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111-412: The lion is a common charge in heraldry . It traditionally symbolises courage , nobility , royalty , strength, stateliness and valour , because historically the lion has been regarded as the "king of beasts". The lion also carries Judeo-Christian symbolism. The Lion of Judah stands in the coat of arms of Jerusalem . Similar-looking lions can be found elsewhere, such as in the coat of arms of

222-487: A Lion, and blazon him by his true name. A lion walking and looking about him, the early Heralds held to be acting the part of a leopard: consequently, when he was in any such attitude, they blazoned him as "a leopard". The animal bearing that name bore it simply as an heraldic title, which distinguished a Lion in a particular attitude. These heraldic "leopards" were drawn in every respect as other heraldic "lions", without spots or any leopardish distinction whatever. This explains

333-453: A blue shield decorated six golden lions rampant and wearing a blue helmet adorned with another lion. A chronicle dated to c. 1175 states that Geoffrey was given a shield of this description when he was knighted by his father-in-law, Henry I , in 1128. Earlier heraldic writers attributed the lions of England to William the Conqueror , but the earliest evidence of the association of lions with

444-417: A buildup of scent attracting the attention of predators, the lioness moves her cubs to a new den site several times a month, carrying them one-by-one by the nape of the neck. Usually, the mother does not integrate herself and her cubs back into the pride until the cubs are six to eight weeks old. Sometimes the introduction to pride life occurs earlier, particularly if other lionesses have given birth at about

555-540: A closer relationship with P. leo melanochaita . Lion samples from some parts of the Ethiopian Highlands cluster genetically with those from Cameroon and Chad, while lions from other areas of Ethiopia cluster with samples from East Africa. Researchers, therefore, assume Ethiopia is a contact zone between the two subspecies. Genome -wide data of a wild-born historical lion sample from Sudan showed that it clustered with P. l. leo in mtDNA-based phylogenies, but with

666-471: A constant lookout for circling vultures, which indicate the death or distress of an animal. Most carrion on which both hyenas and lions feed upon are killed by hyenas rather than lions. Carrion is thought to provide a large part of lion diet. Lions and spotted hyenas occupy a similar ecological niche and compete for prey and carrion; a review of data across several studies indicates a dietary overlap of 58.6%. Lions typically ignore hyenas unless they are on

777-457: A different color, it is said to be armed of that tincture, and if the tongue is a different color it is langued of that tincture. For example, the blazon of the monarch of Scotland is Or, a lion gules, within a double tressure flory counter-flory of the same, armed and langued azure. The arms of the Cinque Ports depict lions dimidiated with the hulks of ships, incorporating the front half of

888-425: A favoured position in the group, either stalking prey on the "wing", then attacking, or moving a smaller distance in the centre of the group and capturing prey fleeing from other lionesses. Males attached to prides do not usually participate in group hunting. Some evidence suggests, however, that males are just as successful as females; they are typically solo hunters who ambush prey in small bushland. They may join in

999-470: A few reported cases of old and wounded lions falling prey to wild dogs. Most lionesses reproduce by the time they are four years of age. Lions do not mate at a specific time of year and the females are polyestrous . Like those of other cats, the male lion's penis has spines that point backward. During withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation . A lioness may mate with more than one male when she

1110-548: A heavier mane. On average, Asiatic lions have sparser manes than African lions. This feature likely evolved to signal the fitness of males to females. Males with darker manes appear to have greater reproductive success and are more likely to remain in a pride for longer. They have longer and thicker hair and higher testosterone levels, but they are also more vulnerable to heat stress. The core body temperature does apparently not increase regardless of sex, season, feeding time, length and colour of mane, but only surface temperature

1221-590: A high affinity to P. l. melanochaita . This result suggested that the taxonomic position of lions in Central Africa may require revision. Other lion subspecies or sister species to the modern lion existed in prehistoric times: The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago . Results of analyses differ in

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1332-473: A key species sought for exhibition in zoological gardens across the world since the late 18th century. Cultural depictions of lions were prominent in Ancient Egypt , and depictions have occurred in virtually all ancient and medieval cultures in the lion's historic and current range. The English word lion is derived via Anglo-Norman liun from Latin leōnem (nominative: leō ), which in turn

1443-495: A kill or are being harassed, while the latter tend to visibly react to the presence of lions with or without the presence of food. In the Ngorongoro crater , lions subsist largely on kills stolen from hyenas, causing them to increase their kill rate. In Botswana's Chobe National Park, the situation is reversed as hyenas there frequently challenge lions and steal their kills, obtaining food from 63% of all lion kills. When confronted on

1554-505: A kill, hyenas may either leave or wait patiently at a distance of 30–100 m (98–328 ft) until the lions have finished. Hyenas may feed alongside lions and force them off a kill. The two species attack one another even when there is no food involved for no apparent reason. Lions can account for up to 71% of hyena deaths in Etosha National Park . Hyenas have adapted by frequently mobbing lions that enter their home ranges. When

1665-519: A kill. Crippled lions and cubs may fall victim to hyenas and leopards or be trampled by buffalo or elephants. Careless lions may be maimed when hunting prey. Nile crocodiles may also kill and eat lions, evidenced by the occasional lion claw found in crocodile stomachs. Ticks commonly infest the ears, neck and groin regions of the lions. Adult forms of several tapeworm species of the genus Taenia have been isolated from lion intestines, having been ingested as larvae in antelope meat. Lions in

1776-558: A lion makes when sniffing chemical signals and involves an open mouth with bared teeth, raised muzzle, wrinkled nose, closed eyes and relaxed ears. Lions also use chemical and visual marking; males spray urine and scrape plots of ground and objects within the territory. The lion's repertoire of vocalisations is large; variations in intensity and pitch appear to be central to communication. Most lion vocalisations are variations of growling , snarling , meowing and roaring. Other sounds produced include puffing, bleating and humming. Roaring

1887-523: A lion rampant guardant (i.e., upright like a lion with its head turned to full face like a leopard) and a Lion léopardé as a lion passant (i.e., walking like a leopard with its head facing dexter like a lion). German-American heraldist Carl-Alexander von Volborth agrees with Rietstap's translations, in contrast to those of Fox-Davies as stated above. As if to clarify the situation, English heraldist Hugh Clark wrote in his Introduction to Heraldry (1829): The true heraldic lion, according to French authors,

1998-495: A lion's whole body is turned to face right, he is to sinister or contourné . If his whole body faces the viewer, he is affronté . If his head only faces the viewer he is guardant or gardant , and if he looks back over his shoulder he is regardant . These adjectives follow any other adjectives of position. A lion (or other beast) coward carries the tail between its hind legs. The tail also may be knotted ( nowed ), forked ( queue fourchée ), doubled ( double-queued ; as in

2109-473: A longer evolutionary history than genetically less diverse lion samples from Asia and West and Central Africa. A whole genome-wide sequence of lion samples showed that samples from West Africa shared alleles with samples from Southern Africa, and samples from Central Africa shared alleles with samples from Asia. This phenomenon indicates that Central Africa was a melting pot of lion populations after they had become isolated, possibly migrating through corridors in

2220-399: A loose pride while females associate with up to 12 other females, forming a stronger pride together with their cubs. Female and male lions associate only when mating. Coalitions of males hold territory for a longer time than single lions. Males in coalitions of three or four individuals exhibit a pronounced hierarchy, in which one male dominates the others and mates more frequently. The lion

2331-491: A lympago, possesses a human face. Both lions and leopards may have been among the earliest beasts to appear in heraldry. The Oxford Guide to Heraldry notes that the earliest English treatise on heraldry, a late-13th or early-14th century Anglo-Norman manuscript titled De Heraudrie , mentions the crow, eagle, griffin, heron, leopard, lion, martlet, popinjay, and swan. Citing Bado Aureo, the Oxford Guide further suggests that

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2442-433: A nomad is a "range". Males associated with a pride patrol the fringes. Both males and females defend the pride against intruders, but the male lion is better-suited for this purpose due to its stockier, more powerful build. Some individuals consistently lead the defense against intruders, while others lag behind. Lions tend to assume specific roles in the pride; slower-moving individuals may provide other valuable services to

2553-544: A nomadic phase before gaining residence in a pride. A study undertaken in the Serengeti National Park revealed that nomadic coalitions gain residency at between 3.5 and 7.3 years of age. In Kruger National Park, dispersing male lions move more than 25 km (16 mi) away from their natal pride in search of their own territory. Female lions stay closer to their natal pride. Therefore, female lions in an area are more closely related to each other than male lions in

2664-541: A number of coats of arms with lions, most of them of ministeriales of the House of Habsburg. The lion in the coat of arms of Bohemia is depicted with two tails ( à la queue fourchée ). According to Ménestrier , this is due to a jest made by Emperor Frederick , who granted Vladislaus II, Duke of Bohemia a coat of arms with a lion coué , that is, with its tail between its legs. Vladislaus' men refused to follow this emblem, calling it an ape, so that Frederick agreed to improve

2775-425: A reed-bed, a cave, or some other sheltered area, usually away from the pride. She will often hunt alone while the cubs are still helpless, staying relatively close to the den. Lion cubs are born blind, their eyes opening around seven days after birth. They weigh 1.2–2.1 kg (2.6–4.6 lb) at birth and are almost helpless, beginning to crawl a day or two after birth and walking around three weeks of age. To avoid

2886-565: A short, rounded head, a reduced neck, and round ears; males have broader heads. The fur varies in colour from light buff to silvery grey, yellowish red, and dark brown. The colours of the underparts are generally lighter. A new-born lion has dark spots , which fade as the cub reaches adulthood, although faint spots may still be seen on the legs and underparts. The tail of all lions ends in a dark, hairy tuft that, in some lions, conceals an approximately 5 mm (0.20 in)-long, hard "spine" or "spur" composed of dermal papillae. The functions of

2997-415: A short, rounded head; round ears; and a dark, hairy tuft at the tip of its tail. It is sexually dimorphic ; adult male lions are larger than females and have a prominent mane. It is a social species , forming groups called prides. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on medium-sized and large ungulates . The lion

3108-429: A year old and begin to hunt effectively when nearing the age of two. Single lions are capable of bringing down zebra and wildebeest, while larger prey like buffalo and giraffe are riskier. In Chobe National Park , large prides have been observed hunting African bush elephants up to around 15 years old in exceptional cases, with the victims being calves, juveniles, and even subadults. In typical hunts, each lioness has

3219-404: Is in heat . Lions of both sexes may be involved in group homosexual and courtship activities. Males will also head-rub and roll around with each other before mounting each other. Generation length of the lion is about seven years. The average gestation period is around 110   days; the female gives birth to a litter of between one and four cubs in a secluded den, which may be a thicket,

3330-905: Is a generalist hypercarnivore and is considered to be both an apex and keystone predator due to its wide prey spectrum. Its prey consists mainly of medium-sized to large ungulates , particularly blue wildebeest , plains zebra , African buffalo , gemsbok and giraffe . It also frequently takes common warthog despite it being much smaller. In India, chital and sambar deer are the most common wild prey, while livestock contributes significantly to lion kills outside protected areas. It usually avoids fully grown adult elephants , rhinoceros and hippopotamus and small prey like dik-dik , hyraxes , hares and monkeys . Unusual prey include porcupines and small reptiles. Lions kill other predators but seldom consume them. Young lions first display stalking behaviour at around three months of age, although they do not participate in hunting until they are almost

3441-538: Is a French term for what the English call a Lion passant gardant . The word leopard is always made use of by the French heralds to express in their language, a lion full-faced, or gardant . Thus, when a lion is placed on an escutcheon in that attitude which we call rampant gardant , the French blazon it a Lion Leopardé . When he is passant only, they call him leopard lioné . English heraldist Charles Boutell wrote in 1890 that

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3552-543: Is a mixture of dry savannah forest and very dry, deciduous scrub forest . In Africa, the range of the lion originally spanned most of the central African rainforest zone and the Sahara desert. In the 1960s, it became extinct in North Africa, except in the southern part of Sudan. During the mid- Holocene , around 8,000-6,000 years ago, the range of lions expanded into Southeastern and Eastern Europe, partially re-occupying

3663-924: Is affected. Unlike in other felid species, female lions consistently interact with multiple males at once. Another hypothesis suggests that the mane also serves to protect the neck in fights, but this is disputed. During fights, including those involving maneless females and adolescents, the neck is not targeted as much as the face, back, and hindquarters. Injured lions also begin to lose their manes. Almost all male lions in Pendjari National Park are either maneless or have very short manes. Maneless lions have also been reported in Senegal, in Sudan 's Dinder National Park and in Tsavo East National Park , Kenya. Castrated lions often have little to no mane because

3774-503: Is always to be represented in profile, or, as the ancient heralds say, showing but one eye and one ear. His attitude, also, should always be rampant or ravaging. When passant and full-faced, they blazoned him a leopard, vide Lion Leopardé : in England, however, the lions in the royal and other achievements have always been blazoned as lions, however depicted since the time of Henry III, in whose reign they were called "Leopards". Lion Leopardé ...

3885-465: Is an apex and keystone predator ; although some lions scavenge when opportunities occur and have been known to hunt humans , lions typically do not actively seek out and prey on humans. The lion inhabits grasslands , savannahs , and shrublands . It is usually more diurnal than other wild cats, but when persecuted, it adapts to being active at night and at twilight . During the Neolithic period,

3996-508: Is commonly referred to as the Lion of Saint Mark , although Arthur Fox-Davies defined as a Lion of Saint Mark one present in a specifically religious context and depicted with a halo . The winged lion is the traditional symbol of Venice , whose patron saint is Mark the Evangelist. A sea-lion, also called a morse, is depicted with the tail of a fish replacing its hindquarters, hind legs and tail. It

4107-417: Is described as naiant when depicted horizontally, and as resurgent when rising from water. They typically appear as supporters, but are also known as charges and crests. The lion-dragon is a lion with the lower body, hind legs, wings and tail of a wyvern , although Fox-Davies doubted the existence of this figure outside of heraldry books and reported not to know of any actual use of it. The man-lion, also called

4218-399: Is generally mutual and the recipient appears to express pleasure. The head and neck are the most common parts of the body licked; this behaviour may have arisen out of utility because lions cannot lick these areas themselves. Lions have an array of facial expressions and body postures that serve as visual gestures. A common facial expression is the "grimace face" or flehmen response , which

4329-521: Is no evidence for gene flow between the two lineages, indicating that they did not share the same geographic area. The Eurasian and American cave lions became extinct at the end of the last glacial period without mitochondrial descendants on other continents. The modern lion was probably widely distributed in Africa during the Middle Pleistocene and started to diverge in sub-Saharan Africa during

4440-415: Is short and powerful; it attempts to catch prey with a fast rush and final leap, usually pulls it down by the rump, and kills with a clamping bite to the throat or muzzle . It can hold the prey's throat for up to 13 minutes, until the prey stops moving. It has a bite force from 1593.8 to 1768 Newtons at the canine tip and up 4167.6 Newtons at the carnassial notch. Lions typically consume prey at

4551-494: Is termed a léopard is always guardant (head turned toward the observer), thus the modern English heraldic terms "lion passant guardant", "lion passant", and "lion rampant guardant" correlate to the Old French terms léopard , lion léopardé , and léopard lionné , respectively. A small group of examples is depicted listed below. Western depictions Heraldic lions have also found their way onto municipal or county seals in

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4662-586: Is the Tsavo lion pride that always has just one adult male. Prides act as fission–fusion societies , and members will split into subgroups that keep in contact with roars . Nomadic lions range widely and move around sporadically, either in pairs or alone. Pairs are more frequent among related males. A lion may switch lifestyles; nomads can become residents and vice versa. Interactions between prides and nomads tend to be hostile, although pride females in estrus allow nomadic males to approach them. Males spend years in

4773-615: Is the most social of all wild felid species, living in groups of related individuals with their offspring. Such a group is called a " pride ". Groups of male lions are called "coalitions". Females form the stable social unit in a pride and do not tolerate outside females. The majority of females remain in their birth prides while all males and some females will disperse . The average pride consists of around 15 lions, including several adult females and up to four males and their cubs of both sexes. Large prides, consisting of up to 30 individuals, have been observed. The sole exception to this pattern

4884-475: Is typically brownish and tinged with yellow, rust, and black hairs. Mutations in the genes microphthalmia-associated transcription factor and tyrosinase are possibly responsible for the colour of manes. It starts growing when lions enter adolescence, when testosterone levels increase, and reach their full size at around four years old. Cool ambient temperatures in European and North American zoos may result in

4995-426: Is unable to consume all of the kill, it rests for a few hours before continuing to eat. On hot days, the pride retreats to shade with one or two males standing guard. Lions defend their kills from scavengers such as vultures and hyenas. Lions scavenge on carrion when the opportunity arises, scavenging animals dead from natural causes such as disease or those that were killed by other predators. Scavenging lions keep

5106-643: Is used to advertise its presence. Lions most often roar at night, a sound that can be heard from a distance of 8 kilometres (5 mi). They tend to roar in a very characteristic manner starting with a few deep, long roars that subside into grunts. The lion is listed as Vulnerable on the IUCN Red List . The Indian population is listed on CITES Appendix I and the African population on CITES Appendix II . Duchy of J%C3%BClich Its territory lies in present-day Germany (part of North Rhine-Westphalia ) and in

5217-515: The 5th century , they were re-interpreted in a Christian context in the western kingdoms of Gaul and Italy in the 6th and 7th centuries. The characteristic of the lion as royal animal in particular is due to the influence of the Physiologus , an early Christian book about animal symbolism, originally written in Greek in the 2nd century and translated into Latin in about AD 400. It was a predecessor of

5328-505: The Narmada River in central India. Lions spend much of their time resting; they are inactive for about twenty hours per day. Although lions can be active at any time, their activity generally peaks after dusk with a period of socialising, grooming, and defecating. Intermittent bursts of activity continue until dawn, when hunting most often takes place. They spend an average of two hours a day walking and fifty minutes eating. The lion

5439-478: The Nile Basin during the early Holocene . In zoos, lions have been bred with tigers to create hybrids for the curiosity of visitors or for scientific purpose. The liger is bigger than a lion and a tiger, whereas most tigons are relatively small compared to their parents because of reciprocal gene effects. The leopon is a hybrid between a lion and leopard. The lion is a muscular, broad-chested cat with

5550-505: The Staufen (Hohenstaufen) and Wittelsbach ( Palatinate ) coats of arms, both deriving from Henry the Lion , the royal coat of arms of Scotland , attributed to William the Lion , the coat of arms of Denmark , first used by Canute VI , the coat of arms of Flanders ( Jülich ), first used by Philip I , the coat of arms of León , an example of canting arms attributed to Alfonso VII (1126), and

5661-457: The University of Birmingham . Alternatively, a lion may be depicted with one head connected to two distinct bodies, in which case it its termed bicorporated. If the conjoined bodies are three, the lion is instead termed tricorporated. These multi-bodied lions, however, are very rare. Also, the claws and tongue of the lion may be described as a different tincture than a "proper" lion. If the claws are

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5772-618: The coat of arms of Bohemia , first granted to Vladislaus II . Coats of arms of the 13th century include those of the House of Sverre ( coat of arms of Norway ), the Ludovingians (the lion of Hesse used by Conrad of Thuringia ), Luxembourg , the kingdom of Ruthenia ( Volhynia ), the House of Habsburg (the Habsburgs all but abandoned their original coat of arms after gaining the Duchy of Austria in

5883-413: The leopard , denoting a lion passant guardant, was a term of French origin that had "long since become obsolete in English armory. In French blazon, however, the old distinction is still observed." Fox-Davies continued, "[French heralds] term our lion passant a léopard-lionné and our lion rampant guardant is their lion-léopardé ." Dutch heraldist Johannes Rietstap , however, defined a Léopard lionné as

5994-459: The 1270s, but it remained in use in derived lineages such as the House of Kyburg ), the kingdom of Bulgaria and the Armenian Kingdom of Cilicia ( Rubenids ). Unlike the eagle , which is comparatively rare in heraldry because it was reserved as an imperial symbol , the lion became a symbol of chivalry and was not restricted to royal coats of arms. The Zürich armorial (14th century) has

6105-749: The Asiatic lion population. In 2017, the Cat Classification Task Force of the Cat Specialist Group revised lion taxonomy , and recognises two subspecies based on results of several phylogeographic studies on lion evolution , namely: However, there seems to be some degree of overlap between both groups in northern Central Africa. DNA analysis from a more recent study indicates that Central African lions are derived from both northern and southern lions, as they cluster with P. leo leo in mtDNA-based phylogenies whereas their genomic DNA indicates

6216-673: The Duchies of Cleves and Berg as well as Guelders and the County of Mark : Duke William II had married Mary, the daughter of Duke Reginald II of Guelders , and duchess herself after the death of her half-brother Reginald III of Guelders in 1371. William II settled the conflict with the Imperial House of Luxembourg and his son William III inherited both duchies, thereby becoming William I of Guelders and Jülich . In 1402, Duke William I Guelders and Jülich died without any legitimate offspring. He

6327-650: The Duchy of Jülich (Duché de Juliers), which became part of the French département of the Roer . The Treaty of Lunéville in 1801 officially acknowledged the cession of Jülich to France. In 1815, following the defeat of Napoleon , the duchy became part of the Prussian Province of Jülich-Cleves-Berg (after 1822 part of the Prussian Rhine Province ), except for the cities Sittard and Tegelen , which became part of

6438-474: The English crown is a seal bearing two lions passant, used by the future King John during the lifetime of his father, Henry II , who died in 1189. Since Henry was the son of Geoffrey Plantagenet, it seems reasonable to suppose that the adoption of lions as an heraldic emblem by Henry or his sons might have been inspired by Geoffrey's shield. John's elder brother, Richard the Lionheart , who succeeded his father on

6549-609: The Late Pleistocene. Lion populations in East and Southern Africa became separated from populations in West and North Africa when the equatorial rainforest expanded 183,500 to 81,800 years ago. They shared a common ancestor probably between 98,000 and 52,000 years ago. Due to the expansion of the Sahara between 83,100 and 26,600 years ago, lion populations in West and North Africa became separated. As

6660-558: The Ngorongoro Crater were afflicted by an outbreak of stable fly ( Stomoxys calcitrans ) in 1962, resulting in lions becoming emaciated and covered in bloody, bare patches. Lions sought unsuccessfully to evade the biting flies by climbing trees or crawling into hyena burrows; many died or migrated and the local population dropped from 70 to 15 individuals. A more recent outbreak in 2001 killed six lions. Captive lions have been infected with canine distemper virus (CDV) since at least

6771-538: The Swedish royal House of Bjälbo , from there in turn derived into the coat of arms of Finland , formerly belonging to Sweden. The animals of the "barbarian" ( Eurasian ) predecessors of heraldic designs are likely to have been used as clan symbols . Symmetrically paired animals in particular find continuation from Migration Period art via Insular art to Romanesque art and heraldry. Adopted in Germanic tradition around

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6882-506: The United States. Outside of classical heraldry, lions have also found their way onto the coats of arms or emblems used by modern states in Asia, often based on traditional depictions of lions in the respective cultures or regions. Lion The lion ( Panthera leo ) is a large cat of the genus Panthera , native to Africa and India . It has a muscular, broad-chested body ;

6993-691: The adjacent Timbavati Private Game Reserve in eastern South Africa. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool . Nevertheless, 17 births have been recorded in five prides between 2007 and 2015. White lions are selected for breeding in captivity. They have reportedly been bred in camps in South Africa for use as trophies to be killed during canned hunts . African lions live in scattered populations across sub-Saharan Africa. The lion prefers grassy plains and savannahs , scrub bordering rivers, and open woodlands with bushes. It rarely enters closed forests. On Mount Elgon ,

7104-424: The age of two. When a new male lion takes over a pride, adolescents both male and female may be evicted. Lions may live 12–17 years in the wild. Although adult lions have no natural predators, evidence suggests most die violently from attacks by humans or other lions. Lions often inflict serious injuries on members of other prides they encounter in territorial disputes or members of the home pride when fighting at

7215-544: The animal's expressive movements are highly developed. The most common peaceful, tactile gestures are head rubbing and social licking , which have been compared with the role of allogrooming among primates. Head rubbing, nuzzling the forehead, face and neck against another lion appears to be a form of greeting and is seen often after an animal has been apart from others or after a fight or confrontation. Males tend to rub other males, while cubs and females rub females. Social licking often occurs in tandem with head rubbing; it

7326-399: The arms by giving the lion not just one but two erect tails. As many attitudes (positions) now exist in heraldry as the heraldist's imagination can conjure, as a result of the ever-increasing need for differentiation, but very few of these were apparently known to medieval heralds. One distinction commonly made (especially among French heralds), although it may be of limited importance, is

7437-447: The arms of the kingdom of Bohemia), or cut off ( defamed ). In addition to the attitudes it is depicted in, a certain variety is present in heraldic lions regarding the presence of additional physical features. Beyond the presence of double or forked tails, heraldic lions are sometimes depicted with two heads, as in the case of the arms of the Mason of Birmingham, from whom they were passed to

7548-499: The attack. They take advantage of factors that reduce visibility; many kills take place near some form of cover or at night. One study in 2018 recorded a lion running at a top speed of 74.1 km/h (46.0 mph). The lion accelerates at the start of the chase by 9.5 m/s² , whereas zebras, wildebeest and Thomson's gazelle accelerate by 5 m/s², 5.6 m/s² and 4.5 m/s², respectively; acceleration appears to be more important than steady displacement speed in lion hunts. The lion's attack

7659-431: The attitude of their "Lions" were heraldic "leopards". In Heraldry: Sources, Symbols and Meaning (1976), German heraldist Ottfried Neubecker explained: When the blazon does not specifically mention a position, the lion may be assumed to be rampant. If he is in a different position, other terminology must be used, referring to the position of his head and limbs. An early heraldic convention found in medieval blazons uses

7770-454: The cause of the decline is not fully understood, habitat loss and conflicts with humans are the greatest causes for concern. One of the most widely recognised animal symbols in human culture, the lion has been extensively depicted in sculptures and paintings, on national flags, and in literature and films. Lions have been kept in menageries since the time of the Roman Empire and have been

7881-411: The cubs varies—one male could patiently let the cubs play with his tail or his mane, while another may snarl and bat the cubs away. Pride lionesses often synchronise their reproductive cycles and communal rearing and suckling of the young, which suckle indiscriminately from any or all of the nursing females in the pride. The synchronisation of births is advantageous because the cubs grow to being roughly

7992-595: The daughter of the last Duke, William IV. She inherited her father's estates: Jülich and Berg with the County of Ravensberg . From 1521 Jülich-Berg and Cleves formed the United Duchies of Jülich-Cleves-Berg in a personal union under Duke John III. When the last duke of Jülich-Cleves-Berg died without direct heirs in 1609, the War of the Jülich Succession broke out. It ended with the 1614 Treaty of Xanten , which divided

8103-452: The distinction between a lion and a leopard previously employed by the ancient Greeks. In antiquity, the lion, having a heavy mane, was generally shown in profile, while the leopard, having less hair, was shown looking towards the observer. A lion looking towards the observer therefore came to be given the name of the animal usually shown in that pose. According to Neubecker, what in Old French

8214-403: The distinction of lions in the walking positions as leopards . The following table summarizes the principal attitudes of heraldic lions: Note: the term segreant denotes the same position, but is only used in reference to winged four-legged beasts like griffins and dragons . Note: A lion thus depicted may be called a "leopard" (see discussion below). Other terms are used to describe

8325-534: The group. Alternatively, there may be rewards associated with being a leader that fends off intruders; the rank of lionesses in the pride is reflected in these responses. The male or males associated with the pride must defend their relationship with the pride from outside males who may attempt to usurp them. Dominance hierarchies do not appear to exist among individuals of either sex in a pride. Asiatic lion prides differ in group composition. Male Asiatic lions are solitary or associate with up to three males, forming

8436-425: The heraldic distinction between lions and leopards is often ambiguous and in some cases may be controversial (as in the case of the royal arms of England , discussed below). Part of the confusion arises from international differences in translation or in the defining characteristics of each, particularly in charges that show some characteristics of each. English herald Arthur Charles Fox-Davies asserted in 1909 that

8547-570: The hunting of large, slower-moving prey like buffalo; and even hunt them on their own. Moderately-sized hunting groups generally have higher success rates than lone females and larger groups. Lions are not particularly known for their stamina. For instance, a lioness's heart comprises only 0.57% of her body weight and a male's is about 0.45% of his body weight, whereas a hyena's heart comprises almost 1% of its body weight. Thus, lions run quickly only in short bursts at about 48–59 km/h (30–37 mph) and need to be close to their prey before starting

8658-424: The leopard, said to be "borne of an adulterous union between a lioness and a pard ," and like a mule incapable of reproducing, may be an appropriate charge for a person born of adultery or barred from reproducing (such as an abbot ). As a general rule, English heralds tend to identify lions as rampant (upright, in profile facing dexter), and leopards as passant guardant (walking, head turned to full face), but

8769-492: The lion and the rear of the vessel. This was originally the result of the joining of the lions or of the royal arms of England with the ships argent of the arms of the townships of the Ports. Over time, the conjoined figure came to be considered as a single heraldic charge in itself, and granted as such as new charges. Winged lions are depicted in arms as both passant and, more commonly, sejant, and also appear as supporters. This figure

8880-705: The lion has been recorded up to an elevation of 3,600 m (11,800 ft) and close to the snow line on Mount Kenya . Savannahs with an annual rainfall of 300 to 1,500 mm (12 to 59 in) make up the majority of lion habitat in Africa, estimated at 3,390,821 km (1,309,203 sq mi) at most, but remnant populations are also present in tropical moist forests in West Africa and montane forests in East Africa. The Asiatic lion now survives only in and around Gir National Park in Gujarat, western India. Its habitat

8991-801: The lion population in Kenya's Masai Mara National Reserve declined, the spotted hyena population increased rapidly. Lions tend to dominate cheetahs and leopards, steal their kills and kill their cubs and even adults when given the chance. Cheetahs often lose their kills to lions or other predators. A study in the Serengeti ecosystem revealed that lions killed at least 17 of 125 cheetah cubs born between 1987 and 1990. Cheetahs avoid their competitors by hunting at different times and habitats. Leopards, by contrast, do not appear to be motivated by an avoidance of lions, as they use heavy vegetation regardless of whether lions are present in an area and both cats are active around

9102-427: The lion ranged throughout Africa and Eurasia, from Southeast Europe to India, but it has been reduced to fragmented populations in sub-Saharan Africa and one population in western India . It has been listed as Vulnerable on the IUCN Red List since 1996 because populations in African countries have declined by about 43% since the early 1990s. Lion populations are untenable outside designated protected areas. Although

9213-410: The lion's position in further detail. Each coat of arms has a right and left (i.e. dexter and sinister ) side - with respect to the person carrying the shield - so the left side of the shield as drawn on the page (thus the right side to the shield bearer) is called the dexter side. The lion's head is normally seen in agreement with the overall position, facing dexter (left) unless otherwise stated. If

9324-478: The lions of England were generally termed leopards until the end of the 14th century, including in the roll of arms of Henry III of England , and in a statute of Edward I of England , dating to 1300, which made reference to " signée de une teste de leopart —marked with the King's lion." In English Heraldry (1867), Boutell explained: Only when he was in this rampant attitude did the early Heralds consider any Lion to be

9435-551: The location of the hunt but sometimes drag large prey into cover. They tend to squabble over kills, particularly the males. Cubs suffer most when food is scarce but otherwise all pride members eat their fill, including old and crippled lions, which can live on leftovers. Large kills are shared more widely among pride members. An adult lioness requires an average of about 5 kg (11 lb) of meat per day while males require about 7 kg (15 lb). Lions gorge themselves and eat up to 30 kg (66 lb) in one session. If it

9546-565: The medieval bestiaries . At the time, few Europeans had a chance to encounter actual lions, so that painters had to rely on traditional depictions and had no actual animals as models. The lion as a heraldic charge is present from the very earliest development of heraldry in the 12th century. One of the earliest known examples of armory as it subsequently came to be practiced can be seen on the tomb of Geoffrey Plantagenet, Count of Anjou , who died in 1151. An enamel, probably commissioned by Geoffrey's widow between 1155 and 1160, depicts him carrying

9657-481: The mid-1970s. CDV is spread by domestic dogs and other carnivores; a 1994 outbreak in Serengeti National Park resulted in many lions developing neurological symptoms such as seizures. During the outbreak, several lions died from pneumonia and encephalitis . Feline immunodeficiency virus and lentivirus also affect captive lions. When resting, lion socialisation occurs through a number of behaviours;

9768-434: The phylogenetic relationship of the lion; it was thought to form a sister group with the jaguar that diverged 3.46 to 1.22 million years ago , but also with the leopard that diverged 3.1 to 1.95 million years ago to 4.32 to 0.02 million years ago . Hybridisation between lion and snow leopard ancestors possibly continued until about 2.1 million years ago. The lion-leopard clade

9879-588: The present-day Netherlands (part of the Limburg province), its population sharing the same Limburgish dialect. In the 9th century a certain Matfried was count of Jülich (pagus Juliacensis). The first mention of a count in the gau of Jülich in Lower Lorraine , is Gerhard I, in 1003; his grandson Gerhard III began to call himself Count of Jülich in 1081. William IV , who became count in 1219, significantly enlarged

9990-612: The previous males associated with a pride, the victors often kill any existing young cubs , perhaps because females do not become fertile and receptive until their cubs mature or die. Females often fiercely defend their cubs from a usurping male but are rarely successful unless a group of three or four mothers within a pride join forces against the male. Cubs also die from starvation and abandonment, and predation by leopards, hyenas and wild dogs. Male cubs are excluded from their maternal pride when they reach maturity at around two or three years of age, while some females may leave when they reach

10101-532: The rainforest decreased and thus gave rise to more open habitats, lions moved from West to Central Africa. Lions from North Africa dispersed to southern Europe and Asia between 38,800 and 8,300 years ago. Extinction of lions in southern Europe, North Africa and the Middle East interrupted gene flow between lion populations in Asia and Africa. Genetic evidence revealed numerous mutations in lion samples from East and Southern Africa, which indicates that this group has

10212-457: The range of the now extinct cave lion . In Hungary , the modern lion was present from about 4,500 to 3,200 years Before Present . In Ukraine , the modern lion was present from about 6,400 to 2,000 years Before Present. In Greece, it was common, as reported by Herodotus in 480 BC; it was considered rare by 300 BC and extirpated by AD 100. In Asia the lion once ranged in regions where climatic conditions supported an abundance of prey. It

10323-488: The removal of the gonads inhibits testosterone production. Rarely, both wild and captive lionesses have manes. Increased testosterone may be the cause of maned lionesses reported in northern Botswana. The white lion is a rare morph with a genetic condition called leucism , which is caused by a double recessive allele . It is not albino; it has normal pigmentation in the eyes and skin. White lions have occasionally been encountered in and around Kruger National Park and

10434-429: The same area. The evolution of sociability in lions was likely driven both by high population density and the clumped resources of savannah habitats. The larger the pride, the more high-quality territory they can defend; "hotspots" being near river confluences , where the cats have better access to water, prey and shelter (via vegetation). The area occupied by a pride is called a "pride area" whereas that occupied by

10545-420: The same size and have an equal chance of survival, and sucklings are not dominated by older cubs. Weaning occurs after six or seven months. Male lions reach maturity at about three years of age and at four to five years are capable of challenging and displacing adult males associated with another pride. They begin to age and weaken at between 10 and 15 years of age at the latest. When one or more new males oust

10656-406: The same time of day. In addition, there is no evidence that lions affect leopard abundance. Leopards take refuge in trees, though lionesses occasionally attempt to climb up and retrieve their kills. Lions similarly dominate African wild dogs , taking their kills and dispatching pups or adult dogs. Population densities of wild dogs are low in areas where lions are more abundant. However, there are

10767-420: The same time. When first introduced to the rest of the pride, lion cubs lack confidence when confronted with adults other than their mother. They soon begin to immerse themselves in the pride life, however, playing among themselves or attempting to initiate play with the adults. Lionesses with cubs of their own are more likely to be tolerant of another lioness's cubs than lionesses without cubs. Male tolerance of

10878-559: The separate duchies between Palatinate-Neuburg and the Margraviate of Brandenburg . Jülich and Berg fell to Count Palatine Wolfgang William of Neuburg and after the last duke of Palatinate-Neuburg (also Elector of the Palatinate from 1685) Charles III Philip had died without issue in 1742, Count Charles Theodore of Palatinate-Sulzbach (after 1777 also Duke of Bavaria ) inherited Jülich and Berg. In 1794 Revolutionary France occupied

10989-422: The spur are unknown. The tuft is absent at birth and develops at around 5 + 1 ⁄ 2 months of age. It is readily identifiable at the age of seven months. Its skull is very similar to that of the tiger, although the frontal region is usually more depressed and flattened and has a slightly shorter postorbital region and broader nasal openings than those of the tiger. Due to the amount of skull variation in

11100-585: The territory and in 1234 granted Jülich town privileges . By 1240 his territorial expansion created conflict on the eastern side of his territory with the Archbishop of Cologne Konrad von Hochstaden , whose troops devastated the city five years later. William IV's son Walram (Count from 1278 to 1297) remained a fierce opponent of the Bishopric, supporting Duke John I of Brabant at the 1288 Battle of Worringen against Archbishop Siegfried II of Westerburg . Walram

11211-533: The throne, is believed to have been the first to have borne the arms of three lions passant-guardant, still the arms of England, having earlier used two lions rampant combatant, which arms may also have belonged to his father. Richard is also credited with having originated the English crest of a lion statant (now statant-guardant). Apart from the lions of the Plantagenet (England and Normandy) coat of arms, 12th-century examples of lions used as heraldic charges include

11322-413: The tiger in size. The size and weight of adult lions vary across its range and habitats. Accounts of a few individuals that were larger than average exist from Africa and India. The male lion's mane is the most recognisable feature of the species. It may have evolved around 320,000–190,000 years ago. It grows downwards and backwards, covering most of the head, neck, shoulders, and chest. The mane

11433-523: The title of margrave from Emperor Louis IV, and in 1356 Emperor Charles IV of Luxembourg raised William V to the rank of duke . His son Duke William II , however, became entangled in a fierce feud with the Emperor's half-brother Wenceslaus of Luxembourg , Duke of Brabant , whom he defeated at the Battle of Baesweiler in 1371. Thereafter Jülich's history became closely intertwined with that of its neighbours:

11544-407: The two species, usually only the structure of the lower jaw can be used as a reliable indicator of species. The skeletal muscles of the lion make up 58.8% of its body weight and represent the highest percentage of muscles among mammals. The lion has a high concentration of fast twitch muscle fibers, giving them quick bursts of speed but less stamina. Among felids, the lion is second only to

11655-620: The usage, retained until late in the 14th century, which assigned to the Lions of the Royal Shield of England the name of "leopards". They were so called, not by the enemies of England for derision and insult, as some persons, in their ignorance of early Heraldry, have been pleased both to imagine and to assert; but the English Kings and Princes, who well knew their "Lions" to be Lions, in blazon styled them "leopards", because they also knew that Lions in

11766-400: Was a borrowing from Ancient Greek λέων léōn . The Hebrew word לָבִיא lavi may also be related. The generic name Panthera is traceable to the classical Latin word 'panthēra' and the ancient Greek word πάνθηρ 'panther'. Felis leo was the scientific name used by Carl Linnaeus in 1758, who described the lion in his work Systema Naturae . The genus name Panthera

11877-593: Was coined by Lorenz Oken in 1816. Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished mostly by the size and colour of their manes and skins. In the 19th and 20th centuries, several lion type specimens were described and proposed as subspecies , with about a dozen recognised as valid taxa until 2017. Between 2008 and 2016, IUCN Red List assessors used only two subspecific names: P. l. leo for African lion populations, and P. l. persica for

11988-511: Was distributed in the Asian and African Palearctic since at least the early Pliocene . The earliest fossils recognisable as lions were found at Olduvai Gorge in Tanzania and are estimated to be up to 2 million years old. Estimates for the divergence time of the modern and cave lion lineages range from 529,000 to 392,000 years ago based on mutation rate per generation time of the modern lion. There

12099-910: Was present in the Caucasus until the 10th century. It lived in Palestine until the Middle Ages and in Southwest Asia until the late 19th century. By the late 19th century, it had been extirpated in most of Turkey. The last live lion in Iran was sighted in 1942, about 65 km (40 mi) northwest of Dezful , although the corpse of a lioness was found on the banks of the Karun river in Khuzestan province in 1944. It once ranged from Sind and Punjab in Pakistan to Bengal and

12210-621: Was succeeded by his younger brother Count Gerhard V who had sided with German king Adolf of Nassau against his rival Albert I of Habsburg .Gerhard managed to retain his territories after Adolf of Nassau lost the Battle of Göllheim in 1298, and in 1314 supported the coronation of Louis IV of Wittelsbach at the nearby City of Aachen , once more against the will of the Cologne bishop. Gerhard died in 1328. His eldest son succeeded him as Count William V . Gerhard's younger son Walram became Archbishop of Cologne in 1332. In 1336 Count William received

12321-472: Was succeeded by his younger brother Reinald IV, Duke of Guelders and Jülich , who also died without heirs in 1423. The Gelderland estates chose Arnold of Egmond as duke, while Jülich amalgamated with Berg and passed to Adolf, Duke of Jülich-Berg , who belonged to a younger branch, and who had obtained Berg by virtue of the marriage of one of his ancestors. In 1511 Duke John III of Cleves inherited Jülich and Berg through marriage with Maria of Jülich-Berg ,

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