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Pollination syndromes are suites of flower traits that have evolved in response to natural selection imposed by different pollen vectors , which can be abiotic (wind and water) or biotic, such as birds, bees, flies, and so forth through a process called pollinator-mediated selection . These traits include flower shape, size, colour, odour , reward type and amount, nectar composition, timing of flowering, etc. For example, tubular red flowers with copious nectar often attract birds; foul smelling flowers attract carrion flies or beetles, etc.

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66-529: Sykia or Sykea (Greek meaning fig tree ) may refer to a number of places in Greece: Sykia, Chalkidiki , a village in the municipal unit Toroni, Chalkidiki Sykia, Corinthia , a village in the municipal unit Xylokastro, Corinthia Sykia, Imathia , a village in the municipal unit Vergina, Imathia Sykia, Laconia , a village in Laconia Sykia, Larissa ,

132-419: A syconium , derived from an arrangement of many small flowers on an inverted, nearly closed receptacle. The many small flowers are unseen unless the fig is cut open. The fruit typically has a bulbous shape with a small opening (the ostiole ) at the outward end that allows access to pollinators . The flowers are pollinated by very small wasps such as Pegoscapus that crawl through the opening in search of

198-404: A constant source of inspiration and wonder to biologists. Notably, three vegetative traits together are unique to figs. All figs present a white to yellowish latex , some in copious quantities; the twig shows paired stipules —or circular scars if the stipules have fallen off; the lateral veins at the base of the leaf are steep, forming a tighter angle with the midrib than the other lateral veins,

264-504: A continuum of greater or lesser specialization or generalization onto particular functional groups of pollinators that exert similar selective pressures" and the frequency with which flowers conform to the expectations of the pollination syndromes is relatively rare. In addition, new types of plant-pollinator interaction, involving "unusual" pollinating animals are regularly being discovered, such as specialized pollination by spider hunting wasps ( Pompilidae ) and fruit chafers ( Cetoniidae ) in

330-597: A distinctive shape or habit, and their fruits distinguish them from other plants. The fruit of Ficus is an inflorescence enclosed in an urn-like structure called a syconium , which is lined on the inside with the fig's tiny flowers that develop into multiple ovaries on the inside surface. In essence, the fig fruit is a fleshy stem with multiple tiny flowers that fruit and coalesce. The unique fig pollination system, involving tiny, highly specific wasps, known as fig wasps that enter via ostiole these subclosed inflorescences to both pollinate and lay their own eggs, has been

396-418: A feature referred to as "triveined". Current molecular clock estimates indicate that Ficus is a relatively ancient genus, being at least 60 million years old, and possibly as old as 80 million years. The main radiation of extant species, however, may have taken place more recently, between 20 and 40 million years ago. Some better-known species that represent the diversity of the genus include, alongside

462-509: A fibrous cloth used for clothing. Figs have figured prominently in some human cultures. There is evidence that figs, specifically the common fig ( F. carica ) and sycamore fig ( Ficus sycomorus ), were among the first – if not the very first – plant species that were deliberately bred for agriculture in the Middle East, starting more than 11,000 years ago. Nine subfossil F. carica figs dated to about 9400–9200 BCE were found in

528-689: A fresh fig, individual fruit will appear as fleshy "threads", each bearing a single seed inside. The genus Dorstenia , also in the fig family (Moraceae), exhibits similar tiny flowers arranged on a receptacle but in this case the receptacle is a more or less flat, open surface. Fig plants can be monoecious ( hermaphrodite ) or gynodioecious (hermaphrodite and female). Nearly half of fig species are gynodioecious, and therefore have some plants with inflorescences (syconium) with long styled pistillate flowers, and other plants with staminate flowers mixed with short styled pistillate flowers. The long-styled flowers tend to prevent wasps from laying their eggs within

594-443: A key resource for some frugivores including fruit bats , and primates including: capuchin monkeys , langurs , gibbons and mangabeys . They are even more important for birds such as Asian barbets , pigeons , hornbills , fig-parrots and bulbuls , which may almost entirely subsist on figs when these are in plenty. Many Lepidoptera caterpillars feed on fig leaves, for example several Euploea species (crow butterflies),

660-443: A large number of pollen grains, while the stamens are generally long and protrude out of flower. Water-pollinated plants are aquatic and pollen is released into the water. Water currents therefore act as a pollen vector in a similar way to wind currents. Their flowers tend to be small and inconspicuous with many pollen grains and large, feathery stigmas to catch the pollen. However, this is relatively uncommon (only 2% of pollination

726-414: A lot of dilute nectar, secreted during the day. Since birds do not have a strong response to scent, they tend to be odorless. Flowers pollinated by generalist birds are often shorter and wider. Hummingbirds are often associated with pendulous flowers, whereas passerines (perching birds) need a landing platform so flowers and surrounding structures are often more robust. Also, many plants have anthers placed in

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792-408: A plant may be at an advantage if it attracts several species or types of pollinators, ensuring pollen transfer every year. Many species of plants have the back-up option of self-pollination , if they are not self-incompatible. Whilst it is clear that pollination syndromes can be observed in nature, there has been much debate amongst scientists as to how frequent they are and to what extent we can use

858-462: A short and stout peduncle or pedicel; bowl-shaped flowers or inflorescences; copious, sucrose-rich nectar usually produced during the night; tough and wiry styles; an adequate distance between the stigma and nectar to fit the rostrum of the pollinating animal; and potentially a winter–spring flowering period. Many non-flying mammals are nocturnal and have an acute sense of smell, so the plants tend not to have bright showy colours, but instead excrete

924-650: A similar tactic, but in this species it is a specialised petal that acts as a sonar reflector. In the New World bat pollinated flowers often have sulphur -scented compounds. Bat-pollinated plants have bigger pollen than their relatives. The characteristics of the pollination syndrome associated with pollination by mammals which are not bats are: a yeasty odour; cryptic, drab, axillary, geoflorous flowers or inflorescences often obscured from sight; large and sturdy flowers, or grouped together as multi-flowered inflorescences; either sessile flowers or inflorescences or subtended by

990-435: A strong odour. These plants also tend to produce large amounts of pollen because mammals are larger than some other pollinators, and lack the precision smaller pollinators can achieve. The Western-Australian endemic Honey possum ( Tarsipes rostratus ) is an unusual non-flying mammal pollinator in that it has adapted to feeding exclusively on pollen and nectar. It is known to forage on a wide variety of plants (particularly in

1056-448: A suitable place to lay eggs. Without this pollinator service fig trees could not reproduce by seed. In turn, the flowers provide a safe haven and nourishment for the next generation of wasps. This accounts for the frequent presence of wasp larvae in the fruit, and has led to a coevolutionary relationship. Technically, a fig fruit proper would be only one of the many tiny matured, seed-bearing gynoecia found inside one fig – if you cut open

1122-770: A then undiscovered moth with a proboscis whose length was unprecedented at the time. His prediction had gone unverified until 21 years after his death, when the moth was discovered and his conjecture vindicated. The story of its postulated pollinator has come to be seen as one of the celebrated predictions of the theory of evolution . Abiotically pollinated flowers do not attract animal pollinators. Nevertheless, they often have suites of shared traits. Wind-pollinated flowers may be small and inconspicuous, as well as green and not showy. They produce enormous numbers of relatively small pollen grains (hence wind-pollinated plants may be allergens , but seldom are animal-pollinated plants allergenic). Their stigmas may be large and feathery to catch

1188-582: A village in the Larissa regional unit Sykia, Phocis , a village in the municipal unit Lidoriki , Phocis Sykia, Voula , a small doline near Voula in Athens See also [ edit ] Sykias or Pelekita Cave , a cave near Zakros beach, Crete, in Greece Sykies [REDACTED] Topics referred to by the same term This disambiguation page lists articles about distinct geographical locations with

1254-570: A wide range of pollinators, and such ecological generalization is frequently found in nature. A study in Tasmania found the syndromes did not usefully predict the pollinators. A critical re-evaluation of the syndromes suggests that on average about one third of the flowering plants can be classified into the classical syndromes. This reflects the fact that nature is much less predictable and straightforward than 19th-century biologists originally thought. Pollination syndromes can be thought of as extremes of

1320-840: Is a genus of about 850 species of woody trees , shrubs , vines , epiphytes and hemiepiphytes in the family Moraceae . Collectively known as fig trees or figs , they are native throughout the tropics with a few species extending into the semi-warm temperate zone. The common fig ( F. carica ) is a temperate species native to southwest Asia and the Mediterranean region (from Afghanistan to Portugal), which has been widely cultivated from ancient times for its fruit , also referred to as figs. The fruit of most other species are also edible though they are usually of only local economic importance or eaten as bushfood . However, they are extremely important food resources for wildlife. Figs are also of considerable cultural importance throughout

1386-519: Is a sura in Quran named "The Fig" or At-Tin (سوره تین). In Asia, figs are important in Buddhism and Hinduism . In Jainism , the consumption of any fruit belonging to this genus is prohibited. The Buddha is traditionally held to have found bodhi (enlightenment) while meditating for 49 days under a sacred fig . The same species was Ashvattha , the " world tree " of Hinduism. The Plaksa Pra-sravana

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1452-432: Is a gynodioecious plant, as well as lofty fig or clown fig ( F. aspera ), Roxburgh fig ( F. auriculata ), mistletoe fig ( F. deltoidea ), F. pseudopalma , creeping fig ( F. pumila ) and related species. The hermaphrodite common figs are called "inedible figs" or "caprifigs"; in traditional culture in the Mediterranean region they were considered food for goats ( Capra aegagrus ). In

1518-519: Is also one of the traditional crops of Israel , and is included in the list of food found in the Promised Land, according to the Torah ( Deut. 8). Jesus cursed a fig tree for bearing no fruit. The fig tree was sacred in ancient Greece and Cyprus , where it was a symbol of fertility . Video Pollination syndrome The "classical" pollination syndromes were first studied in the 19th century by

1584-425: Is an example of mutualism , in which each organism (fig plant and fig wasp ) benefit each other, in this case reproductively. The intimate association between fig species and their wasp pollinators, along with the high incidence of a one-to-one plant-pollinator ratio have long led scientists to believe that figs and wasps are a clear example of coevolution . Morphological and reproductive behavior evidence, such as

1650-560: Is by far the largest genus in the Moraceae, and is one of the largest genera of flowering plants currently described. The species currently classified within Ficus were originally split into several genera in the mid-1800s, providing the basis for a subgeneric classification when reunited into one genus in 1867. This classification put functionally dioecious species into four subgenera based on floral characters. In 1965, E. J. H. Corner reorganized

1716-411: Is frequently found as a pest on figs grown as potted plants and is spread through the export of these plants to other localities. For a list of other diseases common to fig trees, see List of foliage plant diseases (Moraceae) . Many fig species are grown for their fruits, though only Ficus carica is cultivated to any extent for this purpose. A fig "fruit" is a type of multiple fruit known as

1782-643: Is higher because not all wasp species were detected. On the other hand, species of wasps pollinate multiple host fig species. Molecular techniques, like microsatellite markers and mitochondrial sequence analysis, allowed a discovery of multiple genetically distinct, cryptic wasp species. Not all these cryptic species are sister taxa and thus must have experienced a host fig shift at some point. These cryptic species lacked evidence of genetic introgression or backcrosses indicating limited fitness for hybrids and effective reproductive isolation and speciation . The existence of cryptic species suggests that neither

1848-582: Is hydrophily) and most aquatic plants are insect-pollinated, with flowers that emerge into the air. Vallisneria is an example. Bee -pollinated flowers can be very variable in their size, shape and colouration. They can be open and bowl-shaped (' actinomorphic ', radially symmetrical) or more complex and non-radially symmetric (' zygomorphic '), as is the case with many peas and foxgloves . Some bee flowers tend to be yellow or blue, often with ultraviolet nectar guides and scent. Nectar , pollen, or both are offered as rewards in varying amounts. The sugar in

1914-579: Is produced to fuel the high metabolic rates needed to power their flight. Other moths ( Noctuids , Geometrids , Pyralids , for example) fly slowly and settle on the flower. They do not require as much nectar as the fast-flying hawk moths, and the flowers tend to be small (though they may be aggregated in heads). Myophilous plants, those pollinated by flies, tend not to emit a strong scent, are typically purple, violet, blue, and white, and have open dishes or tubes. Sapromyophilous plants attract flies which normally visit dead animals or dung . Flowers mimic

1980-647: Is used to produce barkcloth in Uganda. Pou ( F. religiosa ) leaves' shape inspired one of the standard kbach rachana , decorative elements in Cambodian architecture. Indian banyan ( F. benghalensis ) and the Indian rubber plant, as well as other species, have use in herbalism . The inner bark of an unknown type of wild fig, locally known as urú , was once used by the Moré people  [ es ] of Bolivia to produce

2046-579: The common fig , whose fingered fig leaf is well known in art and iconography : the weeping fig ( F. benjamina ), a hemiepiphyte with thin, tough leaves on pendulous stalks adapted to its rain forest habitat; the rough-leaved sandpaper figs from Australia; and the creeping fig ( F. pumila ), a vine whose small, hard leaves form a dense carpet of foliage over rocks or garden walls. Moreover, figs with different plant habits have undergone adaptive radiation in different biogeographic regions, leading to very high levels of alpha diversity . In

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2112-441: The plain tiger ( Danaus chrysippus ), the giant swallowtail ( Papilio cresphontes ), the brown awl ( Badamia exclamationis ), and Chrysodeixis eriosoma , Choreutidae and Copromorphidae moths . The citrus long-horned beetle ( Anoplophora chinensis ), for example, has larvae that feed on wood , including that of fig trees; it can become a pest in fig plantations. Similarly, the sweet potato whitefly ( Bemisia tabaci )

2178-450: The Italian botanist Federico Delpino . Although they are useful in understanding of plant-pollinator interactions , sometimes the pollinator of a plant species cannot be accurately predicted from the pollination syndrome alone, and caution must be exerted in making assumptions. The naturalist Charles Darwin surmised that the flower of the orchid Angraecum sesquipedale was pollinated by

2244-459: The ability to echolocate . Bat-pollinated flowers in this part of the world tend to be large and showy, white or light coloured, open at night and have strong musty odours. They are often large balls of stamens. In the Americas pollinating bats are tiny creatures called glossophagines which have both the ability to hover as well as echolocate, and have extremely long tongues. Plants in this part of

2310-449: The bats in a recognisable pattern. A number of species of Marcgravia from Caribbean islands have evolved a special leaf just above the inflorescence to attract bats. The leaf petiole is twisted so the leaf sticks upwards, and the leaf is shaped like a concave disc or dish reflector. The leaf reflects echolocation signals from many directions, guiding the pollinating bats towards the flowers. The epiphytic bean Mucuna holtonii employs

2376-512: The beetle longer. The plant's ovaries are usually well protected from the biting mouthparts of their pollinators. A number of cantharophilous plants are thermogenic , with flowers that can increase their temperature. This heat is thought to help further spread the scent, but the infrared light produced by this heat may also be visible to insects during the dark night, and act as a shining beacon to attract them. Flowers pollinated by specialist nectarivores tend to be large, red or orange tubes with

2442-481: The breba crop is often destroyed by spring frosts. Some parthenocarpic cultivars of common figs do not require pollination at all, and will produce a crop of figs (albeit sterile ) in the absence of caprifigs and fig wasps. Depending on the species, each fruit can contain hundreds or even thousand of seeds. Figs can be propagated by seeds, cuttings, air-layering or grafting. However, as with any plant, figs grown from seed are not necessarily genetically identical to

2508-443: The classical accounts of syndromes, such as the details of anther opening. Although a recent review concluded that there is "overwhelming evidence that functional groups exert different selection pressures on floral traits", the sheer complexity and subtlety of plant-pollinator interactions (and the growing recognition that non-pollinating organisms such as seed predators can affect the evolution of flower traits) means that this debate

2574-573: The classical syndromes to classify plant-pollinator interactions. Although some species of plants are visited only by one type of animal (i.e. they are functionally specialized), many plant species are visited by very different pollinators. For example, a flower may be pollinated by bees, butterflies, and birds. Strict specialization of plants relying on one species of pollinator is relatively rare, probably because it can result in variable reproductive success across years as pollinator populations vary significantly. In such cases, plants should generalize on

2640-439: The correspondence between fig and wasp larvae maturation rates, have been cited as support for this hypothesis for many years. Additionally, recent genetic and molecular dating analyses have shown a very close correspondence in the character evolution and speciation phylogenies of these two clades. According to meta-analysis of molecular data for 119 fig species 35% (41) have multiple pollinator wasp species. The real proportion

2706-540: The early Neolithic village Gilgal I (in the Jordan Valley , 13 km, or 8.1 mi, north of Jericho ). These were a parthenogenetic type and thus apparently an early cultivar. This find predates the first known cultivation of grain in the Middle East by many hundreds of years. Numerous species of fig are found in cultivation in domestic and office environments, including: Fig trees have profoundly influenced culture through several religious traditions. Among

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2772-407: The eastern grasslands of South Africa. These plants do not fit into the classical syndromes, though they may show evidence of convergent evolution in their own right. An analysis of flower traits and visitation in 49 species in the plant genus Penstemon found that it was possible to separate bird- and bee- pollinated species quite well, but only by using floral traits which were not considered in

2838-415: The ecological specialization (i.e. the number of species of pollinators within that functional group). They are responses to common selection pressures exerted by shared pollinators or abiotic pollen vectors, which generate correlations among traits. That is, if two distantly related plant species are both pollinated by nocturnal moths, for example, their flowers will converge on a form which is recognised by

2904-433: The families Proteaceae and Myrtaceae ) including many with typical bird-pollinated flowers such as Calothamnus quadrifidus and many species of Banksia . Pollination syndromes reflect convergent evolution towards forms ( phenotypes ) that limit the number of species of pollinators visiting the plant. They increase the functional specialization of the plant with regard to pollination, though this may not affect

2970-411: The family Agaonidae for pollination. Adult plants vary in size from Ficus benghalensis which may cover a hectare (2.5 acres) or more of ground to Ficus nana of New Guinea which never exceeds one meter (forty inches) in height and width. Specific identification of many of the species can be difficult, but members of the genus Ficus are relatively easy to recognize. Many have aerial roots and

3036-420: The female fig trees, the male flower parts fail to develop; they produce the "'edible figs". Fig wasps grow in common fig caprifigs but not in the female syconiums because the female flower is too long for the wasp to successfully lay her eggs in them. Nonetheless, the wasp pollinates the flower with pollen from the caprifig it grew up in. In many situations, the wasp pollinator is unable to escape and dies within

3102-558: The flower so that pollen rubs against the birds head/back as the bird reaches in for nectar. There are major differences between bat pollination in the New World as opposed to the Old World . In the Old World pollinating bats are large fruit bats of the family Pteropodidae which do not have the ability to hover and must perch in the plant to lap the nectar; these bats furthermore do not have

3168-457: The fruit. When the wasp dies, it is broken down by enzymes ( Ficain ) inside the fig. Fig wasps are not known to transmit any diseases harmful to humans. When a caprifig ripens, another caprifig must be ready to be pollinated. In temperate climes, wasps hibernate in figs, and there are distinct crops. Caprifigs have three crops per year; common figs have two. The first crop ( breba ) is larger and juicier, and usually eaten fresh. In cold climates

3234-421: The genus on the basis of breeding system, uniting these four dioecious subgenera into a single dioecious subgenus Ficus . Monoecious figs were classified within the subgenera Urostigma , Pharmacosycea and Sycomorus . This traditional classification has been called into question by recent phylogenetic studies employing genetic methods to investigate the relationships between representative members of

3300-443: The long tongue of the butterflies. Among the more important moth pollinators are the hawk moths ( Sphingidae ). Their behaviour is similar to hummingbirds : they hover in front of flowers with rapid wingbeats. Most are nocturnal or crepuscular . So moth-pollinated flowers tend to be white, night-opening, large and showy with tubular corollas and a strong, sweet scent produced in the evening, night or early morning. Much nectar

3366-502: The more famous species are the sacred fig tree (Pipal, bodhi, bo, or po, Ficus religiosa ) and other banyan figs such as Ficus benghalensis . The oldest living plant of known planting date is a Ficus religiosa tree known as the Sri Maha Bodhi planted in the temple at Anuradhapura , Sri Lanka by King Tissa in 288 BCE. The common fig is one of two significant trees in Islam , and there

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3432-412: The moths (e.g. pale colour, sweet scent, nectar released at the base of a long tube, night-flowering). These honeybees selectively visit flowers from only one species for a period of time, as can be seen by the colour of the pollen in their baskets. Pollinators fluctuate in abundance and activity independently of their plants, and any one species may fail to pollinate a plant in a particular year. Thus

3498-456: The nectar tends to be sucrose -dominated. A few bees collect oil from special glands on the flower. Butterfly -pollinated flowers tend to be large and showy, pink or lavender in colour, frequently have a landing area, and are usually scented. Since butterflies do not digest pollen (with one exception), more nectar is offered than pollen. The flowers have simple nectar guides with the nectaries usually hidden in narrow tubes or spurs, reached by

3564-400: The number of symbionts nor their evolutionary relationships are necessarily fixed ecologically. While the morphological characteristics that facilitate the fig-wasp mutualisms are likely to be shared more fully in closer relatives, the absence of unique pairings would make it impossible to do a one-to-one tree comparison and difficult to determine cospeciation. With over 800 species, Ficus

3630-486: The odor of such objects. The plant provides them with no reward and they leave quickly unless it has traps to slow them down. Such plants are far less common than myophilous ones. Beetle -pollinated flowers are usually large, greenish or off-white in color and heavily scented. Scents may be spicy, fruity, or similar to decaying organic material. Most beetle-pollinated flowers are flattened or dish shaped, with pollen easily accessible, although they may include traps to keep

3696-464: The other section of Pharmacosycea , the rest of the monoecious species, and all of the dioecious species. These remaining species are divided into two main monophyletic lineages (though the statistical support for these lineages is not as strong as for the monophyly of the more derived clades within them). One consists of all sections of Urostigma except for section Urostigma s. s. . The other includes section Urostigma s. s. , subgenus Sycomorus , and

3762-579: The ovules, while the short-styled flowers are accessible for egg laying. All the native fig trees of the American continent are hermaphrodites, as well as species like Indian banyan ( F. benghalensis ), weeping fig ( F. benjamina ), Indian rubber plant ( F. elastica ), fiddle-leaved fig ( F. lyrata ), Moreton Bay fig ( F. macrophylla ), Chinese banyan ( F. microcarpa ), sacred fig ( F. religiosa ) and sycamore fig ( F. sycomorus ). The common fig ( Ficus carica )

3828-522: The parent and are only propagated this way for breeding purposes. Each species of fig is pollinated by one or a few specialised wasp species, and therefore plantings of fig species outside of their native range results in effectively sterile individuals. For example, in Hawaii , some 60 species of figs have been introduced, but only four of the wasps that fertilize them, so only those species of figs produce viable seeds there and can become invasive species . This

3894-405: The pollen grains. Insects may visit them to collect pollen; in some cases, these are ineffective pollinators and exert little natural selection on the flowers, but there are also examples of ambophilous flowers which are both wind and insect pollinated. Anemophilous, or wind pollinated flowers, are usually small and inconspicuous, and do not possess a scent or produce nectar. The anthers may produce

3960-541: The same name. If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Sykia&oldid=945552088 " Category : Place name disambiguation pages Hidden categories: Short description is different from Wikidata All article disambiguation pages All disambiguation pages Fig tree About 800, see text Ficus ( / ˈ f aɪ k ə s / or / ˈ f iː k ə s / )

4026-839: The species of subgenus Ficus , though the relationships of the sections of these groups to one another are not well resolved. As of April 2024, there are 880 accepted Ficus species according to Plants of the World Online . The following species are typically spreading or climbing lianas : The wood of fig trees is often soft and the latex precludes its use for many purposes. It was used to make mummy caskets in Ancient Egypt . Certain fig species (mainly F. cotinifolia , F. insipida and F. padifolia ) are traditionally used in Mesoamerica to produce papel amate ( Nahuatl : āmatl ). Mutuba ( F. natalensis )

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4092-499: The tropics, Ficus commonly is the most species-rich plant genus in a particular forest. In Asia, as many as 70 or more species can co-exist. Ficus species richness declines with an increase in latitude in both hemispheres. A description of fig tree cultivation is set out in Ibn al-'Awwam 's 12th-century agricultural work titled, Book on Agriculture . Figs are keystone species in many tropical forest ecosystems . Their fruit are

4158-443: The tropics, both as objects of worship and for their many practical uses. Ficus is a pantropical genus of trees, shrubs, and vines occupying a wide variety of ecological niches ; most are evergreen , but some deciduous species are found in areas outside of the tropics and to higher elevations. Fig species are characterized by their unique inflorescence and distinctive pollination syndrome , which uses wasp species belonging to

4224-407: The various sections of each subgenus. Of Corner's original subgeneric divisions of the genus, only Sycomorus is supported as monophyletic in the majority of phylogenetic studies. Notably, there is no clear split between dioecious and monoecious lineages. One of the two sections of Pharmacosycea , a monoecious group, form a monophyletic clade basal to the rest of the genus, which includes

4290-426: The world are often pollinated by both bats and hummingbirds, and have long tubular flowers. Flowers in this part of the world are typically borne away from the trunk or other obstructions, and offer nectar for extended periods of time. In one essay, von Helversen et al . speculate that maybe some bell-shaped flowers have evolved to attract bats in the Americas, as the bell-shape might reflect the sonar pulses emitted by

4356-666: Was said to be a fig tree between the roots of which the Sarasvati River sprang forth; it is usually held to be a sacred fig but more probably is Ficus virens . According to the Kikuyu people , sacrifices to Ngai were performed under a sycomore tree (Mũkũyũ) and if one was not available, a fig tree (Mũgumo) would be used. The common fig tree is cited in the Bible , where in Genesis 3:7, Adam and Eve cover their nakedness with fig leaves. The fig fruit

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