most seeds are the product of sexual reproduction which produces a remixing of genetic material and phenotype variability on which natural selection acts. Plant seeds hold endophytic microorganisms that can perform various functions, the most important of which

Walnut - Misplaced Pages Continue

2720-421: The flowering plants , the ovary ripens into a fruit which contains the seed and serves to disseminate it. Many structures commonly referred to as "seeds" are actually dry fruits. Sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the so-called stone fruits (such as

2805-424: The legumes (such as beans and peas ), trees such as the oak and walnut , vegetables such as squash and radish , and sunflowers . According to Bewley and Black (1978), Brazil nut storage is in hypocotyl and this place of storage is uncommon among seeds. All gymnosperm seeds are albuminous. The seed coat develops from the maternal tissue, the integuments , originally surrounding the ovule. The seed coat in

2890-416: The peach ) have a hardened fruit layer (the endocarp ) fused to and surrounding the actual seed. Nuts are the one-seeded, hard-shelled fruit of some plants with an indehiscent seed, such as an acorn or hazelnut . The first land plants evolved around 468 million years ago, and reproduced using spores. The earliest seed bearing plants to appear were the gymnosperms , which have no ovaries to contain

2975-555: The tonne and the spelling of ton in English is the same, though they differ in mass. One tonne is equivalent to: A tonne is the mass of one cubic metre of pure water at 4 °C (39 °F). As a non-SI unit, the use of SI metric prefixes with the tonne does not fall within the SI standard. For multiples of the tonne, it is more usual to speak of thousands or millions of tonnes. Kilotonne, megatonne, and gigatonne are more usually used for

3060-464: The Arizona walnut. Other sources list J. californica californica as native to southern California, and Juglans californica hindsii , or just J. hindsii , as native to northern California; in at least one case, these are given as "geographic variants" instead of subspecies (Botanica). Numerous walnut cultivars have been developed commercially, which are nearly all hybrids of the English walnut. During

3145-452: The US Food and Drug Administration (FDA) provided a qualified health claim allowing products containing walnuts to state: "Supportive but not conclusive research shows that eating 1.5 ounces (43 g) per day of walnuts, as part of a low saturated fat and low cholesterol diet and not resulting in increased caloric intake, may reduce the risk of coronary heart disease ." At the same time,

3230-552: The United States. It traditionally referred to a metric ton of ore containing 1% (i.e. 10 kg) of metal. The following excerpt from a mining geology textbook describes its usage in the particular case of tungsten: Tungsten concentrates are usually traded in metric tonne units (originally designating one tonne of ore containing 1% of WO 3 , today used to measure WO 3 quantities in 10 kg units. One metric tonne unit (mtu) of tungsten (VI) contains 7.93 kilograms of tungsten. In

3315-570: The accident and the discontinuation of walnut shells as a cleaning agent. Commercially, crushed walnut shells are still used outside of aviation for low-abrasive, less-toxic cleaning and blasting applications. In the oil and gas industry, deep bed filters of ground walnut shell are used for "polishing" (filtering) oily contaminates from water. At least two companies, LitterMaid and Naturally Fresh, make cat litter from ground walnut shells. Advantages cited over conventional clay litter include environmental sustainability of using what would otherwise be

3400-472: The agency refused to authorize the claim that "Diets including walnuts can reduce the risk of heart disease" and in 2010, it sent a warning letter to Diamond Foods stating there is "not sufficient evidence to identify a biologically active substance in walnuts that reduces the risk of coronary heart disease." In 2011, a scientific panel for the European Food Safety Authority recommended

3485-641: The case of uranium , MTU is sometimes used in the sense of metric ton of uranium (1,000 kg [2,200 lb]). The tonne of trinitrotoluene (TNT) is used as a proxy for energy, usually of explosions (TNT is a common high explosive ). Prefixes are used: kiloton(ne), megaton(ne), gigaton(ne), especially for expressing nuclear weapon yield , based on a specific combustion energy of TNT of about 4.2  MJ / kg (or one thermochemical calorie per milligram ). Hence, 1 t TNT = approx. 4.2  GJ , 1 kt TNT = approx. 4.2  TJ , 1 Mt TNT = approx. 4.2  PJ . The SI unit of energy

Walnut - Misplaced Pages Continue

3570-631: The cells are filled with starch , as for instance cereal grains , or not (non-farinaceous). The endosperm may also be referred to as "fleshy" or "cartilaginous" with thicker soft cells such as coconut , but may also be oily as in Ricinus (castor oil), Croton and Poppy . The endosperm is called "horny" when the cell walls are thicker such as date and coffee , or "ruminated" if mottled, as in nutmeg , palms and Annonaceae . In most monocotyledons (such as grasses and palms ) and some ( endospermic or albuminous ) dicotyledons (such as castor beans )

3655-405: The cells of the outer epidermis enlarge radially and their walls thicken, with nucleus and cytoplasm compressed into the outer layer. these cells which are broader on their inner surface are called palisade cells. In the inner epidermis, the cells also enlarge radially with plate like thickening of the walls. The mature inner integument has a palisade layer, a pigmented zone with 15–20 layers, while

3740-515: The common and recommended pronunciation is / t ɒ n / . In the United States, metric ton is the name for this unit used and recommended by NIST; an unqualified mention of a ton typically refers to a short ton of 2,000 lb (907.2 kg) and to a lesser extent to a long ton of 2,240 lb (1,016 kg), with the term tonne rarely used in speech or writing. Both terms are acceptable in Canadian English . Ton and tonne are both derived from

3825-407: The early 19th century, influential works being: Angiosperm seeds are "enclosed seeds", produced in a hard or fleshy structure called a fruit that encloses them for protection. Some fruits have layers of both hard and fleshy material. In gymnosperms, no special structure develops to enclose the seeds, which begin their development "naked" on the bracts of cones. However, the seeds do become covered by

3910-482: The effect of walnut supplementation on blood pressure found insufficient evidence to support walnut consumption as a blood pressure-lowering strategy. As of 2021, the relationship between walnut consumption and cognitive health is inconclusive. Walnut meats are available in two forms: in their shells or de-shelled. Due to processing, the meats may be whole, halved, or in smaller portions. All walnuts can be eaten on their own (raw, toasted, or pickled), or as part of

3995-440: The elaiosomes are eaten. The remainder of the seed, which is hard and inedible to the ants, then germinates either within the nest or at a removal site where the seed has been discarded by the ants. This dispersal relationship is an example of mutualism , since the plants depend upon the ants to disperse seeds, while the ants depend upon the plants seeds for food. As a result, a drop in numbers of one partner can reduce success of

4080-429: The embryo is embedded in the endosperm (and nucellus), which the seedling will use upon germination . In the non-endospermic dicotyledons the endosperm is absorbed by the embryo as the latter grows within the developing seed, and the cotyledons of the embryo become filled with stored food. At maturity, seeds of these species have no endosperm and are also referred to as exalbuminous seeds. The exalbuminous seeds include

4165-401: The energy of nuclear explosions and other events in equivalent mass of TNT , often loosely as approximate figures. When used in this context, there is little need to distinguish between metric and other tons, and the unit is spelled either as ton or tonne with the relevant prefix attached. A metric ton unit (mtu) can mean 10 kg (22 lb) within metal trading, particularly within

4250-579: The extended storage of walnuts is −3 to 0 °C (27 to 32 °F) with low humidity for industrial and home storage. However, such refrigeration technologies are unavailable in developing countries where walnuts are produced in large quantities; walnuts are best stored below 25 °C (77 °F) with low humidity. Temperatures above 30 °C (86 °F) and humidity levels above 70 percent can lead to rapid and high spoilage losses. Above 75 percent humidity threshold, fungal molds that release aflatoxin can form. In 2022, world production of walnuts (in shell)

4335-452: The fertilised ovule, an immature plant from which a new plant will grow under proper conditions. The embryo has one cotyledon or seed leaf in monocotyledons , two cotyledons in almost all dicotyledons and two or more in gymnosperms. In the fruit of grains (caryopses) the single monocotyledon is shield shaped and hence called a scutellum . The scutellum is pressed closely against the endosperm from which it absorbs food and passes it to

SECTION 50

#1732844119775

4420-645: The first few years of their lives deriving energy from the fungi and do not produce green leaves. At up to 55 pounds (25 kilograms) the largest seed is the coco de mer (Lodoicea maldivica). This indicates a 25 Billion fold difference in seed weight. Plants that produce smaller seeds can generate many more seeds per flower, while plants with larger seeds invest more resources into those seeds and normally produce fewer seeds. Small seeds are quicker to ripen and can be dispersed sooner, so autumn all blooming plants often have small seeds. Many annual plants produce great quantities of smaller seeds; this helps to ensure at least

4505-507: The fruits achenes , caryopses , nuts , samaras , and utricles . Other seeds are enclosed in fruit structures that aid wind dispersal in similar ways: Myrmecochory is the dispersal of seeds by ants . Foraging ants disperse seeds which have appendages called elaiosomes (e.g. bloodroot , trilliums , acacias , and many species of Proteaceae ). Elaiosomes are soft, fleshy structures that contain nutrients for animals that eat them. The ants carry such seeds back to their nest, where

4590-408: The fusion of two male gametes with the egg cell and the central cell to form the primary endosperm and the zygote. Right after fertilization, the zygote is mostly inactive, but the primary endosperm divides rapidly to form the endosperm tissue. This tissue becomes the food the young plant will consume until the roots have developed after germination . After fertilization, the ovules develop into

4675-446: The grasses, are not distinct structures, but are fused with the fruit wall to form a pericarp .) The testae of both monocots and dicots are often marked with patterns and textured markings, or have wings or tufts of hair. When the seed coat forms from only one layer, it is also called the testa, though not all such testae are homologous from one species to the next. The funiculus abscisses (detaches at fixed point – abscission zone),

4760-431: The growing parts. Embryo descriptors include small, straight, bent, curved, and curled. Within the seed, there usually is a store of nutrients for the seedling that will grow from the embryo. The form of the stored nutrition varies depending on the kind of plant. In angiosperms, the stored food begins as a tissue called the endosperm , which is derived from the mother plant and the pollen via double fertilization . It

4845-434: The inner epidermis may remain a single layer, it may also divide to produce two to three layers and accumulates starch, and is referred to as the colourless layer. By contrast, the outer epidermis becomes tanniferous . The inner integument may consist of eight to fifteen layers. As the cells enlarge, and starch is deposited in the outer layers of the pigmented zone below the outer epidermis, this zone begins to lignify, while

4930-444: The innermost layer is known as the fringe layer. In gymnosperms, which do not form ovaries, the ovules and hence the seeds are exposed. This is the basis for their nomenclature – naked seeded plants. Two sperm cells transferred from the pollen do not develop the seed by double fertilization, but one sperm nucleus unites with the egg nucleus and the other sperm is not used. Sometimes each sperm fertilizes an egg cell and one zygote

5015-588: The introduction of the SI in 1960; it has been used with this meaning in France since 1842, when there were no metric prefixes for multiples of 10 and above, and is now used as the standard spelling for the metric mass measurement in most English -speaking countries. In the United States, the unit was originally referred to using the French words millier or tonneau , but these terms are now obsolete. The British imperial and United States customary units are comparable to

5100-424: The latter example these hairs are the source of the textile crop cotton . Other seed appendages include the raphe (a ridge), wings, caruncles (a soft spongy outgrowth from the outer integument in the vicinity of the micropyle), spines, or tubercles. A scar also may remain on the seed coat, called the hilum , where the seed was attached to the ovary wall by the funicle. Just below it is a small pore, representing

5185-402: The main area of growth of the embryo, while the lower or micropylar pole produces the stalk-like suspensor that attaches to the micropyle. The suspensor absorbs and manufactures nutrients from the endosperm that are used during the embryo's growth. The main components of the embryo are: Monocotyledonous plants have two additional structures in the form of sheaths. The plumule is covered with

SECTION 60

#1732844119775

5270-422: The mature seed can be a paper-thin layer (e.g. peanut ) or something more substantial (e.g. thick and hard in honey locust and coconut ), or fleshy as in the sarcotesta of pomegranate . The seed coat helps protect the embryo from mechanical injury, predators, and drying out. Depending on its development, the seed coat is either bitegmic or unitegmic . Bitegmic seeds form a testa from the outer integument and

5355-509: The mechanical layer is the endotegmen, then the seed is endotestal. The exotesta may consist of one or more rows of cells that are elongated and pallisade like (e.g. Fabaceae ), hence 'palisade exotesta'. In addition to the three basic seed parts, some seeds have an appendage, an aril , a fleshy outgrowth of the funicle ( funiculus ), (as in yew and nutmeg ) or an oily appendage, an elaiosome (as in Corydalis ), or hairs (trichomes). In

5440-464: The megatonne (one teragram) and millitonne (one kilogram). If describing TNT equivalent units of energy, one megatonne of TNT is equivalent to approximately 4.184 petajoules . In English, tonne is an established spelling alternative to metric ton . In the United States and United Kingdom, tonne is usually pronounced the same as ton ( / t ʌ n / ), but the final "e" can also be pronounced, i.e. " tunnie " ( / ˈ t ʌ n i / ). In Australia,

5525-532: The metric ton in the United States, having been adopted by the United States National Institute of Standards and Technology (NIST). It is a symbol, not an abbreviation, and should not be followed by a period . Use of lower case is significant, and use of other letter combinations can lead to ambiguity. For example, T, MT, mT, are the SI symbols for the tesla , megatesla, and millitesla, respectively, while Mt and mt are SI-compatible symbols for

5610-456: The micropyle of the ovule. Seeds are very diverse in size. The dust-like orchid seeds are the smallest, with about one million seeds per gram; they are often embryonic seeds with immature embryos and no significant energy reserves. Orchids and a few other groups of plants are mycoheterotrophs which depend on mycorrhizal fungi for nutrition during germination and the early growth of the seedling. Some terrestrial orchid seedlings, in fact, spend

5695-399: The monocotyledons, ten in the dicotyledons, and two in the gymnosperms (linear and spatulate). This classification is based on three characteristics: embryo morphology, amount of endosperm and the position of the embryo relative to the endosperm. In endospermic seeds, there are two distinct regions inside the seed coat, an upper and larger endosperm and a lower smaller embryo. The embryo is

5780-403: The optimal conditions for survival of the resulting seedling; the second is spreading germination of a batch of seeds over time so a catastrophe (e.g. late frosts, drought, herbivory ) does not result in the death of all offspring of a plant ( bet-hedging ). Seed dormancy is defined as a seed failing to germinate under environmental conditions optimal for germination, normally when the environment

5865-472: The other. In South Africa , the Argentine ant ( Linepithema humile ) has invaded and displaced native species of ants. Unlike the native ant species, Argentine ants do not collect the seeds of Mimetes cucullatus or eat the elaiosomes. In areas where these ants have invaded, the numbers of Mimetes seedlings have dropped. Seed dormancy has two main functions: the first is synchronizing germination with

5950-410: The parent. The large, heavy root allows the seed to penetrate into the ground when it falls. Many garden plant seeds will germinate readily as soon as they have water and are warm enough; though their wild ancestors may have had dormancy, these cultivated plants lack it. After many generations of selective pressure by plant breeders and gardeners, dormancy has been selected out. For annuals , seeds are

6035-455: The plumule and radicle, acting as a protective covering. The maturing ovule undergoes marked changes in the integuments, generally a reduction and disorganization but occasionally a thickening. The seed coat forms from the two integuments or outer layers of cells of the ovule, which derive from tissue from the mother plant, the inner integument forms the tegmen and the outer forms the testa . (The seed coats of some monocotyledon plants, such as

6120-405: The rate of germination. This is given as a percent of germination over a certain amount of time, 90% germination in 20 days, for example. 'Dormancy' is covered above; many plants produce seeds with varying degrees of dormancy, and different seeds from the same fruit can have different degrees of dormancy. It's possible to have seeds with no dormancy if they are dispersed right away and do not dry (if

6205-486: The ripening process, the husk becomes brittle and the shell hard. The shell encloses the kernel or meat, which is usually in two halves separated by a membranous partition. The seed kernels – commonly available as shelled walnuts – are enclosed in a brown seed coat which contains antioxidants. The antioxidants protect the oil-rich seed from atmospheric oxygen, preventing rancidity . Walnut trees are late to grow leaves , typically not doing so until more than halfway through

6290-418: The same nut, such as: German Walnuss , Dutch walnoot , Danish valnød , and Swedish valnöt . Walnuts, like other tree nuts, must be processed and stored properly. Poor storage makes walnuts susceptible to insect and fungal mold infestations; the latter produces aflatoxin – a potent carcinogen . A batch that contains mold-infested walnuts should be entirely discarded. The ideal temperature for

6375-415: The scar forming an oval depression, the hilum . Anatropous ovules have a portion of the funiculus that is adnate (fused to the seed coat), and which forms a longitudinal ridge, or raphe , just above the hilum. In bitegmic ovules (e.g. Gossypium described here) both inner and outer integuments contribute to the seed coat formation. With continuing maturation the cells enlarge in the outer integument. While

6460-413: The seed affects its health and germination ability: since the seed has a living embryo, over time cells die and cannot be replaced. Some seeds can live for a long time before germination, while others can only survive for a short period after dispersal before they die. Seed vigor is a measure of the quality of seed, and involves the viability of the seed, the germination percentage, germination rate, and

6545-418: The seeds dry they go into physiological dormancy). There is great variation amongst plants and a dormant seed is still a viable seed even though the germination rate might be very low. Environmental conditions affecting seed germination include; water, oxygen, temperature and light. Tonnes The BIPM symbol for the tonne is t, adopted at the same time as the unit in 1879. Its use is also official for

6630-405: The seeds. The ovule consists of a number of components: The shape of the ovules as they develop often affects the final shape of the seeds. Plants generally produce ovules of four shapes: the most common shape is called anatropous , with a curved shape. Orthotropous ovules are straight with all the parts of the ovule lined up in a long row producing an uncurved seed. Campylotropous ovules have

6715-500: The seeds. They arose during the late Devonian period (416 million to 358 million years ago). From these early gymnosperms, seed ferns evolved during the Carboniferous period (359 to 299 million years ago); they had ovules that were borne in a cupule, which consisted of groups of enclosing branches likely used to protect the developing seed. Published literature about seed storage, viability and its hygrometric dependence began in

6800-420: The soil surface is the next phase of the plant's growth and is called seedling establishment. Three fundamental conditions must exist before germination can occur. (1) The embryo must be alive, called seed viability. (2) Any dormancy requirements that prevent germination must be overcome. (3) The proper environmental conditions must exist for germination. Far red light can prevent germination. Seed viability

6885-482: The spring. Walnut hulls contain diverse phytochemicals , such as polyphenols , that stain hands and can cause skin irritation. Seven phenolic compounds, including ferulic acid , vanillic acid , coumaric acid , syringic acid , myricetin , and juglone , were identified in walnut husks; juglone had concentrations of 2-4% fresh weight. Walnuts also contain the ellagitannin, pedunculagin . Regiolone has been isolated with juglone, betulinic acid and sitosterol from

6970-602: The stem bark of J. regia . The three species of walnuts most commonly grown for their seeds are the Persian (or English) walnut ( J. regia ), originating from Iran, the black walnut ( J. nigra ) – native to eastern North America – and the Japanese walnut, also known as the heartnut ( J. ailantifolia ). Other species include J. californica , the California black walnut (often used as a rootstock for commercial propagation of J. regia ), J. cinerea (butternuts), and J. major ,

7055-507: The strength of the seedlings produced. The germination percentage is simply the proportion of seeds that germinate from all seeds subject to the right conditions for growth. The germination rate is the length of time it takes for the seeds to germinate. Germination percentages and rates are affected by seed viability, dormancy and environmental effects that impact on the seed and seedling. In agriculture and horticulture quality seeds have high viability, measured by germination percentage plus

7140-459: Was 3.9 million tonnes , with China contributing 36% of the total (table). Other significant producers (in the order of decreasing harvest) were the United States, Iran , and Turkey . English walnuts without shells are 4% water, 15% protein , 65% fat, and 14% carbohydrates , including 7% dietary fiber (table). In a reference amount of 100 g (3.5 oz), walnuts provide 654 kilocalories (2,740 kJ) and rich contents (20% or more of

7225-426: Was referred to as albumen , and the inner endosperm layer as vitellus. Although misleading, the term began to be applied to all the nutrient matter. This terminology persists in referring to endospermic seeds as "albuminous". The nature of this material is used in both describing and classifying seeds, in addition to the embryo to endosperm size ratio. The endosperm may be considered to be farinaceous (or mealy) in which

#774225