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94-659: Sensation (psychology) refers to the processing of the senses by the sensory system. Sensation or sensations may also refer to: Sensation (psychology) A sense is a biological system used by an organism for sensation , the process of gathering information about the surroundings through the detection of stimuli . Although, in some cultures, five human senses were traditionally identified as such (namely sight , smell , touch , taste , and hearing ), many more are now recognized. Senses used by non-human organisms are even greater in variety and number. During sensation, sense organs collect various stimuli (such as

188-456: A peripheral neuropathy involving peripheral nerves of the somatosensory system. This may present as numbness or paresthesia . Haptic technology can provide touch sensation in virtual and real environments. In the field of speech therapy , tactile feedback can be used to treat speech disorders . Affectionate touch is present in everyday life and can take multiple forms. These actions, however, seem to carry specific functions even though

282-465: A central point. An example would be when we use parentheses in writing. We tend to perceive all of the words in the parentheses as one section instead of individual words within the parentheses. The Law of Continuity tells us that objects are grouped together by their elements and then perceived as a whole. This usually happens when we see overlapping objects. We will see the overlapping objects with no interruptions. The Law of Past Experience refers to

376-403: A group, but we can also perceive three groups of two lines with seven objects in each line. The Law of Closure is the idea that we as humans still see a full picture even if there are gaps within that picture. There could be gaps or parts missing from a section of a shape, but we would still perceive the shape as whole. The Law of Symmetry refers to a person's preference to see symmetry around

470-485: A human), close your eyes (preferably in a dark room) and press gently on the outside corner of one eye through the eyelid. You will see a visual spot toward the inside of your visual field, near your nose.) All stimuli received by the receptors are transduced to an action potential , which is carried along one or more afferent neurons towards a specific area ( cortex ) of the brain . Just as different nerves are dedicated to sensory and motors tasks, different areas of

564-586: A materialistic view of the mind. Some examples of human absolute thresholds for the nine to 21 external senses . Humans respond more strongly to multimodal stimuli compared to the sum of each single modality together, an effect called the superadditive effect of multisensory integration . Neurons that respond to both visual and auditory stimuli have been identified in the superior temporal sulcus . Additionally, multimodal "what" and "where" pathways have been proposed for auditory and tactile stimuli. External receptors that respond to stimuli from outside

658-495: A mechanoreceptor. Photoreceptors convert light (visible electromagnetic radiation ) into signals. Chemical stimuli can be interpreted by a chemoreceptor that interprets chemical stimuli, such as an object's taste or smell, while osmoreceptors respond to a chemical solute concentrations of body fluids. Nociception (pain) interprets the presence of tissue damage, from sensory information from mechano-, chemo-, and thermoreceptors. Another physical stimulus that has its own type of receptor

752-468: A particular color . Visible light is electromagnetic radiation with a wavelength between 380 and 720 nm. Wavelengths of electromagnetic radiation longer than 720 nm fall into the infrared range, whereas wavelengths shorter than 380 nm fall into the ultraviolet range. Light with a wavelength of 380 nm is blue whereas light with a wavelength of 720 nm is dark red . All other colors fall between red and blue at various points along

846-471: A respective visual system (sense of vision), auditory system (sense of hearing), somatosensory system (sense of touch), olfactory system (sense of smell), and gustatory system (sense of taste). Those systems, in turn, contribute to vision , hearing , touch , smell , and the ability to taste . Internal sensation, or interoception, detects stimuli from internal organs and tissues. Many internal sensory and perceptual systems exist in humans, including

940-439: A sense, but it is generally regarded as a cognitive (that is, post-sensory) function of the visual cortex of the brain where patterns and objects in images are recognized and interpreted based on previously learned information. This is called visual memory . The inability to see is called blindness . Blindness may result from damage to the eyeball, especially to the retina, damage to the optic nerve that connects each eye to

1034-498: A series of tiny bones to hair-like fibers in the inner ear , which detect mechanical motion of the fibers within a range of about 20 to 20,000  hertz , with substantial variation between individuals. Hearing at high frequencies declines with an increase in age. Inability to hear is called deafness or hearing impairment. Sound can also be detected as vibrations conducted through the body. Lower frequencies that can be heard are detected this way. Some deaf people are able to determine

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1128-486: A small receptive field (extremely detailed information), they are used in areas like fingertips the most; they are not covered (shelled) and thus respond to pressures over long periods. Tactile corpuscles react to moderate vibration (10–50 Hz) and light touch. They are located in the dermal papillae ; due to their reactivity, they are primarily located in fingertips and lips. They respond in quick action potentials , unlike Merkel nerve endings. They are responsible for

1222-416: A sound or smell) for transduction , meaning transformation into a form that can be understood by the brain. Sensation and perception are fundamental to nearly every aspect of an organism's cognition , behavior and thought . In organisms, a sensory organ consists of a group of interrelated sensory cells that respond to a specific type of physical stimulus. Via cranial and spinal nerves (nerves of

1316-478: A specific type of stimulus. For example, the general sensation and perception of touch, which is known as somatosensation, can be separated into light pressure, deep pressure, vibration, itch, pain, temperature, or hair movement, while the general sensation and perception of taste can be separated into submodalities of sweet , salty , sour , bitter , spicy, and umami , all of which are based on different chemicals binding to sensory neurons . Sensory receptors are

1410-662: A stimulus is constant and unchanging, perceptual sensory adaptation occurs. During that process, the subject becomes less sensitive to the stimulus. Biological auditory (hearing), vestibular and spatial, and visual systems (vision) appear to break down real-world complex stimuli into sine wave components, through the mathematical process called Fourier analysis. Many neurons have a strong preference for certain sine frequency components in contrast to others. The way that simpler sounds and images are encoded during sensation can provide insight into how perception of real-world objects happens. Perception occurs when nerves that lead from

1504-669: A stronger sense of smell than humans. Some animal species lack one or more human sensory system analogues and some have sensory systems that are not found in humans, while others process and interpret the same sensory information in very different ways. For example, some animals are able to detect electrical fields and magnetic fields , air moisture , or polarized light . Others sense and perceive through alternative systems such as echolocation . Recent theory suggests that plants and artificial agents such as robots may be able to detect and interpret environmental information in an analogous manner to animals. Sensory modality refers to

1598-411: Is a molecule called propylthiouracil (PROP) that some humans experience as bitter, some as almost tasteless, while others experience it as somewhere between tasteless and bitter. There is a genetic basis for this difference between perception given the same sensory stimulus. This subjective difference in taste perception has implications for individuals' food preferences, and consequently, health. When

1692-429: Is a perception resulting from activation of neural receptors , generally in the skin including hair follicles , but also in the tongue , throat , and mucosa . A variety of pressure receptors respond to variations in pressure (firm, brushing, sustained, etc.). The touch sense of itching caused by insect bites or allergies involves special itch-specific neurons in the skin and spinal cord. The loss or impairment of

1786-430: Is a type of sensory information that elicits an emotional reaction and is usually social in nature, such as a physical human touch. This type of information is actually coded differently than other sensory information. Intensity of affective touch is still encoded in the primary somatosensory cortex and is processed in a similar way to emotions invoked by sight and sound, as exemplified by the increase of adrenaline caused by

1880-476: Is attached to the tympanic membrane and articulates with the incus. The incus, in turn, articulates with the stapes. The stapes is then attached to the inner ear , where the sound waves will be transduced into a neural signal. The middle ear is connected to the pharynx through the Eustachian tube , which helps equilibrate air pressure across the tympanic membrane. The tube is normally closed but will pop open when

1974-438: Is based on their location relative to the stimuli. An exteroceptor is a receptor that is located near a stimulus of the external environment, such as the somatosensory receptors that are located in the skin. An interoceptor is one that interprets stimuli from internal organs and tissues, such as the receptors that sense the increase in blood pressure in the aorta or carotid sinus . The cells that interpret information about

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2068-453: Is better among adults with smaller index fingertips; this effect of finger size has been shown to underlie the better passive tactile spatial acuity of women, on average, compared to men. The density of tactile corpuscles , a type of mechanoreceptor that detects low-frequency vibrations, is greater in smaller fingers; the same may hold for Merkel cells , which detect the static indentations important for fine spatial acuity. Among children of

2162-489: Is concerned with the nature of perceptual experience and the status of perceptual data , in particular how they relate to beliefs about, or knowledge of, the world. Historical inquiries into the underlying mechanisms of sensation and perception have led early researchers to subscribe to various philosophical interpretations of perception and the mind , including panpsychism , dualism , and materialism . The majority of modern scientists who study sensation and perception take on

2256-431: Is dedicated to the auditory system . The main point is to understand why humans are able to use sound in thinking outside of actually saying it. Relating to auditory cognitive psychology is psychoacoustics . Psychoacoustics is more directed at people interested in music. Haptics , a word used to refer to both taction and kinesthesia, has many parallels with psychoacoustics. Most research around these two are focused on

2350-465: Is described by Steven's power law . Signal detection theory quantifies the experience of the subject to the presentation of a stimulus in the presence of noise . There is internal noise and there is external noise when it comes to signal detection. The internal noise originates from static in the nervous system. For example, an individual with closed eyes in a dark room still sees something—a blotchy pattern of grey with intermittent brighter flashes—this

2444-549: Is encoded or transduced. Multimodality integrates different senses into one unified perceptual experience. For example, information from one sense has the potential to influence how information from another is perceived. Sensation and perception are studied by a variety of related fields, most notably psychophysics , neurobiology , cognitive psychology , and cognitive science . Sensory organs are organs that sense and transduce stimuli. Humans have various sensory organs (i.e. eyes, ears, skin, nose, and mouth) that correspond to

2538-472: Is important amongst some animals. Usually, tactile contact between two animals occurs through stroking, licking, or grooming. These behaviours are essential for the individual's social healthcare, as in the hypothalamus they induce the release of oxytocin, a hormone that decreases stress and anxiety and increases social bonding between animals. More precisely, the consistency of oxytocin neuron activation in rats stroked by humans has been observed, especially in

2632-577: Is in the parietal lobe and its cortex is the primary somatosensory cortex ( Brodmann areas 3, 2 and 1 ) collectively referred to as S1. BA3 receives the densest projections from the thalamus . BA3a is involved with the sense of relative position of neighboring body parts and amount of effort being used during movement. BA3b is responsible for distributing somatosensory information, it projects texture information to BA1 and shape and size information to BA2. Region S2 ( secondary somatosensory cortex ) divides into Area S2 and parietal ventral area. Area S2

2726-410: Is internal noise. External noise is the result of noise in the environment that can interfere with the detection of the stimulus of interest. Noise is only a problem if the magnitude of the noise is large enough to interfere with signal collection. The nervous system calculates a criterion, or an internal threshold, for the detection of a signal in the presence of noise. If a signal is judged to be above

2820-494: Is involved in numerous functions. The somatosensory system is spread through all major parts of the vertebrate body. It consists both of sensory receptors and sensory neurons in the periphery (skin, muscle and organs for example), to deeper neurons within the central nervous system . All afferent touch/vibration information ascends the spinal cord via the dorsal column-medial lemniscus pathway via gracilis (T7 and below) or cuneatus (T6 and above). Cuneatus sends signals to

2914-508: Is involved with specific touch perception and is thus integrally linked with the amygdala and hippocampus to encode and reinforce memories. Parietal ventral area is the somatosensory relay to the premotor cortex and somatosensory memory hub, BA5. BA5 is the topographically organized somato memory field and association area. BA1 processes texture info while BA2 processes size and shape information. Area S2 processes light touch, pain, visceral sensation, and tactile attention. S1 processes

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3008-425: Is often multimodal. Multimodality integrates different senses into one unified perceptual experience. Information from one sense has the potential to influence how information from another is perceived. Multimodal perception is qualitatively different from unimodal perception. There has been a growing body of evidence since the mid-1990s on the neural correlates of multimodal perception. The philosophy of perception

3102-594: Is seen into patterns or groups: Common Fate, Similarity, Proximity, Closure, Symmetry, Continuity, and Past Experience. The Law of Common fate says that objects are led along the smoothest path. People follow the trend of motion as the lines/dots flow. The Law of Similarity refers to the grouping of images or objects that are similar to each other in some aspect. This could be due to shade, colour, size, shape, or other qualities you could distinguish. The Law of Proximity states that our minds like to group based on how close objects are to each other. We may see 42 objects in

3196-428: Is temperature, which is sensed through a thermoreceptor that is either sensitive to temperatures above (heat) or below (cold) normal body temperature. Each sense organ (eyes or nose, for instance) requires a minimal amount of stimulation in order to detect a stimulus. This minimum amount of stimulus is called the absolute threshold. The absolute threshold is defined as the minimum amount of stimulation necessary for

3290-538: Is that received from pain and temperature that is deemed as harmful (noxious). Thermoreceptors relay temperature information in normal circumstances. Nociceptors are specialised receptors for signals of pain. The sense of touch in perceiving the environment uses special sensory receptors in the skin called cutaneous receptors . They include mechanoreceptors such as tactile corpuscles that relay information about pressure and vibration; nociceptors, and thermoreceptors for temperature perception. Stimulation of

3384-409: Is the pathway responsible for the sending of fine touch information to the cerebral cortex of the brain. Crude touch (non-discriminating) is a sensory modality that allows the subject to sense that something has touched them, without being able to localize where they were touched (contrasting "fine touch"). Its fibres are carried in the spinothalamic tract , unlike the fine touch, which is carried in

3478-523: Is the sensation of potentially damaging stimuli. Mechanical, chemical, or thermal stimuli beyond a set threshold will elicit painful sensations. Stressed or damaged tissues release chemicals that activate receptor proteins in the nociceptors. For example, the sensation of heat associated with spicy foods involves capsaicin , the active molecule in hot peppers. Low frequency vibrations are sensed by mechanoreceptors called Merkel cells , also known as type I cutaneous mechanoreceptors. Merkel cells are located in

3572-489: The cell membrane that mediates a physiological change in a neuron, most often through the opening of ion channels or changes in the cell signaling processes. Transmembrane receptors are activated by chemicals called ligands . For example, a molecule in food can serve as a ligand for taste receptors. Other transmembrane proteins, which are not accurately called receptors, are sensitive to mechanical or thermal changes. Physical changes in these proteins increase ion flow across

3666-410: The sensory organs (e.g. eye) to the brain are stimulated, even if that stimulation is unrelated to the target signal of the sensory organ. For example, in the case of the eye, it does not matter whether light or something else stimulates the optic nerve, that stimulation will results in visual perception, even if there was no visual stimulus to begin with. (To prove this point to yourself (and if you are

3760-433: The stratum basale of the epidermis . Deep pressure and vibration is transduced by lamellated ( Pacinian ) corpuscles, which are receptors with encapsulated endings found deep in the dermis, or subcutaneous tissue. Light touch is transduced by the encapsulated endings known as tactile ( Meissner ) corpuscles. Follicles are also wrapped in a plexus of nerve endings known as the hair follicle plexus. These nerve endings detect

3854-594: The vestibular system (sense of balance) sensed by the inner ear and providing the perception of spatial orientation ; proprioception (body position); and nociception (pain). Further internal chemoreception - and osmoreception -based sensory systems lead to various perceptions, such as hunger , thirst , suffocation , and nausea , or different involuntary behaviors, such as vomiting . Nonhuman animals experience sensation and perception, with varying levels of similarity to and difference from humans and other animal species. For example, other mammals in general have

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3948-416: The "blue" cones predominantly. The relative activation of the three different cones is calculated by the brain, which perceives the color as blue. However, cones cannot react to low-intensity light, and rods do not sense the color of light. Therefore, our low-light vision is—in essence—in grayscale . In other words, in a dark room, everything appears as a shade of gray . If you think that you can see colors in

4042-673: The Central and Peripheral nervous systems that relay sensory information to and from the brain and body), the different types of sensory receptor cells (such as mechanoreceptors , photoreceptors , chemoreceptors , thermoreceptors ) in sensory organs transduct sensory information from these organs towards the central nervous system, finally arriving at the sensory cortices in the brain , where sensory signals are processed and interpreted (perceived). Sensory systems, or senses, are often divided into external (exteroception) and internal ( interoception ) sensory systems. Human external senses are based on

4136-618: The ability to feel anything touched is called tactile anesthesia . Paresthesia is a sensation of tingling, pricking, or numbness of the skin that may result from nerve damage and may be permanent or temporary. Two types of somatosensory signals that are transduced by free nerve endings are pain and temperature. These two modalities use thermoreceptors and nociceptors to transduce temperature and pain stimuli, respectively. Temperature receptors are stimulated when local temperatures differ from body temperature . Some thermoreceptors are sensitive to just cold and others to just heat. Nociception

4230-509: The ability to read Braille and feel gentle stimuli. Pacinian corpuscles determine gross touch and distinguish rough and soft substances. They react in quick action potentials, especially to vibrations around 250 Hz (even up to centimeters away). They are the most sensitive to vibrations and have large receptor fields. Pacinian corpuscles react only to sudden stimuli so pressures like clothes that are always compressing their shape are quickly ignored. They have also been implicated in detecting

4324-404: The basis of the type of stimuli they transduce. The different types of functional receptor cell types are mechanoreceptors , photoreceptors , chemoreceptors ( osmoreceptor ), thermoreceptors , electroreceptors (in certain mammals and fish), and nociceptors . Physical stimuli, such as pressure and vibration, as well as the sensation of sound and body position (balance), are interpreted through

4418-415: The basis of three different criteria: cell type , position, and function. Receptors can be classified structurally on the basis of cell type and their position in relation to stimuli they sense. Receptors can further be classified functionally on the basis of the transduction of stimuli, or how the mechanical stimulus, light, or chemical changed the cell membrane potential . One way to classify receptors

4512-452: The body are called exteroceptors . Human external sensation is based on the sensory organs of the eyes , ears , skin , vestibular system , nose , and mouth , which contribute, respectively, to the sensory perceptions of vision , hearing , touch , balance , smell , and taste . Smell and taste are both responsible for identifying molecules and thus both are types of chemoreceptors . Both olfaction (smell) and gustation (taste) require

4606-546: The brain (cortices) are similarly dedicated to different sensory and perceptual tasks. More complex processing is accomplished across primary cortical regions that spread beyond the primary cortices. Every nerve, sensory or motor , has its own signal transmission speed. For example, nerves in the frog's legs have a 90 ft/s (99 km/h) signal transmission speed, while sensory nerves in humans, transmit sensory information at speeds between 165 ft/s (181 km/h) and 330 ft/s (362 km/h). Perceptual experience

4700-515: The brain, and/or from stroke ( infarcts in the brain). Temporary or permanent blindness can be caused by poisons or medications. People who are blind from degradation or damage to the visual cortex, but still have functional eyes, are actually capable of some level of vision and reaction to visual stimuli but not a conscious perception; this is known as blindsight . People with blindsight are usually not aware that they are reacting to visual sources, and instead just unconsciously adapt their behavior to

4794-408: The caudal paraventricular nucleus. It was found that this affiliative relationship induced by tactile contact is common no matter the relationship between the two individuals (mother-infant, male-female, human-animal). It has also been discovered that the level of oxytocin release through this behaviour correlates with the time course of social interaction as longer stroking induced a greater release of

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4888-432: The cells or structures that detect sensations. Stimuli in the environment activate specialized receptor cells in the peripheral nervous system . During transduction, physical stimulus is converted into action potential by receptors and transmitted towards the central nervous system for processing. Different types of stimuli are sensed by different types of receptor cells . Receptor cells can be classified into types on

4982-551: The cochlear nucleus indirectly via spinal grey matter, this info is used in determining if a perceived sound is just villi noise/irritation. All fibers cross (left becomes right) in the medulla. A somatosensory pathway will typically have three neurons: first-order, second-order, and third-order. Photoreceptors, similar to those found in the retina of the eye , detect potentially damaging ultraviolet radiation ( ultraviolet A specifically), inducing increased production of melanin by melanocytes . Thus tanning potentially offers

5076-524: The criterion, thus the signal is differentiated from the noise, the signal is sensed and perceived. Errors in signal detection can potentially lead to false positives and false negatives . The sensory criterion might be shifted based on the importance of the detecting the signal. Shifting of the criterion may influence the likelihood of false positives and false negatives. Subjective visual and auditory experiences appear to be similar across humans subjects. The same cannot be said about taste. For example, there

5170-453: The dark, it is most likely because your brain knows what color something is and is relying on that memory. There is some disagreement as to whether the visual system consists of one, two, or three submodalities. Neuroanatomists generally regard it as two submodalities, given that different receptors are responsible for the perception of color and brightness. Some argue that stereopsis , the perception of depth using both eyes, also constitutes

5264-401: The dermis of the skin are examples of neurons that have free nerve endings (1). Also located in the dermis of the skin are lamellated corpuscles , neurons with encapsulated nerve endings that respond to pressure and touch (2). The cells in the retina that respond to light stimuli are an example of a specialized receptor (3), a photoreceptor . A transmembrane protein receptor is a protein in

5358-425: The detection of a stimulus 50% of the time. Absolute threshold is measured by using a method called signal detection . This process involves presenting stimuli of varying intensities to a subject in order to determine the level at which the subject can reliably detect stimulation in a given sense. Differential threshold or just noticeable difference (JDS) is the smallest detectable difference between two stimuli, or

5452-459: The direction and location of vibrations picked up through the feet. Studies pertaining to audition started to increase in number towards the latter end of the nineteenth century. During this time, many laboratories in the United States began to create new models, diagrams, and instruments that all pertained to the ear. Auditory cognitive psychology is a branch of cognitive psychology that

5546-420: The dorsal column. As fine touch normally works in parallel to crude touch, a person will be able to localize touch until fibres carrying fine touch (in the dorsal column–medial lemniscus pathway) have been disrupted. Then the subject will feel the touch, but be unable to identify where they were touched. The somatosensory cortex encodes incoming sensory information from receptors all over the body. Affective touch

5640-462: The environment can be either (1) a neuron that has a free nerve ending , with dendrites embedded in tissue that would receive a sensation; (2) a neuron that has an encapsulated ending in which the sensory nerve endings are encapsulated in connective tissue that enhances their sensitivity; or (3) a specialized receptor cell , which has distinct structural components that interpret a specific type of stimulus. The pain and temperature receptors in

5734-445: The environment that affect receptor cell membrane potentials. Other stimuli include the electromagnetic radiation from visible light. For humans, the only electromagnetic energy that is perceived by our eyes is visible light. Some other organisms have receptors that humans lack, such as the heat sensors of snakes, the ultraviolet light sensors of bees, or magnetic receptors in migratory birds. Receptor cells can be further categorized on

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5828-476: The evaluation of risks that occur during conducting such behaviours in the wild life, and further research is required to unveil the connection between tactile caring and fitness level. Studies show a correlation between touching a soft or hard object and how a person thinks or even makes decisions. Further, between the firmness of a touch and the evoking of gender stereotyping. Tactile memories as part of haptic memory , are organized somatotopically , following

5922-468: The evolutionary benefit from such a wide range of behaviours is not entirely understood. Researchers investigated the expression patterns and characteristics of 8 different affectionate touch actions - embracing, holding, kissing, leaning, petting, squeezing, stroking, and tickling - in a self-report study. It was found that the affectionate touch has distinct target areas on the body, different associated affect, comfort-value, and expression frequency based on

6016-402: The hormone. The importance of somatosensory stimulation in social animals such as primates has also been observed. Grooming is part of the social interaction primates exert on their conspecifics. This interaction is required between individuals to maintain the affiliative relationship within the group, avoid internal conflict and increase group bonding. However, such social interaction requires

6110-481: The instrument, the listener, and the player of the instrument. Somatosensation is considered a general sense, as opposed to the special senses discussed in this section. Somatosensation is the group of sensory modalities that are associated with touch and interoception. The modalities of somatosensation include pressure , vibration , light touch, tickle , itch , temperature , pain , kinesthesia . Somatosensation , also called tactition (adjectival form: tactile)

6204-512: The lateral aspect of the head is known as the auricle . At the end of the auditory canal is the tympanic membrane, or ear drum , which vibrates after it is struck by sound waves. The auricle, ear canal, and tympanic membrane are often referred to as the external ear . The middle ear consists of a space spanned by three small bones called the ossicles . The three ossicles are the malleus , incus , and stapes , which are Latin names that roughly translate to hammer, anvil, and stirrup. The malleus

6298-428: The location of objects which are sensed by the visual system (which provides confirmation of the place of those objects relative to the body), as input to the mechanical reflexes of the body. Fine touch (or discriminative touch) is a sensory modality that allows a subject to sense and localize touch. The form of touch where localization is not possible is known as crude touch. The dorsal column–medial lemniscus pathway

6392-509: The location of touch sensations on handheld tools. Bulbous corpuscles react slowly and respond to sustained skin stretch. They are responsible for the feeling of object slippage and play a major role in the kinesthetic sense and control of finger position and movement. Merkel and bulbous cells - slow-response - are myelinated ; the rest - fast-response - are not. All of these receptors are activated upon pressures that distort their shape causing an action potential. The postcentral gyrus

6486-438: The membrane, and can generate an action potential or a graded potential in the sensory neurons . A third classification of receptors is by how the receptor transduces stimuli into membrane potential changes. Stimuli are of three general types. Some stimuli are ions and macromolecules that affect transmembrane receptor proteins when these chemicals diffuse across the cell membrane. Some stimuli are physical variations in

6580-408: The molecular level, visual stimuli cause changes in the photopigment molecule that lead to changes in membrane potential of the photoreceptor cell. A single unit of light is called a photon , which is described in physics as a packet of energy with properties of both a particle and a wave. The energy of a photon is represented by its wavelength , with each wavelength of visible light corresponding to

6674-466: The most rigorous being the grating orientation task. In this task subjects identify the orientation of a grooved surface presented in two different orientations, which can be applied manually or with automated equipment. Many studies have shown a decline in passive tactile spatial acuity with age; the reasons for this decline are unknown, but may include loss of tactile receptors during normal aging. Remarkably, index finger passive tactile spatial acuity

6768-437: The movement of hair at the surface of the skin, such as when an insect may be walking along the skin . Stretching of the skin is transduced by stretch receptors known as bulbous corpuscles . Bulbous corpuscles are also known as Ruffini corpuscles, or type II cutaneous mechanoreceptors. The heat receptors are sensitive to infrared radiation and can occur in specialized organs, for instance in pit vipers . The thermoceptors in

6862-403: The muscles of the pharynx contract during swallowing or yawning . Mechanoreceptors turn motion into electrical nerve pulses, which are located in the inner ear. Since sound is vibration, propagating through a medium such as air, the detection of these vibrations, that is the sense of the hearing, is a mechanical sense because these vibrations are mechanically conducted from the eardrum through

6956-416: The organization of the somatosensory cortex. A variety of studies have measured and investigated the causes for differences between individuals in the sense of fine touch. One well-studied area is passive tactile spatial acuity, the ability to resolve the fine spatial details of an object pressed against the stationary skin. A variety of methods have been used to measure passive tactile spatial acuity, perhaps

7050-418: The physical sensation to its related conceptual processing. Indeed, it was found that different physical properties - weight, texture, and hardness - of a touched object can influence social judgement and decision-making. For example, participants described a passage of a social interaction to be harsher when they touched a hard wooden block instead of a soft blanket prior to the task. Building on these findings,

7144-580: The physical stimulus is preserved. In this way, neighboring neurons in the somatosensory cortex represent nearby locations on the skin or in the body, creating a map or sensory homunculus . Tactile signing is a common means of communication used by people with deafblindness . It is based on a sign language or another system of manual communication. Humans can communicate specific emotions through touch alone including anger, fear, disgust, love, gratitude, and sympathy via touch at much-better-than-chance levels. The two different types of mechanoreceptor in

7238-506: The prefrontal cortex is highly correlated with pleasantness scores of an affective touch. Inhibitory transcranial magnetic stimulation (TMS) of the primary somatosensory cortex inhibits the perception of affective touch intensity, but not affective touch pleasantness. Therefore, the S1 is not directly involved in processing socially affective touch pleasantness, but still plays a role in discriminating touch location and intensity. Tactile interaction

7332-420: The proprioceptors in the skin, muscles, and joints. The receptor for the sense of balance resides in the vestibular system in the ear (for the three-dimensional orientation of the head, and by inference, the rest of the body). Balance is also mediated by the kinesthetic reflex fed by proprioception (which senses the relative location of the rest of the body to the head). In addition, proprioception estimates

7426-414: The receptors activate peripheral sensory neurons that convey signals to the spinal cord that may drive a responsive reflex , and may also be conveyed to the brain for conscious perception. Somatosensory information from the face and head enter the brain via cranial nerves such as the trigeminal nerve . The neural pathways that go to the brain are structured such that information about the location of

7520-505: The recognition of every member in the group. As such, it has been observed that the size of the neocortex is positively correlated with the size of the group, reflecting a limit to the number of recognizable members amongst which grooming can occur. Furthermore, the time course of grooming is related to vulnerability due to predation to which animals are exposed to whilst performing such social interaction. The relationship between tactile interaction, stress reduction and social bonding depends on

7614-411: The remaining info (crude touch, pain, temperature). BA7 integrates visual and proprioceptive info to locate objects in space. The insular cortex (insula) plays a role in the sense of bodily-ownership, bodily self-awareness, and perception. Insula also plays a role in conveying info about sensual touch, pain, temperature, itch, and local oxygen status. Insula is a highly connected relay and thus

7708-531: The same age, those with smaller fingers also tend to have better tactile acuity. Many studies have shown that passive tactile spatial acuity is enhanced among blind individuals compared to sighted individuals of the same age, possibly because of cross modal plasticity in the cerebral cortex of blind individuals. Perhaps also due to cortical plasticity, individuals who have been blind since birth reportedly consolidate tactile information more rapidly than sighted people. A somatosensory deficiency may be caused by

7802-714: The sensory organs of the eyes , ears , skin , nose , mouth and the vestibular system . Internal sensation detects stimuli from internal organs and tissues. Internal senses possessed by humans include spatial orientation , proprioception (body position) and nociception (pain). Further internal senses lead to signals such as hunger , thirst , suffocation , and nausea , or different involuntary behaviors, such as vomiting . Some animals are able to detect electrical and magnetic fields , air moisture , or polarized light , while others sense and perceive through alternative systems, such as echolocation . Sensory modalities or sub modalities are different ways sensory information

7896-491: The skin are quite different from the homeostatic thermoceptors in the brain ( hypothalamus ), which provide feedback on internal body temperature. Somatosensory system Mechanosensory information includes that of light touch, vibration, pressure and tension in the skin. Much of this information belongs to the sense of touch which is a general somatic sense in contrast to the special senses of sight , smell , taste , hearing and balance . Nociceptory information

7990-504: The skin are termed low-threshold mechanoreceptors , and high threshold mechanoreceptors . The four mechanoreceptors in glabrous skin are low-threshold that respond to harmless stimuli. They are innervated by four different afferent fibers. High-threshold mechanoreceptors, respond to harmful stimuli . Merkel cell nerve endings are found in the basal epidermis and hair follicles ; they react to low vibrations (5–15  Hz ) and deep static touch such as shapes and edges. Due to having

8084-409: The skin rapid protection from DNA damage and sunburn caused by ultraviolet radiation (DNA damage caused by ultraviolet B ). However, whether this offers protection is debatable, because the amount of melanin released by this process is modest in comparison to the amounts released in response to DNA damage caused by ultraviolet B radiation. The tactile feedback from proprioception is derived from

8178-482: The smallest difference in stimuli that can be judged to be different from each other. Weber's Law is an empirical law that states that the difference threshold is a constant fraction of the comparison stimulus. According to Weber's Law, bigger stimuli require larger differences to be noticed. Magnitude estimation is a psychophysical method in which subjects assign perceived values of given stimuli. The relationship between stimulus intensity and perceptive intensity

8272-419: The social touch of a loved one, as opposed to the physical inability to touch someone you do not love. Meanwhile, the feeling of pleasantness associated with affective touch activates the anterior cingulate cortex more than the primary somatosensory cortex. Functional magnetic resonance imaging (fMRI) data shows that increased blood-oxygen-level contrast (BOLD) signal in the anterior cingulate cortex as well as

8366-465: The stimulus. On February 14, 2013, researchers developed a neural implant that gives rats the ability to sense infrared light which for the first time provides living creatures with new abilities, instead of simply replacing or augmenting existing abilities. According to Gestalt Psychology, people perceive the whole of something even if it is not there. The Gestalt's Law of Organization states that people have seven factors that help to group what

8460-449: The tendency humans have to categorize objects according to past experiences under certain circumstances. If two objects are usually perceived together or within close proximity of each other the Law of Past Experience is usually seen. Hearing, or audition, is the transduction of sound waves into a neural signal that is made possible by the structures of the ear . The large, fleshy structure on

8554-587: The transduction of chemical stimuli into electrical potentials. The visual system, or sense of sight, is based on the transduction of light stimuli received through the eyes and contributes to visual perception . The visual system detects light on photoreceptors in the retina of each eye that generates electrical nerve impulses for the perception of varying colors and brightness. There are two types of photoreceptors: rods and cones . Rods are very sensitive to light but do not distinguish colors. Cones distinguish colors but are less sensitive to dim light. At

8648-466: The type of touch action that is performed. Besides the rather obvious sensory consequences of touch, it can also affect higher-level aspects of cognition such as social judgements and decision-making. This effect might arise due to a physical-to-mental scaffolding process in early development, whereby sensorimotor experiences are linked to the emergence of conceptual knowledge. Such links might be maintained throughout life, and so touching an object may cue

8742-416: The wavelength scale. The three types of cone opsins , being sensitive to different wavelengths of light, provide us with color vision. By comparing the activity of the three different cones, the brain can extract color information from visual stimuli. For example, a bright blue light that has a wavelength of approximately 450 nm would activate the "red" cones minimally, the "green" cones marginally, and

8836-413: The way that information is encoded, which is similar to the idea of transduction . The main sensory modalities can be described on the basis of how each is transduced. Listing all the different sensory modalities, which can number as many as 17, involves separating the major senses into more specific categories, or submodalities, of the larger sense. An individual sensory modality represents the sensation of

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