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99-736: The Temperate Northern Pacific is a biogeographic region of the Earth's seas, comprising the temperate waters of the northern Pacific Ocean . The Temperate Northern Pacific connects, via the Bering Sea , to the Arctic marine realm , which includes the polar waters of the Arctic Ocean . To the south, it transitions to the tropical marine realms of the Pacific, including the Tropical Eastern Pacific along

198-451: A "commuting" distance to an island that has enough resources to be sustainable. The presence of an island is marked by seabirds , differences in cloud and weather patterns, as well as changes in the direction of waves. It is also possible for human populations to have gone extinct on islands, evidenced by explorers finding islands that show evidence of habitation but no life. Not all islands were or are inhabited by maritime cultures. In

297-509: A "rafting event". This phenomenon is known as oceanic dispersal . Tropical storms have the capacity to transport species over great distances. Animals like tortoises can live for weeks without food or water, and are able to survive floating on debris in the sea. One case study showed that in 1995, fifteen iguanas survived a 300 km journey to Anguilla in the Caribbean , an island which no iguana had lived on previously. They survived floating on

396-476: A combination of historical factors such as: speciation , extinction , continental drift , and glaciation . Through observing the geographic distribution of species, we can see associated variations in sea level , river routes, habitat, and river capture . Additionally, this science considers the geographic constraints of landmass areas and isolation, as well as the available ecosystem energy supplies. Over periods of ecological changes, biogeography includes

495-426: A detailed review of 1,288 islands found that they were home to 1,189 highly-threatened vertebrate species, which was 41 percent of the global figure. Coral bleaching is expected to occur with more frequency, threatening marine ecosystems, some of which island economies are dependent on. Some islands that are low-lying may cease to exist given high enough amounts of sea level rise. Tuvalu received media attention for

594-609: A few meters high, and are usually temporary. Changes in the flow speed, water level, and sediment content of the river may effect the rate of fluvial island formation and depletion. Permanent river islands also exist, the largest of which (that is completely inland) is Bananal Island in the Tocantins of Brazil, which has a maximum width of 55 kilometers. Lakes form for a variety of reasons, including glaciers , plate tectonics, and volcanism. Lake islands can form as part of these processes. The field of insular biogeography studies

693-421: A global scale. GIS can show certain processes on the earth's surface like whale locations, sea surface temperatures , and bathymetry. Current scientists also use coral reefs to delve into the history of biogeography through the fossilized reefs. Two global information systems are either dedicated to, or have strong focus on, biogeography (in the form of the spatial location of observations of organisms), namely

792-419: A great impact on Charles Darwin , who was inspired to consider species adaptations and evolution after learning about botanical geography. De Candolle was the first to describe the differences between the small-scale and large-scale distribution patterns of organisms around the globe. Several additional scientists contributed new theories to further develop the concept of biogeography. Charles Lyell developed

891-402: A higher carrying capacity also have more genetic diversity , which promotes speciation . Oceanic islands, ones that have never been connected to shore, are only populated by life that can cross the sea. This means that any animals present on the island had to have flown there, in the case of birds or bats , were carried by such animals, or were carried in a sea current in what is known as

990-472: A loss of almost all the infrastructure in Dominica. Sea level rise and other climate changes can reduce freshwater reserves, resulting in droughts . These risks are expected to decrease the habitability of islands, especially small ones. Beyond risks to human life, plant and animal life are threatened. It has been estimated that almost 50 percent of land species threatened by extinction live on islands. In 2017,

1089-403: A mass of uprooted trees from a storm. Plant species are thought to be able to travel great distances of ocean. New Zealand and Australia share 200 native plant species, despite being separated by 1500 km. Continental islands, islands that were at one point connected to a continent, are expected to share a common history of plant and animal life up until the point that the island broke away from

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1188-782: A minimum. Some islands became host to humans due to a land bridge or a continental island splitting from the mainland. Today, up to 10% of the world's population lives on islands. Islands are popular targets for tourism due to their perceived natural beauty, isolation, and unique cultures. Islands became the target of colonization by Europeans, resulting in the majority of islands in the Pacific being put under European control. Decolonization has resulted in some but not all island nations becoming self-governing , with lasting effects related to industrialisation , invasive species , nuclear weapons testing , and tourism . Islands and island countries are threatened by climate change . Sea level rise threatens to submerge nations such as Maldives ,

1287-743: A minority. Cultural attitudes related to communal ownership of land as well as a lack of individualistic decision-making may make some island cultures less compatible with the global capitalist economy, causing these nations to experience less economic growth. Islands have long been a popular target for tourism , thanks to their unique climates, cultures, and natural beauty. However, islands may suffer from poor transportation connectivity from airplanes and boats and strains on infrastructure from tourist activity. Islands in colder climates often rely on seasonal tourists seeking to enjoy nature or local cultures, and may only be one aspect of an island's economy. In contrast, tourism on tropical islands can often make up

1386-413: A number of methods have been developed to produce arguably more complete "predictive" or "modelled" distributions for species based on their associated environmental or other preferences (such as availability of food or other habitat requirements); this approach is known as either Environmental niche modelling (ENM) or Species distribution modelling (SDM). Depending on the reliability of the source data and

1485-599: A permanent caisson , a steel or concrete structure built in a closed loop and then filled with sand. Some modern islands have been constructed by pouring millions of tons of sand into the sea, such as with Pearl Island in Qatar or the Palm Islands in Dubai. These islands are usually created for real estate development , and are sold for private ownership or construction of housing. Offshore oil platforms have also been described as

1584-524: A phenomenon where new oceanic crust is formed, pushing away older crust. Islands sitting on the continental shelf may be called continental islands. Other islands, like those that make up New Zealand , are what remains of continents that shrank and sunk beneath the sea. It was estimated that Zealandia , the continent-like area of crust that New Zealand sits on, has had 93% of its original surface area submerged. Some islands are formed when coral reefs grow on volcanic islands that have submerged beneath

1683-423: A press conference publicizing the ongoing submerging of the island country. Tuvalu signed a cooperation agreement with Australia agreeing to annually allow 280 of its citizens to become permanent residents of Australia. The Marshall Islands, a country of 1,156 islands, have also been identified as a country that may be existentially threatened by rising seas. Increasing intensity of tropical storms also increases

1782-483: A priority target of conservation efforts, to prevent the extinction of these species. Despite high levels of endemism, the total species richness , the total number of unique species in a region, is lower on islands than on mainlands. The level of species richness on islands is proportional to the area of that island, a phenomenon known as the species-area relationship . This is because larger areas have more resources and thus can support more organisms. Populations with

1881-411: A restricted set of natural resources. However, these nations control some of the largest fisheries in the world, deposits of copper , gold , and nickel , as well as oil deposits . The natural beauty of island nations also makes them a magnet for tourism . Islands also have geopolitical value for naval bases , weapons testing , and general territorial control. One such example is French Polynesia ,

1980-987: A result of tectonic uplift (or subsidence ), natural damming created by a landslide , or headward or lateral erosion of the watershed between adjacent basins. Biogeography is a synthetic science, related to geography , biology , soil science , geology , climatology , ecology and evolution . Some fundamental concepts in biogeography include: The study of comparative biogeography can follow two main lines of investigation: There are many types of biogeographic units used in biogeographic regionalisation schemes, as there are many criteria ( species composition , physiognomy , ecological aspects) and hierarchization schemes: biogeographic realms (ecozones), bioregions ( sensu stricto ), ecoregions , zoogeographical regions , floristic regions , vegetation types, biomes , etc. The terms biogeographic unit, biogeographic area can be used for these categories, regardless of rank. In 2008, an International Code of Area Nomenclature

2079-468: A selection pressure for smaller animals that require less food. Having fewer predators would mean these animals did not need not be large to survive. Charles Darwin formulated the theory of natural selection through the study of island ecology. The species he observed on the Galápagos Islands , including tanager birds, contributed to his understanding of how evolution works. He first traveled to

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2178-509: A specific property known as adaptive radiation . In this process, a species that arrives on a group of islands rapidly becomes more diverse over time, splitting off into new species or subspecies. A species that reaches an island ecosystem may face little competition for resources, or may find that the resources that they found in their previous habitat are not available. These factors together result in individual evolutionary branches with different means of survival. The classical example of this

2277-414: A territory that receives substantial military expenditure and aid from France. Since the first discoveries of Polynesian, Micronesian , and other islands by Westerners, these nations have been the subject of colonization. Islands were the target of Christian missionaries . These missionaries faced resistance, but found success when some local chiefs used European support to centralize power. Beginning in

2376-514: A tree that is present in Australia, New Zealand, parts of South American, and New Guinea, places that today are geographically distant. A possible explanation for this phenomenon is that these landmasses were once all part of the continent Gondwana and separated by tectonic drift. However, there are competing theories that suggest this species may have reached faraway places by way of oceanic dispersal. Species that colonize island archipelagos exhibit

2475-603: A type of island. Some atolls have been covered in concrete to create artificial islands for military purposes, such as those created by China in the South China Sea . These atolls were previously low-tide elevations, landmasses that are only above water during low tide . The United Nations Convention on the Law of the Sea indicates that these islands may not have the same legal status as a naturally occurring island, and as such may not confer

2574-440: A whole suite of predictor variables for biogeographic analysis, including satellite imaging and processing of the Earth. Two main types of satellite imaging that are important within modern biogeography are Global Production Efficiency Model (GLO-PEM) and Geographic Information Systems (GIS). GLO-PEM uses satellite-imaging gives "repetitive, spatially contiguous, and time specific observations of vegetation". These observations are on

2673-507: Is Darwin's finches , a group of up to fifteen tanager species that are endemic to the Galápagos Islands. These birds evolved different beaks in order to eat different kinds of food available on the islands. The large ground finch has a large bill used to crack seeds and eat fruit. The Genovesa cactus finch prefers cacti as a food source, and has a beak adapted for removing pulp and flowers from cacti. The green warbler-finch (in

2772-726: Is a piece of land, distinct from a continent, completely surrounded by water. There are continental islands, which were formed by being split from a continent by plate tectonics , and oceanic islands, which have never been part of a continent. Oceanic islands can be formed from volcanic activity, grow into atolls from coral reefs , and form from sediment along shorelines, creating barrier islands . River islands can also form from sediment and debris in rivers. Artificial islands are those made by humans, including small rocky outcroppings built out of lagoons and large-scale land reclamation projects used for development. Islands are host to diverse plant and animal life. Oceanic islands have

2871-588: Is actually a cognate of Swedish ö and German Aue , and more distantly related to Latin aqua (water). The spelling of the word was modified in the 15th century because of a false etymology caused by an association with the Old French loanword isle , which itself comes from the Latin word insula . Islands often are found in archipelagos or island chains, which are collections of islands. These chains are thought to form from volcanic hotspots , areas of

2970-424: Is an alternate view than that of Linnaeus. Buffon's law eventually became a principle of biogeography by explaining how similar environments were habitats for comparable types of organisms. Buffon also studied fossils which led him to believe that the Earth was over tens of thousands of years old, and that humans had not lived there long in comparison to the age of the Earth. Following the period of exploration came

3069-631: Is further subdivided into marine provinces, and the marine provinces divided into marine ecoregions: Biogeography Knowledge of spatial variation in the numbers and types of organisms is as vital to us today as it was to our early human ancestors , as we adapt to heterogeneous but geographically predictable environments . Biogeography is an integrative field of inquiry that unites concepts and information from ecology , evolutionary biology , taxonomy , geology , physical geography , palaeontology , and climatology . Modern biogeographic research combines information and ideas from many fields, from

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3168-404: Is in the geological similarities between varying locations around the globe, the geographic distribution of some fossils (including the mesosaurs ) on various continents, and the jigsaw puzzle shape of the landmasses on Earth. Though Wegener did not know the mechanism of this concept of Continental Drift, this contribution to the study of biogeography was significant in the way that it shed light on

3267-473: Is on how the environment and humans affect the distribution of species as well as other manifestations of Life such as species or genetic diversity. Biogeography is being applied to biodiversity conservation and planning, projecting global environmental changes on species and biomes, projecting the spread of infectious diseases, invasive species, and for supporting planning for the establishment of crops. Technological evolving and advances have allowed for generating

3366-497: Is the apple snail , initially introduced to the U.S. by aquarium owners. It has since been transported by hurricanes across the Gulf Coast and neighboring islands. These species compete for resources with native animals, and some may grow so densely that they displace other forms of existing life. For hundreds of years, islands have been created through land reclamation . One of the first recorded instances of this when people of

3465-628: Is the giant tortoise of the Seychelles , though it is unknown if it grew in size before or after reaching the island. Larger animals such as the hippopotamus tend to become smaller, such as in the case of the pygmy hippopotamus . This is known as insular dwarfism . In the case of smaller animals, it has been hypothesized that animals on islands may have fewer predators and competitors, resulting in selection pressure towards larger animals. Larger animals may exhaust food resources quickly due to their size, causing malnutrition in their young, resulting in

3564-607: The Age of Enlightenment in Europe, which attempted to explain the patterns of biodiversity observed by Buffon and Linnaeus. At the birth of the 19th century, Alexander von Humboldt, known as the "founder of plant geography", developed the concept of physique generale to demonstrate the unity of science and how species fit together. As one of the first to contribute empirical data to the science of biogeography through his travel as an explorer, he observed differences in climate and vegetation. The Earth

3663-529: The Atlantic Ocean in search of an alternate route to the East Indies . These historians theorize that successful explorers were rewarded with recognition and wealth, leading others to attempt possibly dangerous expeditions to discover more islands, usually with poor results. About 10% of the world's population lives on islands. The study of the culture of islands is known as island studies . The interest in

3762-667: The Global Biodiversity Information Facility (GBIF: 2.57 billion species occurrence records reported as at August 2023) and, for marine species only, the Ocean Biodiversity Information System (OBIS, originally the Ocean Biogeographic Information System : 116 million species occurrence records reported as at August 2023), while at a national scale, similar compilations of species occurrence records also exist such as

3861-569: The Marshall Islands , and Tuvalu completely. Increases in the frequency and intensity of tropical cyclones can cause widespread destruction of infrastructure and animal habitats. Species that live exclusively on islands are some of those most threatened by extinction . An island is an area of land surrounded by water on all sides that is distinct from a continent. There is no standard of size that distinguishes islands and continents . Continents have an accepted geological definition – they are

3960-501: The Polynesians . Many of the previous island settlements required traveling distances of less than 100 km (62 mi), whereas Polynesians may have traveled 2,000–3,200 km (1,200–2,000 mi) to settle islands such as Tahiti . They would send navigators to sail the ocean without the aid of navigational instruments to discover new islands for settlement. Between 1100 and 800 BC, Polynesians sailed East from New Guinea and

4059-869: The Solomon Islands and reached the islands that make up the modern-day Fiji and Samoa . The furthest extent of this migration would be Easter Island in the East, and New Zealand in the South, with New Zealand's first settlements between 1250 and 1300. Historians have sought to understand why some remote islands have always been uninhabited, while others, especially in the Pacific Ocean , have long been populated by humans. Generally, larger islands are more likely to be able to sustain humans and thus are more likely to have been settled. Small islands that cannot sustain populations on their own can still be habitable if they are within

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4158-493: The Solomon Islands created eighty such islands by piling coral and rock in the Lau Lagoon . One traditional way of constructing islands is with the use of a revetment . Sandbags or stones are dropped with a barge into the sea to bring the land level slightly out of the water. The island area is then filled with sand or gravel, followed by a construction of this revetment to hold it together. Islands have also been constructed with

4257-414: The lithosphere where the mantle is hotter than the surrounding area. These hotspots would give rise to volcanoes whose lava would form the rock the islands are made of. For some islands, the movement of tectonic plates above stationary hotspots would form islands in a linear chain, with the islands further away from the hotspot being progressively older and more eroded , before disappearing under

4356-450: The sea as a natural barrier to the introduction of new species, causing the species that do reach the island to evolve in isolation. Continental islands share animal and plant life with the continent they split from. Depending on how long ago the continental island formed, the life on that island may have diverged greatly from the mainland due to natural selection . Humans have lived on and traveled between islands for thousands of years at

4455-522: The 16th century, European states placed most of Oceania in under colonial administration . Pohnpei was colonized by Spain as early as 1526. It changed hands from Germany to Japan to the United States before joining the Federated States of Micronesia in 1982, maintaining a "free association" status with the U.S. Guam was a Spanish territory until 1898, and now is a unincorporated territory of

4554-476: The 36 volume Histoire Naturelle, générale et particulière , in which he argued that varying geographical regions would have different forms of life. This was inspired by his observations comparing the Old and New World, as he determined distinct variations of species from the two regions. Buffon believed there was a single species creation event, and that different regions of the world were homes for varying species, which

4653-505: The Amazon basin, Orinoco basin, and Guianas ) with an exceptionally low (flat) topographic relief, the many waterways have had a highly reticulated history over geological time . In such a context, stream capture is an important factor affecting the evolution and distribution of freshwater organisms. Stream capture occurs when an upstream portion of one river drainage is diverted to the downstream portion of an adjacent basin. This can happen as

4752-588: The Gómez Farias Region, Tamaulipas, Mexico , which has been described as "ground-breaking" and "a classic treatise in historical biogeography". Martin applied several disciplines including ecology , botany , climatology , geology , and Pleistocene dispersal routes to examine the herpetofauna of a relatively small and largely undisturbed area, but ecologically complex, situated on the threshold of temperate – tropical (nearctic and neotropical) regions, including semiarid lowlands at 70 meters elevation and

4851-506: The Hawaiian islands being home to irrigated fields of taro, whereas in some islands, like Tahiti, breadfruit was more widely cultivated and fermented in order to preserve it. There is archeological evidence that Canary Islanders would chew the roots of ferns for sustenance, a practice that wore heavily on their molars . These islanders would also grow barley and raised livestock such as goats . Many island nations have little land and

4950-604: The Indian Ocean was much narrower than it is today, and that South America was closer to the Antarctic, one would be hard pressed to explain the presence of many "ancient" lineages of perching birds in Africa, as well as the mainly South American distribution of the suboscines . Paleobiogeography also helps constrain hypotheses on the timing of biogeographic events such as vicariance and geodispersal , and provides unique information on

5049-448: The Origin of Species were devoted to geographical distribution. The first discoveries that contributed to the development of biogeography as a science began in the mid-18th century, as Europeans explored the world and described the biodiversity of life. During the 18th century most views on the world were shaped around religion and for many natural theologists, the bible. Carl Linnaeus , in

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5148-617: The Origin of Species . The first evidence of humans colonizing islands probably occurred in the Paleolithic era, 100,000 to 200,000 years ago. Reaching the Indonesian islands of Flores and Timor would have required crossing distances of water of at least 29 km (18 mi). Some islands, such as Honshu , were probably connected to the mainland with a land bridge that allowed humans to colonize it before it became an island. The first people to colonize distant oceanic islands were

5247-780: The Pacific coast of the Americas, the Eastern Indo-Pacific in the central Pacific Ocean, and the Central Indo-Pacific of the western Pacific basin. The Taiwan Strait forms the boundary between the Temperate Northern Pacific and the Central Indo-Pacific. Characteristic fauna include the Pacific salmon and trout ( Oncorhynchus spp.), gray whale (Eschrichtius robustus) , and North Pacific right whale (Eubalaena japonica). The Temperate Northern Pacific

5346-478: The Theory of Continental Drift in 1912, though it was not widely accepted until the 1960s. This theory was revolutionary because it changed the way that everyone thought about species and their distribution around the globe. The theory explained how continents were formerly joined in one large landmass, Pangea , and slowly drifted apart due to the movement of the plates below Earth's surface. The evidence for this theory

5445-603: The Theory of Uniformitarianism after studying fossils. This theory explained how the world was not created by one sole catastrophic event, but instead from numerous creation events and locations. Uniformitarianism also introduced the idea that the Earth was actually significantly older than was previously accepted. Using this knowledge, Lyell concluded that it was possible for species to go extinct. Since he noted that Earth's climate changes, he realized that species distribution must also change accordingly. Lyell argued that climate changes complemented vegetation changes, thus connecting

5544-631: The U.K. National Biodiversity Network , the Atlas of Living Australia , and many others. In the case of the oceans, in 2017 Costello et al. analyzed the distribution of 65,000 species of marine animals and plants as then documented in OBIS, and used the results to distinguish 30 distinct marine realms, split between continental-shelf and offshore deep-sea areas. Since it is self evident that compilations of species occurrence records cannot cover with any completeness, areas that have received either limited or no sampling,

5643-481: The U.S. The decolonization era saw many island states achieve independence or some form of self-governance . Nuclear weapons testing on the Marshall Islands left many atolls destroyed or uninhabitable, causing the forced displacement of people from their home islands as well as increases in cancer rates due to radiation . Colonization has resulted in a decline of observance of traditional cultural practices in places such as Hawaii, where Native Hawaiians are now

5742-414: The anticipated effects of climate change can also be used to show potential changes in species distributions that may occur in the future based on such scenarios. Paleobiogeography goes one step further to include paleogeographic data and considerations of plate tectonics . Using molecular analyses and corroborated by fossils , it has been possible to demonstrate that perching birds evolved first in

5841-421: The biotic and abiotic features of the Earth in his book, Cosmos . Augustin de Candolle contributed to the field of biogeography as he observed species competition and the several differences that influenced the discovery of the diversity of life. He was a Swiss botanist and created the first Laws of Botanical Nomenclature in his work, Prodromus. He discussed plant distribution and his theories eventually had

5940-406: The continent. For example, the presence of freshwater fish on an island surrounded by ocean would indicate that it once was attached to a continent, since these fish cannot traverse the ocean on their own. Over the course of time, evolution and extinction changes the nature of animal life on a continental island, but only once it splits from the mainland. An example is that of the southern beech ,

6039-451: The deposition of sediment by waves . These islands erode and grow as the wind and waves shift. Barrier islands have the effect of protecting coastal areas from severe weather because they absorb some of the energy of large waves before they can reach the shore. A fluvial island is an island that forms from the erosion and sedimentation of debris in rivers; almost all rivers have some form of fluvial islands. These islands may only be

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6138-475: The development of biogeography as a science. The scientific theory of biogeography grows out of the work of Alexander von Humboldt (1769–1859), Francisco Jose de Caldas (1768–1816), Hewett Cottrell Watson (1804–1881), Alphonse de Candolle (1806–1893), Alfred Russel Wallace (1823–1913), Philip Lutley Sclater (1829–1913) and other biologists and explorers. The patterns of species distribution across geographical areas can usually be explained through

6237-420: The development of the fields of conservation biology and landscape ecology . Classic biogeography has been expanded by the development of molecular systematics , creating a new discipline known as phylogeography . This development allowed scientists to test theories about the origin and dispersal of populations, such as island endemics . For example, while classic biogeographers were able to speculate about

6336-426: The distances and frequency with which invasive species may be transported to islands. Floodwaters from these storms may also wash plants further inland than they would travel on their own, introducing them to new habitats. Agriculture and trade also have introduced non-native life to islands. These processes result in an introduction of invasive species to ecosystems that are especially small and fragile. One example

6435-400: The ecological processes that take place on islands, with a focus on what factors effect the evolution , extinction , and richness of species. Scientists often study islands as an isolated model of how the process of natural selection takes place. Island ecology studies organisms on islands and their environment. It has yielded important insights for its parent field of ecology since

6534-496: The environmental surroundings to varying species. This largely influenced Charles Darwin in his development of the theory of evolution. Charles Darwin was a natural theologist who studied around the world, and most importantly in the Galapagos Islands . Darwin introduced the idea of natural selection, as he theorized against previously accepted ideas that species were static or unchanging. His contributions to biogeography and

6633-597: The factors affecting organism distribution, and to predict future trends in organism distribution. Often mathematical models and GIS are employed to solve ecological problems that have a spatial aspect to them. Biogeography is most keenly observed on the world's islands . These habitats are often much more manageable areas of study because they are more condensed than larger ecosystems on the mainland. Islands are also ideal locations because they allow scientists to look at habitats that new invasive species have only recently colonized and can observe how they disperse throughout

6732-453: The formation of regional biotas. For example, data from species-level phylogenetic and biogeographic studies tell us that the Amazonian teleost fauna accumulated in increments over a period of tens of millions of years, principally by means of allopatric speciation, and in an arena extending over most of the area of tropical South America (Albert & Reis 2011). In other words, unlike some of

6831-477: The former Lifemapper project at the University of Kansas (now continued as a part of BiotaPhy ) and AquaMaps , which as at 2023 contain modelled distributions for around 200,000 terrestrial, and 33,000 species of teleosts , marine mammals and invertebrates, respectively. One advantage of ENM/SDM is that in addition to showing current (or even past) modelled distributions, insertion of changed parameters such as

6930-641: The geographical distribution of organisms around the globe. Alfred Russel Wallace studied the distribution of flora and fauna in the Amazon Basin and the Malay Archipelago in the mid-19th century. His research was essential to the further development of biogeography, and he was later nicknamed the "father of Biogeography". Wallace conducted fieldwork researching the habits, breeding and migration tendencies, and feeding behavior of thousands of species. He studied butterfly and bird distributions in comparison to

7029-498: The habit of true warbler species) consumes spiders and insects that live on plants. Other examples of this phenomenon exist worldwide, including in Hawaii and Madagascar, and are not limited to island ecosystems. Species endemic to islands show a common evolutionary trajectory. Foster's rule (also known as the island rule), states that small mammals such as rodents evolve to become larger, known as island gigantism . One such example

7128-411: The importance of environmental and geographic similarities or differences as a result of climate and other pressures on the planet. Importantly, late in his career Wegener recognised that testing his theory required measurement of continental movement rather than inference from fossils species distributions. In 1958 paleontologist Paul S. Martin published A Biogeography of Reptiles and Amphibians in

7227-556: The island and change it. They can then apply their understanding to similar but more complex mainland habitats. Islands are very diverse in their biomes , ranging from the tropical to arctic climates. This diversity in habitat allows for a wide range of species study in different parts of the world. One scientist who recognized the importance of these geographic locations was Charles Darwin , who remarked in his journal "The Zoology of Archipelagoes will be well worth examination". Two chapters in On

7326-411: The islands as a naturalist on HMS Beagle in 1835, as part of a five-year circumnavigation of Earth. He wrote that "the different islands to a considerable extent are inhabited by a different set of beings". Through the study of the finches and other animals he realized that organisms survive by changing to adapt to their habitat. It would be over twenty years before he published his theories in On

7425-643: The largest landmass of a particular tectonic plate . Islands can occur in any body of water, including lakes , rivers , seas . Low-tide elevations , areas of land that are not above the surface during a high tide , are generally not considered islands. Islands that have been bridged or otherwise joined to a mainland with land reclamation are sometimes considered "de-islanded", but not in every case. The word island derives from Middle English iland , from Old English igland (from ig or ieg , similarly meaning 'island' when used independently, and -land carrying its contemporary meaning. Old English ieg

7524-404: The mainland, they had no need for boats. The motivation for island exploration has been the subject of research and debate. Some early historians previously argued that early island colonization was unintentional, perhaps by a raft being swept out to sea. Others compare the motivations of Polynesian and similar explorers with those of Christopher Columbus , the explorer who sailed westward over

7623-589: The majority of the local economy and built environment . These islands sometimes also require consistent foreign aid on top of tourism in order to ensure economic growth. This reliance can result in social inequality and environmental degradation . During tourism downturns, these economies struggle to make up the lost inflow of cash with other industries. Climate change threatens human development on islands due to sea level rise , more dangerous tropical cyclones , coral bleaching , and an increase in invasive species . For example, in 2017 Hurricane Maria caused

7722-479: The mid-18th century, improved our classifications of organisms through the exploration of undiscovered territories by his students and disciples. When he noticed that species were not as perpetual as he believed, he developed the Mountain Explanation to explain the distribution of biodiversity; when Noah's ark landed on Mount Ararat and the waters receded, the animals dispersed throughout different elevations on

7821-515: The most important and consequential developments in biogeography has been to show how multiple organisms, including mammals like monkeys and reptiles like squamates , overcame barriers such as large oceans that many biogeographers formerly believed were impossible to cross. See also Oceanic dispersal . Biogeography now incorporates many different fields including but not limited to physical geography, geology, botany and plant biology, zoology, general biology, and modelling. A biogeographer's main focus

7920-499: The mountain. This showed different species in different climates proving species were not constant. Linnaeus' findings set a basis for ecological biogeography. Through his strong beliefs in Christianity, he was inspired to classify the living world, which then gave way to additional accounts of secular views on geographical distribution. He argued that the structure of an animal was very closely related to its physical surroundings. This

8019-683: The nature of the models employed (including the scales for which data are available), maps generated from such models may then provide better representations of the "real" biogeographic distributions of either individual species, groups of species, or biodiversity as a whole, however it should also be borne in mind that historic or recent human activities (such as hunting of great whales , or other human-induced exterminations) may have altered present-day species distributions from their potential "full" ecological footprint. Examples of predictive maps produced by niche modelling methods based on either GBIF (terrestrial) or OBIS (marine, plus some freshwater) data are

8118-476: The northernmost cloud forest in the western hemisphere at over 2200 meters. The publication of The Theory of Island Biogeography by Robert MacArthur and E.O. Wilson in 1967 showed that the species richness of an area could be predicted in terms of such factors as habitat area, immigration rate and extinction rate. This added to the long-standing interest in island biogeography . The application of island biogeography theory to habitat fragments spurred

8217-571: The origins of species in the Hawaiian Islands , phylogeography allows them to test theories of relatedness between these populations and putative source populations on various continents, notably in Asia and North America . Biogeography continues as a point of study for many life sciences and geography students worldwide, however it may be under different broader titles within institutions such as ecology or evolutionary biology. In recent years, one of

8316-515: The past, some societies were found to have lost their seafaring ability over time, such as the case of the Canary Islands , which were occupied by an indigenous people since the island's first discovery in the first century until being conquered by the Spanish Empire in 1496. It has been hypothesized that since the inhabitants had little incentive for trade and had little to any contact with

8415-495: The physiological and ecological constraints on organismal dispersal to geological and climatological phenomena operating at global spatial scales and evolutionary time frames. The short-term interactions within a habitat and species of organisms describe the ecological application of biogeography. Historical biogeography describes the long-term, evolutionary periods of time for broader classifications of organisms. Early scientists, beginning with Carl Linnaeus , contributed to

8514-448: The presence or absence of geographical barriers. His observations led him to conclude that the number of organisms present in a community was dependent on the amount of food resources in the particular habitat. Wallace believed species were dynamic by responding to biotic and abiotic factors. He and Philip Sclater saw biogeography as a source of support for the theory of evolution as they used Darwin's conclusion to explain how biogeography

8613-420: The region of Australia or the adjacent Antarctic (which at that time lay somewhat further north and had a temperate climate). From there, they spread to the other Gondwanan continents and Southeast Asia – the part of Laurasia then closest to their origin of dispersal – in the late Paleogene , before achieving a global distribution in the early Neogene . Not knowing that at the time of dispersal,

8712-717: The sea entirely. An example is the Hawaiian Islands , with the oldest island being 25 million years old, and the youngest, Hawaii , still being an active volcano . However, not all island chains are formed this way. Some may be formed all at once by fractures in the tectonic plates themselves, simultaneously creating multiple islands. One supporting piece of evidence is that of the Line Islands , which are all estimated to be 8 million years old, rather than being different ages. Other island chains form due to being separated from existing continents. The Japanese archipelago may have been separated from Eurasia due to seafloor spreading ,

8811-549: The study of islands is due to their unique cultures and natural environments that differ from mainland cultures. This is for a few reasons: First, the obvious political and geographic isolation from mainland cultures. Second, unique restraints on resources and ecology creating marine-focused cultures with a focus on fishing and sailing. Third, a lasting historical and political significance of islands. The Polynesian diet got most of its protein from fishing. Polynesians were known to fish close to shore, as well as in deep water. It

8910-428: The study of plant and animal species in: their past and/or present living refugium habitat ; their interim living sites; and/or their survival locales. As writer David Quammen put it, "...biogeography does more than ask Which species? and Where . It also asks Why? and, what is sometimes more crucial, Why not? ." Modern biogeography often employs the use of Geographic Information Systems (GIS), to understand

9009-450: The surface. When these coral islands encircle a central lagoon , the island is known as an atoll . The formation of reefs and islands related to those reefs is aided by the buildup of sediment in shallow patches of water. In some cases, tectonic movements lifting a reef out of the water by as little as 1 meter can cause sediment to accumulate and an island to form. Barrier islands are long, sandy bars that form along shorelines due to

9108-416: The theory of evolution were different from those of other explorers of his time, because he developed a mechanism to describe the ways that species changed. His influential ideas include the development of theories regarding the struggle for existence and natural selection. Darwin's theories started a biological segment to biogeography and empirical studies, which enabled future scientists to develop ideas about

9207-438: The time of Charles Darwin . In biology, endemism is defined as the phenomenon where species or genus is only found in a certain geographical area. Islands isolate land organisms from others with water, and isolate aquatic organisms living on them with land. Island ecosystems have the highest rates of endemism globally. This means that islands contribute heavily to global biodiversity . Areas with high lives of biodiversity are

9306-447: The well-known insular faunas ( Galapagos finches , Hawaiian drosophilid flies, African rift lake cichlids ), the species-rich Amazonian ichthyofauna is not the result of recent adaptive radiations . For freshwater organisms, landscapes are divided naturally into discrete drainage basins by watersheds , episodically isolated and reunited by erosional processes. In regions like the Amazon Basin (or more generally Greater Amazonia,

9405-400: Was divided into regions which he defined as tropical, temperate, and arctic and within these regions there were similar forms of vegetation. This ultimately enabled him to create the isotherm, which allowed scientists to see patterns of life within different climates. He contributed his observations to findings of botanical geography by previous scientists, and sketched this description of both

9504-529: Was important to a George Louis Buffon's rival theory of distribution. Closely after Linnaeus, Georges-Louis Leclerc, Comte de Buffon observed shifts in climate and how species spread across the globe as a result. He was the first to see different groups of organisms in different regions of the world. Buffon saw similarities between some regions which led him to believe that at one point continents were connected and then water separated them and caused differences in species. His hypotheses were described in his work,

9603-660: Was proposed for biogeography. It achieved limited success; some studies commented favorably on it, but others were much more critical, and it "has not yet gained a significant following". Similarly, a set of rules for paleobiogeography has achieved limited success. In 2000, Westermann suggested that the difficulties in getting formal nomenclatural rules established in this field might be related to "the curious fact that neither paleo- nor neobiogeographers are organized in any formal groupings or societies, nationally (so far as I know) or internationally — an exception among active disciplines." Island An island or isle

9702-470: Was reported that Rapa Nui people were known to fish as far as 500 km (310 mi) from shore at coral reefs. Spear , line , and net fishing were all used, to catch tuna as well as sharks and stingrays . Island cultures also cultivate native and non-native crops. Polynesians grew the native yam , taro , breadfruit , banana , coconut and other fruits and vegetables. Different island climates made different resources more important, such as

9801-496: Was similar to a record of species inheritance. Key findings, such as the sharp difference in fauna either side of the Wallace Line , and the sharp difference that existed between North and South America prior to their relatively recent faunal interchange , can only be understood in this light. Otherwise, the field of biogeography would be seen as a purely descriptive one. Moving on to the 20th century, Alfred Wegener introduced

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