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123-574: The hard clam ( Mercenaria mercenaria ), also known as the round clam , hard-shell (or hard-shelled ) clam , or the quahog , is an edible marine bivalve mollusk that is native to the eastern shores of North America and Central America from Prince Edward Island to the Yucatán Peninsula . It is one of many unrelated edible bivalves that in the United States are frequently referred to simply as clams . Older literature sources may use
246-444: A cocktail sauce with horseradish , and often with lemon . Sometimes littlenecks are steamed and dipped in butter, though not as commonly as their soft-shelled clam cousin the "steamer". Littlenecks are often found in-the-shell in sauces, soups, stews, and clams casino , or substituted for European varieties such as the cockle in southern European seafood dishes. The largest clams are quahogs or chowders and cherrystones; they have
369-410: A halocline . If a zone undergoes a strong, vertical chemistry gradient with depth, it contains a chemocline . Temperature and salinity control ocean water density. Colder and saltier water is denser, and this density plays a crucial role in regulating the global water circulation within the ocean. The halocline often coincides with the thermocline, and the combination produces a pronounced pycnocline ,
492-438: A holoparasite such as Cuscuta derives all of its nutrients from another plant. Parasitic plants make up about one per cent of angiosperms and are in almost every biome in the world. All these plants have modified roots, haustoria , which penetrate the host plants, connecting them to the conductive system—either the xylem , the phloem , or both. This provides them with the ability to extract water and nutrients from
615-489: A mid-ocean ridge , which creates a long mountain range beneath the ocean. Together they form the global mid-oceanic ridge system that features the longest mountain range in the world. The longest continuous mountain range is 65,000 km (40,000 mi). This underwater mountain range is several times longer than the longest continental mountain range – the Andes . Oceanographers state that less than 20% of
738-469: A boundary between less dense surface water and dense deep water. Parasite Parasitism is a close relationship between species , where one organism, the parasite, lives on or inside another organism, the host , causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson characterised parasites as "predators that eat prey in units of less than one". Parasites include single-celled protozoans such as
861-511: A cause of gastroenteritis , is spread by the fecal–oral route from animals, or by eating insufficiently cooked poultry , or by contaminated water. Haemophilus influenzae , an agent of bacterial meningitis and respiratory tract infections such as influenza and bronchitis , is transmitted by droplet contact. Treponema pallidum , the cause of syphilis , is spread by sexual activity . Viruses are obligate intracellular parasites, characterised by extremely limited biological function, to
984-422: A depth of a few hundred meters or less. Human activity often has negative impacts on marine life within the continental shelf. Ocean temperatures depend on the amount of solar radiation reaching the ocean surface. In the tropics, surface temperatures can rise to over 30 °C (86 °F). Near the poles where sea ice forms, the temperature in equilibrium is about −2 °C (28 °F). In all parts of
1107-627: A faster rate than their hosts. Classic examples include interactions between vertebrate hosts and tapeworms , flukes , and those between the malaria-causing Plasmodium species, and fleas . Parasites reduce host fitness by general or specialised pathology , that ranges from parasitic castration to modification of host behaviour . Parasites increase their own fitness by exploiting hosts for resources necessary for their survival, in particular by feeding on them and by using intermediate (secondary) hosts to assist in their transmission from one definitive (primary) host to another. Although parasitism
1230-579: A gentle breeze on a pond causes ripples to form. A stronger gust blowing over the ocean causes larger waves as the moving air pushes against the raised ridges of water. The waves reach their maximum height when the rate at which they are travelling nearly matches the speed of the wind. In open water, when the wind blows continuously as happens in the Southern Hemisphere in the Roaring Forties , long, organized masses of water called swell roll across
1353-595: A hormone or by diverting nutrients. For example, the trematode Zoogonus lasius , whose sporocysts lack mouths, castrates the intertidal marine snail Tritia obsoleta chemically, developing in its gonad and killing its reproductive cells. Directly transmitted parasites, not requiring a vector to reach their hosts, include such parasites of terrestrial vertebrates as lice and mites; marine parasites such as copepods and cyamid amphipods; monogeneans ; and many species of nematodes, fungi, protozoans, bacteria, and viruses. Whether endoparasites or ectoparasites, each has
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#17328590711491476-836: A parasite employs to identify and approach a potential host are known as "host cues". Such cues can include, for example, vibration, exhaled carbon dioxide , skin odours, visual and heat signatures, and moisture. Parasitic plants can use, for example, light, host physiochemistry, and volatiles to recognize potential hosts. There are six major parasitic strategies , namely parasitic castration ; directly transmitted parasitism; trophically -transmitted parasitism; vector -transmitted parasitism; parasitoidism ; and micropredation. These apply to parasites whose hosts are plants as well as animals. These strategies represent adaptive peaks ; intermediate strategies are possible, but organisms in many different groups have consistently converged on these six, which are evolutionarily stable. A perspective on
1599-618: A parasitic alien species. First used in English in 1539, the word parasite comes from the Medieval French parasite , from the Latinised form parasitus , from Ancient Greek παράσιτος (parasitos) 'one who eats at the table of another' in turn from παρά (para) 'beside, by' and σῖτος (sitos) 'wheat, food'. The related term parasitism appears in English from 1611. Parasitism
1722-432: A phenomenon termed the biotrophy-necrotrophy switch . Pathogenic fungi are well-known causative agents of diseases on animals as well as humans. Fungal infections ( mycosis ) are estimated to kill 1.6 million people each year. One example of a potent fungal animal pathogen are Microsporidia - obligate intracellular parasitic fungi that largely affect insects, but may also affect vertebrates including humans, causing
1845-533: A predator, the European sparrowhawk , giving her time to lay her eggs in the host's nest unobserved. Host species often combat parasitic egg mimicry through egg polymorphism , having two or more egg phenotypes within a single population of a species. Multiple phenotypes in host eggs decrease the probability of a parasitic species accurately "matching" their eggs to host eggs. In kleptoparasitism (from Greek κλέπτης ( kleptēs ), "thief"), parasites steal food gathered by
1968-496: A shallow area and this, coupled with a low pressure system, can raise the surface of the ocean dramatically above a typical high tide. The average depth of the oceans is about 4 km. More precisely the average depth is 3,688 meters (12,100 ft). Nearly half of the world's marine waters are over 3,000 meters (9,800 ft) deep. "Deep ocean," which is anything below 200 meters (660 ft), covers about 66% of Earth's surface. This figure does not include seas not connected to
2091-573: A single host-species. Within that species, most individuals are free or almost free of parasites, while a minority carry a large number of parasites; this is known as an aggregated distribution . Trophically -transmitted parasites are transmitted by being eaten by a host. They include trematodes (all except schistosomes ), cestodes , acanthocephalans , pentastomids , many roundworms , and many protozoa such as Toxoplasma . They have complex life cycles involving hosts of two or more species. In their juvenile stages they infect and often encyst in
2214-430: A suitable fungus soon after germinating. Parasitic fungi derive some or all of their nutritional requirements from plants, other fungi, or animals. Plant pathogenic fungi are classified into three categories depending on their mode of nutrition: biotrophs, hemibiotrophs and necrotrophs. Biotrophic fungi derive nutrients from living plant cells, and during the course of infection they colonise their plant host in such
2337-1047: A variety of methods to infect animal hosts, including physical contact, the fecal–oral route , free-living infectious stages, and vectors, suiting their differing hosts, life cycles, and ecological contexts. Examples to illustrate some of the many possible combinations are given in the table. social behaviour (grooming) Among the many variations on parasitic strategies are hyperparasitism, social parasitism, brood parasitism, kleptoparasitism, sexual parasitism, and adelphoparasitism. Hyperparasites feed on another parasite, as exemplified by protozoa living in helminth parasites, or facultative or obligate parasitoids whose hosts are either conventional parasites or parasitoids. Levels of parasitism beyond secondary also occur, especially among facultative parasitoids. In oak gall systems, there can be up to five levels of parasitism. Hyperparasites can control their hosts' populations, and are used for this purpose in agriculture and to some extent in medicine . The controlling effects can be seen in
2460-469: A variety of routes. To give a few examples, Bacillus anthracis , the cause of anthrax , is spread by contact with infected domestic animals ; its spores , which can survive for years outside the body, can enter a host through an abrasion or may be inhaled. Borrelia , the cause of Lyme disease and relapsing fever , is transmitted by vectors, ticks of the genus Ixodes , from the diseases' reservoirs in animals such as deer . Campylobacter jejuni ,
2583-403: A wave-cut platform develops at the foot of the cliff and this has a protective effect, reducing further wave-erosion. Material worn from the margins of the land eventually ends up in the sea. Here it is subject to attrition as currents flowing parallel to the coast scour out channels and transport sand and pebbles away from their place of origin. Sediment carried to the sea by rivers settles on
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#17328590711492706-474: A way as to keep it alive for a maximally long time. One well-known example of a biotrophic pathogen is Ustilago maydis , causative agent of the corn smut disease. Necrotrophic pathogens on the other hand, kill host cells and feed saprophytically , an example being the root-colonising honey fungi in the genus Armillaria . Hemibiotrophic pathogens begin their colonising their hosts as biotrophs, and subsequently killing off host cells and feeding as necrotrophs,
2829-406: A zone undergoes dramatic changes in temperature with depth, it contains a thermocline , a distinct boundary between warmer surface water and colder deep water. In tropical regions, the thermocline is typically deeper compared to higher latitudes. Unlike polar waters , where solar energy input is limited, temperature stratification is less pronounced, and a distinct thermocline is often absent. This
2952-572: Is quahog ( / ˈ k oʊ h ɒ ɡ / KOH -hog , / ˈ k w ɔː h ɒ ɡ / KWAW -hog , or / k w ə ˈ h ɒ ɡ / kwə- HOG , also spelt quahaug , quohog or cohog ). The word comes from the Narragansett word "poquauhock", which is similar in Wampanoag and some other Algonquian languages ; it is first attested in North American English in 1794. Native polities on
3075-439: Is a bay , a small bay with a narrow inlet is a cove and a large bay may be referred to as a gulf . Coastlines are influenced by several factors including the strength of the waves arriving on the shore, the gradient of the land margin, the composition and hardness of the coastal rock, the inclination of the off-shore slope and the changes of the level of the land due to local uplift or submergence. Normally, waves roll towards
3198-563: Is a kind of symbiosis , a close and persistent long-term biological interaction between a parasite and its host. Unlike saprotrophs , parasites feed on living hosts, though some parasitic fungi, for instance, may continue to feed on hosts they have killed. Unlike commensalism and mutualism , the parasitic relationship harms the host, either feeding on it or, as in the case of intestinal parasites, consuming some of its food. Because parasites interact with other species, they can readily act as vectors of pathogens, causing disease . Predation
3321-675: Is aggregated. Coinfection by multiple parasites is common. Autoinfection , where (by exception) the whole of the parasite's life cycle takes place in a single primary host, can sometimes occur in helminths such as Strongyloides stercoralis . Vector-transmitted parasites rely on a third party, an intermediate host, where the parasite does not reproduce sexually, to carry them from one definitive host to another. These parasites are microorganisms, namely protozoa , bacteria , or viruses , often intracellular pathogens (disease-causers). Their vectors are mostly hematophagic arthropods such as fleas, lice, ticks, and mosquitoes. For example,
3444-464: Is also known as the Northern quahog, round clam, or chowder clam. In fish markets, there are specialist names for different sizes of this species of clam. The smallest legally harvestable clams are called countnecks or peanuts , next size up are littlenecks , then topnecks . Above that are the cherrystones , and the largest are called quahogs or chowder clams . The most distinctive of these names
3567-438: Is an important commercial form of aquaculture . For example, the species is an important member of the suspension-feeding, benthic fauna of the lower Chesapeake Bay . Rhode Island is situated right in the middle of "quahog country" and has supplied a quarter of the U.S.'s total annual commercial quahog catch. The quahog is the official shellfish of the U.S. state of Rhode Island . The species has also been introduced and
3690-444: Is an important reference point for oceanography and geography, particularly as mean sea level . The ocean surface has globally little, but measurable topography , depending on the ocean's volumes. The ocean surface is a crucial interface for oceanic and atmospheric processes. Allowing interchange of particles, enriching the air and water, as well as grounds by some particles becoming sediments . This interchange has fertilized life in
3813-443: Is by definition not a symbiosis, as the interaction is brief, but the entomologist E. O. Wilson has characterised parasites as "predators that eat prey in units of less than one". Within that scope are many possible strategies. Taxonomists classify parasites in a variety of overlapping schemes, based on their interactions with their hosts and on their life cycles , which can be complex. An obligate parasite depends completely on
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3936-411: Is customarily divided into five principal oceans – listed below in descending order of area and volume: The ocean fills Earth's oceanic basins . Earth's oceanic basins cover different geologic provinces of Earth's oceanic crust as well as continental crust . As such it covers mainly Earth's structural basins , but also continental shelfs . In mid-ocean, magma is constantly being thrust through
4059-399: Is due to the fact that surface waters in polar latitudes are nearly as cold as deeper waters. Below the thermocline, water everywhere in the ocean is very cold, ranging from −1 °C to 3 °C. Because this deep and cold layer contains the bulk of ocean water, the average temperature of the world ocean is 3.9 °C. If a zone undergoes dramatic changes in salinity with depth, it contains
4182-512: Is farmed on the Pacific coast of North America and in Great Britain and continental Europe . It reproduces sexually by females and males shedding gametes into the water. Quahog parasite unknown (QPX) is a parasite that affects the hard shell clam Mercenaria mercenaria . While little is known about the disease, research is currently under way in several laboratories. This research is fueled by
4305-645: Is found in the ant Tetramorium inquilinum , an obligate parasite which lives exclusively on the backs of other Tetramorium ants. A mechanism for the evolution of social parasitism was first proposed by Carlo Emery in 1909. Now known as " Emery's rule ", it states that social parasites tend to be closely related to their hosts, often being in the same genus. Intraspecific social parasitism occurs in parasitic nursing, where some individual young take milk from unrelated females. In wedge-capped capuchins , higher ranking females sometimes take milk from low ranking females without any reciprocation. In brood parasitism ,
4428-500: Is irregular, unevenly dominating the Earth's surface . This leads to the distinction of the Earth's surface into a water and land hemisphere , as well as the division of the ocean into different oceans. Seawater covers about 361,000,000 km (139,000,000 sq mi) and the ocean's furthest pole of inaccessibility , known as " Point Nemo ", in a region known as spacecraft cemetery of
4551-585: Is often unambiguous, it is part of a spectrum of interactions between species , grading via parasitoidism into predation, through evolution into mutualism , and in some fungi, shading into being saprophytic . Human knowledge of parasites such as roundworms and tapeworms dates back to ancient Egypt , Greece , and Rome . In early modern times, Antonie van Leeuwenhoek observed Giardia lamblia with his microscope in 1681, while Francesco Redi described internal and external parasites including sheep liver fluke and ticks . Modern parasitology developed in
4674-454: Is produced and magma is forced up creating underwater mountains, some of which may form chains of volcanic islands near to deep trenches. Near some of the boundaries between the land and sea, the slightly denser oceanic plates slide beneath the continental plates and more subduction trenches are formed. As they grate together, the continental plates are deformed and buckle causing mountain building and seismic activity. Every ocean basin has
4797-407: Is pushed across the surface of the ocean by the wind, but this represents a transfer of energy and not horizontal movement of water. As waves approach land and move into shallow water , they change their behavior. If approaching at an angle, waves may bend ( refraction ) or wrap around rocks and headlands ( diffraction ). When the wave reaches a point where its deepest oscillations of the water contact
4920-423: Is reflected back out of the water. Red light is most easily absorbed and thus does not reach great depths, usually to less than 50 meters (164 ft). Blue light, in comparison, can penetrate up to 200 meters (656 ft). Second, water molecules and very tiny particles in ocean water preferentially scatter blue light more than light of other colors. Blue light scattering by water and tiny particles happens even in
5043-419: Is seen in some species of anglerfish , such as Ceratias holboelli , where the males are reduced to tiny sexual parasites , wholly dependent on females of their own species for survival, permanently attached below the female's body, and unable to fend for themselves. The female nourishes the male and protects him from predators, while the male gives nothing back except the sperm that the female needs to produce
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5166-467: Is sometimes referred to as the World Ocean, global ocean or the great ocean . The concept of a continuous body of water with relatively unrestricted exchange between its components is critical in oceanography . The word ocean comes from the figure in classical antiquity , Oceanus ( / oʊ ˈ s iː ə n ə s / ; Ancient Greek : Ὠκεανός Ōkeanós , pronounced [ɔːkeanós] ),
5289-469: Is then carried to a nest, sometimes alongside other prey if it is not large enough to support a parasitoid throughout its development. An egg is laid on top of the prey and the nest is then sealed. The parasitoid develops rapidly through its larval and pupal stages, feeding on the provisions left for it. Koinobiont parasitoids, which include flies as well as wasps, lay their eggs inside young hosts, usually larvae. These are allowed to go on growing, so
5412-670: The North Sea or the Red Sea . There is no sharp distinction between seas and oceans, though generally seas are smaller, and are often partly (as marginal seas ) or wholly (as inland seas ) bordered by land. The contemporary concept of the World Ocean was coined in the early 20th century by the Russian oceanographer Yuly Shokalsky to refer to the continuous ocean that covers and encircles most of Earth. The global, interconnected body of salt water
5535-472: The South Pacific Ocean , at 48°52.6′S 123°23.6′W / 48.8767°S 123.3933°W / -48.8767; -123.3933 ( Point Nemo ) . This point is roughly 2,688 km (1,670 mi) from the nearest land. There are different customs to subdivide the ocean and are adjourned by smaller bodies of water such as, seas , gulfs , bays , bights , and straits . The ocean
5658-522: The Thames Barrier is designed to protect London from a storm surge, while the failure of the dykes and levees around New Orleans during Hurricane Katrina created a humanitarian crisis in the United States. Most of the ocean is blue in color, but in some places the ocean is blue-green, green, or even yellow to brown. Blue ocean color is a result of several factors. First, water preferentially absorbs red light, which means that blue light remains and
5781-410: The carbon cycle , and the water cycle by acting as a huge heat reservoir . Ocean scientists split the ocean into vertical and horizontal zones based on physical and biological conditions. The pelagic zone is the open ocean's water column from the surface to the ocean floor. The water column is further divided into zones based on depth and the amount of light present. The photic zone starts at
5904-406: The coastline and structure of the world ocean. A global ocean has existed in one form or another on Earth for eons. Since its formation the ocean has taken many conditions and shapes with many past ocean divisions and potentially at times covering the whole globe. During colder climatic periods, more ice caps and glaciers form, and enough of the global water supply accumulates as ice to lessen
6027-591: The ocean floor , they begin to slow down. This pulls the crests closer together and increases the waves' height , which is called wave shoaling . When the ratio of the wave's height to the water depth increases above a certain limit, it " breaks ", toppling over in a mass of foaming water. This rushes in a sheet up the beach before retreating into the ocean under the influence of gravity. Earthquakes , volcanic eruptions or other major geological disturbances can set off waves that can lead to tsunamis in coastal areas which can be very dangerous. The ocean's surface
6150-505: The snubnosed eel is probably a facultative endoparasite (i.e., it is semiparasitic) that opportunistically burrows into and eats sick and dying fish. Plant-eating insects such as scale insects , aphids , and caterpillars closely resemble ectoparasites, attacking much larger plants; they serve as vectors of bacteria, fungi and viruses which cause plant diseases . As female scale insects cannot move, they are obligate parasites, permanently attached to their hosts. The sensory inputs that
6273-504: The 19th century. In human culture, parasitism has negative connotations. These were exploited to satirical effect in Jonathan Swift 's 1733 poem "On Poetry: A Rhapsody", comparing poets to hyperparasitical "vermin". In fiction, Bram Stoker 's 1897 Gothic horror novel Dracula and its many later adaptations featured a blood-drinking parasite. Ridley Scott 's 1979 film Alien was one of many works of science fiction to feature
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#17328590711496396-442: The Earth's biosphere . Oceanic evaporation , as a phase of the water cycle, is the source of most rainfall (about 90%), causing a global cloud cover of 67% and a consistent oceanic cloud cover of 72%. Ocean temperatures affect climate and wind patterns that affect life on land. One of the most dramatic forms of weather occurs over the oceans: tropical cyclones (also called "typhoons" and "hurricanes" depending upon where
6519-452: The Moon are 20x stronger than the Moon's tidal forces on the Earth.) The primary effect of lunar tidal forces is to bulge Earth matter towards the near and far sides of the Earth, relative to the moon. The "perpendicular" sides, from which the Moon appears in line with the local horizon, experience "tidal troughs". Since it takes nearly 25 hours for the Earth to rotate under the Moon (accounting for
6642-403: The Moon's 28 day orbit around Earth), tides thus cycle over a course of 12.5 hours. However, the rocky continents pose obstacles for the tidal bulges, so the timing of tidal maxima may not actually align with the Moon in most localities on Earth, as the oceans are forced to "dodge" the continents. Timing and magnitude of tides vary widely across the Earth as a result of the continents. Thus, knowing
6765-468: The Moon's gravity, oceanic tides are also substantially modulated by the Sun's tidal forces, by the rotation of the Earth, and by the shape of the rocky continents blocking oceanic water flow. (Tidal forces vary more with distance than the "base" force of gravity: the Moon's tidal forces on Earth are more than double the Sun's, despite the latter's much stronger gravitational force on Earth. Earth's tidal forces upon
6888-400: The Moon's position does not allow a local to predict tide timings, instead requiring precomputed tide tables which account for the continents and the Sun, among others. During each tidal cycle, at any given place the tidal waters rise to maximum height, high tide, before ebbing away again to the minimum level, low tide. As the water recedes, it gradually reveals the foreshore , also known as
7011-643: The Okeanos is represented with a dragon-tail on some early Greek vases. Scientists believe that a sizable quantity of water would have been in the material that formed Earth. Water molecules would have escaped Earth's gravity more easily when it was less massive during its formation. This is called atmospheric escape . During planetary formation , Earth possibly had magma oceans . Subsequently, outgassing , volcanic activity and meteorite impacts , produced an early atmosphere of carbon dioxide , nitrogen and water vapor , according to current theories. The gases and
7134-496: The QPX parasite among the thraustochytrid stramenopiles . The QPX parasite is unicellular and possesses at least one flagellum, like most other stramenopile organisms. Thraustochytrids are common protists in marine sediments and the water column, but only a few thraustochytrids are known as parasites of marine animals. Although QPX disease was first recorded on the Atlantic coast of Canada in
7257-706: The World Ocean, such as the Caspian Sea . The deepest region of the ocean is at the Mariana Trench , located in the Pacific Ocean near the Northern Mariana Islands . The maximum depth has been estimated to be 10,971 meters (35,994 ft). The British naval vessel Challenger II surveyed the trench in 1951 and named the deepest part of the trench the " Challenger Deep ". In 1960, the Trieste successfully reached
7380-635: The agents of malaria , sleeping sickness , and amoebic dysentery ; animals such as hookworms , lice , mosquitoes , and vampire bats ; fungi such as honey fungus and the agents of ringworm ; and plants such as mistletoe , dodder , and the broomrapes . There are six major parasitic strategies of exploitation of animal hosts, namely parasitic castration , directly transmitted parasitism (by contact), trophically-transmitted parasitism (by being eaten), vector-transmitted parasitism, parasitoidism , and micropredation. One major axis of classification concerns invasiveness: an endoparasite lives inside
7503-696: The air or soil given off by host shoots or roots , respectively. About 4,500 species of parasitic plant in approximately 20 families of flowering plants are known. Species within the Orobanchaceae (broomrapes) are among the most economically destructive of all plants. Species of Striga (witchweeds) are estimated to cost billions of dollars a year in crop yield loss, infesting over 50 million hectares of cultivated land within Sub-Saharan Africa alone. Striga infects both grasses and grains, including corn , rice , and sorghum , which are among
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#17328590711497626-421: The amounts in other parts of the water cycle. The reverse is true during warm periods. During the last ice age, glaciers covered almost one-third of Earth's land mass with the result being that the oceans were about 122 m (400 ft) lower than today. During the last global "warm spell," about 125,000 years ago, the seas were about 5.5 m (18 ft) higher than they are now. About three million years ago
7749-479: The atmosphere are thought to have accumulated over millions of years. After Earth's surface had significantly cooled, the water vapor over time would have condensed, forming Earth's first oceans. The early oceans might have been significantly hotter than today and appeared green due to high iron content. Geological evidence helps constrain the time frame for liquid water existing on Earth. A sample of pillow basalt (a type of rock formed during an underwater eruption)
7872-407: The bottom of the trench, manned by a crew of two men. Oceanographers classify the ocean into vertical and horizontal zones based on physical and biological conditions. The pelagic zone consists of the water column of the open ocean, and can be divided into further regions categorized by light abundance and by depth. The ocean zones can be grouped by light penetration into (from top to bottom):
7995-453: The clams' tissues and is toxic to humans when they are consumed. The toxin affects the human central nervous system . Eating contaminated shellfish, raw or cooked, can be fatal. Some other kinds of algal blooms make the seawater appear red, but red tide blooms do not always discolor the water, nor are they related to tides. Marine (ocean) The ocean is the body of salt water that covers approximately 70.8% of Earth . In English ,
8118-864: The deer tick Ixodes scapularis acts as a vector for diseases including Lyme disease , babesiosis , and anaplasmosis . Protozoan endoparasites, such as the malarial parasites in the genus Plasmodium and sleeping-sickness parasites in the genus Trypanosoma , have infective stages in the host's blood which are transported to new hosts by biting insects. Parasitoids are insects which sooner or later kill their hosts, placing their relationship close to predation. Most parasitoids are parasitoid wasps or other hymenopterans ; others include dipterans such as phorid flies . They can be divided into two groups, idiobionts and koinobionts, differing in their treatment of their hosts. Idiobiont parasitoids sting their often-large prey on capture, either killing them outright or paralysing them immediately. The immobilised prey
8241-549: The early 1960s, it did not become a major economic problem until its appearance in cultured clams at Prince Edward Island, Massachusetts in 1992, and Virginia in 1997. Infected clams are characterized by the presence of blisters or pustules in the mantle and later by gaping and death. In coastal areas of the New England states, Long Island, and New Jersey, restaurants known as raw bars or clam bars specialize in serving littlenecks and topnecks raw on an opened half-shell, usually with
8364-529: The eastern Atlantic seaboard made valuable beads called wampum from the shells, especially those colored purple; the species name mercenaria is related to the Latin word for commerce. Today people living in coastal New England still use Algonquian words for the clam, as they have done for thousands of years. In many areas where aquaculture is important, clam farmers have bred specialized versions of these clams with distinctions needed for them to be distinguished in
8487-501: The elder of the Titans in classical Greek mythology . Oceanus was believed by the ancient Greeks and Romans to be the divine personification of an enormous river encircling the world. The concept of Ōkeanós has an Indo-European connection. Greek Ōkeanós has been compared to the Vedic epithet ā-śáyāna-, predicated of the dragon Vṛtra-, who captured the cows/rivers. Related to this notion,
8610-409: The energy that would have gone into reproduction into host and parasite growth, sometimes causing gigantism in the host. The host's other systems remain intact, allowing it to survive and to sustain the parasite. Parasitic crustaceans such as those in the specialised barnacle genus Sacculina specifically cause damage to the gonads of their many species of host crabs . In the case of Sacculina ,
8733-491: The evolutionary options can be gained by considering four key questions: the effect on the fitness of a parasite's hosts; the number of hosts they have per life stage; whether the host is prevented from reproducing; and whether the effect depends on intensity (number of parasites per host). From this analysis, the major evolutionary strategies of parasitism emerge, alongside predation. Parasitic castrators partly or completely destroy their host's ability to reproduce, diverting
8856-444: The family Cuculidae , over 40% of cuckoo species are obligate brood parasites, while others are either facultative brood parasites or provide parental care. The eggs of some brood parasites mimic those of their hosts, while some cowbird eggs have tough shells, making them hard for the hosts to kill by piercing, both mechanisms implying selection by the hosts against parasitic eggs. The adult female European cuckoo further mimics
8979-412: The formation of unusually high rogue waves . Most waves are less than 3 m (10 ft) high and it is not unusual for strong storms to double or triple that height. Rogue waves, however, have been documented at heights above 25 meters (82 ft). The top of a wave is known as the crest, the lowest point between waves is the trough and the distance between the crests is the wavelength. The wave
9102-424: The host and parasitoid develop together for an extended period, ending when the parasitoids emerge as adults, leaving the prey dead, eaten from inside. Some koinobionts regulate their host's development, for example preventing it from pupating or making it moult whenever the parasitoid is ready to moult. They may do this by producing hormones that mimic the host's moulting hormones ( ecdysteroids ), or by regulating
9225-442: The host to complete its life cycle, while a facultative parasite does not. Parasite life cycles involving only one host are called "direct"; those with a definitive host (where the parasite reproduces sexually) and at least one intermediate host are called "indirect". An endoparasite lives inside the host's body; an ectoparasite lives outside, on the host's surface. Mesoparasites—like some copepods , for example—enter an opening in
9348-422: The host's body and remain partly embedded there. Some parasites can be generalists, feeding on a wide range of hosts, but many parasites, and the majority of protozoans and helminths that parasitise animals, are specialists and extremely host-specific. An early basic, functional division of parasites distinguished microparasites and macroparasites. These each had a mathematical model assigned in order to analyse
9471-453: The host's body; an ectoparasite lives outside, on the host's surface. Like predation, parasitism is a type of consumer–resource interaction , but unlike predators , parasites, with the exception of parasitoids, are much smaller than their hosts, do not kill them, and often live in or on their hosts for an extended period. Parasites of animals are highly specialised , each parasite species living on one given animal species, and reproduce at
9594-656: The host's endocrine system. A micropredator attacks more than one host, reducing each host's fitness by at least a small amount, and is only in contact with any one host intermittently. This behavior makes micropredators suitable as vectors, as they can pass smaller parasites from one host to another. Most micropredators are hematophagic , feeding on blood. They include annelids such as leeches , crustaceans such as branchiurans and gnathiid isopods, various dipterans such as mosquitoes and tsetse flies , other arthropods such as fleas and ticks, vertebrates such as lampreys , and mammals such as vampire bats . Parasites use
9717-425: The host. A parasitic plant is classified depending on where it latches onto the host, either the stem or the root, and the amount of nutrients it requires. Since holoparasites have no chlorophyll and therefore cannot make food for themselves by photosynthesis , they are always obligate parasites, deriving all their food from their hosts. Some parasitic plants can locate their host plants by detecting chemicals in
9840-477: The host. The parasitism is often on close relatives, whether within the same species or between species in the same genus or family. For instance, the many lineages of cuckoo bees lay their eggs in the nest cells of other bees in the same family. Kleptoparasitism is uncommon generally but conspicuous in birds; some such as skuas are specialised in pirating food from other seabirds, relentlessly chasing them down until they disgorge their catch. A unique approach
9963-423: The hosts suffer increased parental investment and energy expenditure to feed parasitic young, which are commonly larger than host young. The growth rate of host nestlings is slowed, reducing the host's fitness. Brood parasites include birds in different families such as cowbirds , whydahs , cuckoos , and black-headed ducks . These do not build nests of their own, but leave their eggs in nests of other species . In
10086-421: The interface between water and air is called swell – a term used in sailing , surfing and navigation . These motions profoundly affect ships on the surface of the ocean and the well-being of people on those ships who might suffer from sea sickness . Wind blowing over the surface of a body of water forms waves that are perpendicular to the direction of the wind. The friction between air and water caused by
10209-472: The intermediate host. When the intermediate-host animal is eaten by a predator, the definitive host, the parasite survives the digestion process and matures into an adult; some live as intestinal parasites . Many trophically transmitted parasites modify the behaviour of their intermediate hosts, increasing their chances of being eaten by a predator. As with directly transmitted parasites, the distribution of trophically transmitted parasites among host individuals
10332-482: The intertidal zone. The difference in height between the high tide and low tide is known as the tidal range or tidal amplitude. When the sun and moon are aligned (full moon or new moon), the combined effect results in the higher "spring tides", while the sun and moon misaligning (half moons) result in lesser tidal ranges. In the open ocean tidal ranges are less than 1 meter, but in coastal areas these tidal ranges increase to more than 10 meters in some areas. Some of
10455-471: The intestinal infection microsporidiosis . Protozoa such as Plasmodium , Trypanosoma , and Entamoeba are endoparasitic. They cause serious diseases in vertebrates including humans—in these examples, malaria, sleeping sickness, and amoebic dysentery —and have complex life cycles. Many bacteria are parasitic, though they are more generally thought of as pathogens causing disease. Parasitic bacteria are extremely diverse, and infect their hosts by
10578-441: The large blue butterfly, Phengaris arion , its larvae employing ant mimicry to parasitise certain ants, Bombus bohemicus , a bumblebee which invades the hives of other bees and takes over reproduction while their young are raised by host workers, and Melipona scutellaris , a eusocial bee whose virgin queens escape killer workers and invade another colony without a queen. An extreme example of interspecific social parasitism
10701-567: The largest group is the parasitoid wasps in the Hymenoptera. The phyla and classes with the largest numbers of parasitic species are listed in the table. Numbers are conservative minimum estimates. The columns for Endo- and Ecto-parasitism refer to the definitive host, as documented in the Vertebrate and Invertebrate columns. A hemiparasite or partial parasite such as mistletoe derives some of its nutrients from another living plant, whereas
10824-710: The largest tidal ranges in the world occur in the Bay of Fundy and Ungava Bay in Canada, reaching up to 16 meters. Other locations with record high tidal ranges include the Bristol Channel between England and Wales, Cook Inlet in Alaska, and the Río Gallegos in Argentina. Tides are not to be confused with storm surges , which can occur when high winds pile water up against the coast in
10947-417: The majority of Earth's surface. It includes the Pacific , Atlantic , Indian , Southern/Antarctic , and Arctic oceans. As a general term, "the ocean" and "the sea" are often interchangeable. Strictly speaking, a "sea" is a body of water (generally a division of the world ocean) partly or fully enclosed by land. The word "sea" can also be used for many specific, much smaller bodies of seawater, such as
11070-537: The marine worm Bonellia viridis has a similar reproductive strategy, although the larvae are planktonic. Examples of the major variant strategies are illustrated. Parasitism has an extremely wide taxonomic range, including animals, plants, fungi, protozoans, bacteria, and viruses. Parasitism is widespread in the animal kingdom, and has evolved independently from free-living forms hundreds of times. Many types of helminth including flukes and cestodes have complete life cycles involving two or more hosts. By far
11193-610: The marketplace. These are quite similar to common "wild type" Mercenaria clams, except that their shells bear distinctive markings. These are known as the notata strain of quahogs, which occur naturally in low numbers wherever quahogs are found. Hard clams are quite common throughout New England , north into Canada , and all down the Eastern seaboard of the United States to Florida ; but they are particularly abundant between Cape Cod and New Jersey , where seeding and harvesting them
11316-454: The need to inform aquaculturists , who suffer financially because of the mortality rates in clams that QPX inflicts and the ensuing years in which runs must be left fallow to clear the disease. It was discovered along the coast of Cape Cod , Massachusetts in 1995. Quahog parasite X (or quahog parasite unknown [QPX]) disease of the hard clam Mercenaria mercenaria is caused by a poorly known protistan parasite. Its DNA sequence analysis places
11439-407: The next generation. Adelphoparasitism, (from Greek ἀδελφός ( adelphós ), brother ), also known as sibling-parasitism, occurs where the host species is closely related to the parasite, often in the same family or genus. In the citrus blackfly parasitoid, Encarsia perplexa , unmated females may lay haploid eggs in the fully developed larvae of their own species, producing male offspring, while
11562-408: The ocean faces many environmental threats, such as marine pollution , overfishing , and the effects of climate change . Those effects include ocean warming , ocean acidification and sea level rise . The continental shelf and coastal waters are most affected by human activity. The terms "the ocean" or "the sea" used without specification refer to the interconnected body of salt water covering
11685-688: The ocean, deep ocean temperatures range between −2 °C (28 °F) and 5 °C (41 °F). Constant circulation of water in the ocean creates ocean currents . Those currents are caused by forces operating on the water, such as temperature and salinity differences, atmospheric circulation (wind), and the Coriolis effect . Tides create tidal currents, while wind and waves cause surface currents. The Gulf Stream , Kuroshio Current , Agulhas Current and Antarctic Circumpolar Current are all major ocean currents. Such currents transport massive amounts of water, gases, pollutants and heat to different parts of
11808-508: The ocean, on land and air. All these processes and components together make up ocean surface ecosystems . Tides are the regular rise and fall in water level experienced by oceans, primarily driven by the Moon 's gravitational tidal forces upon the Earth. Tidal forces affect all matter on Earth, but only fluids like the ocean demonstrate the effects on human timescales. (For example, tidal forces acting on rock may produce tidal locking between two planetary bodies.) Though primarily driven by
11931-469: The ocean. If the wind dies down, the wave formation is reduced, but already-formed waves continue to travel in their original direction until they meet land. The size of the waves depends on the fetch , the distance that the wind has blown over the water and the strength and duration of that wind. When waves meet others coming from different directions, interference between the two can produce broken, irregular seas. Constructive interference can lead to
12054-419: The oceans absorb CO 2 from the atmosphere , a higher concentration leads to ocean acidification (a drop in pH value ). The ocean provides many benefits to humans such as ecosystem services , access to seafood and other marine resources , and a means of transport . The ocean is known to be the habitat of over 230,000 species , but may hold considerably more – perhaps over two million species. Yet,
12177-425: The oceans could have been up to 50 m (165 ft) higher. The entire ocean, containing 97% of Earth's water, spans 70.8% of Earth 's surface, making it Earth's global ocean or world ocean . This makes Earth, along with its vibrant hydrosphere a "water world" or " ocean world ", particularly in Earth's early history when the ocean is thought to have possibly covered Earth completely. The ocean's shape
12300-434: The oceans have been mapped. The zone where land meets sea is known as the coast , and the part between the lowest spring tides and the upper limit reached by splashing waves is the shore . A beach is the accumulation of sand or shingle on the shore. A headland is a point of land jutting out into the sea and a larger promontory is known as a cape . The indentation of a coastline, especially between two headlands,
12423-533: The oceans may have always been on the Earth since the beginning of the planet's formation. In this model, atmospheric greenhouse gases kept the oceans from freezing when the newly forming Sun had only 70% of its current luminosity . The origin of Earth's oceans is unknown. Oceans are thought to have formed in the Hadean eon and may have been the cause for the emergence of life . Plate tectonics , post-glacial rebound , and sea level rise continually change
12546-451: The photic zone, the mesopelagic zone and the aphotic deep ocean zone: The pelagic part of the aphotic zone can be further divided into vertical regions according to depth and temperature: Distinct boundaries between ocean surface waters and deep waters can be drawn based on the properties of the water. These boundaries are called thermoclines (temperature), haloclines (salinity), chemoclines (chemistry), and pycnoclines (density). If
12669-417: The point where, while they are evidently able to infect all other organisms from bacteria and archaea to animals, plants and fungi, it is unclear whether they can themselves be described as living. They can be either RNA or DNA viruses consisting of a single or double strand of genetic material ( RNA or DNA , respectively), covered in a protein coat and sometimes a lipid envelope. They thus lack all
12792-535: The population movements of the host–parasite groupings. The microorganisms and viruses that can reproduce and complete their life cycle within the host are known as microparasites. Macroparasites are the multicellular organisms that reproduce and complete their life cycle outside of the host or on the host's body. Much of the thinking on types of parasitism has focused on terrestrial animal parasites of animals, such as helminths. Those in other environments and with other hosts often have analogous strategies. For example,
12915-418: The power of a storm wave impacting on the foot of a cliff has a shattering effect as air in cracks and crevices is compressed and then expands rapidly with release of pressure. At the same time, sand and pebbles have an erosive effect as they are thrown against the rocks. This tends to undercut the cliff, and normal weathering processes such as the action of frost follows, causing further destruction. Gradually,
13038-455: The seabed between adjoining plates to form mid-oceanic ridges and here convection currents within the mantle tend to drive the two plates apart. Parallel to these ridges and nearer the coasts, one oceanic plate may slide beneath another oceanic plate in a process known as subduction . Deep trenches are formed here and the process is accompanied by friction as the plates grind together. The movement proceeds in jerks which cause earthquakes, heat
13161-407: The seabed causing deltas to form in estuaries. All these materials move back and forth under the influence of waves, tides and currents. Dredging removes material and deepens channels but may have unexpected effects elsewhere on the coastline. Governments make efforts to prevent flooding of the land by the building of breakwaters , seawalls , dykes and levees and other sea defences. For instance,
13284-401: The shore at the rate of six to eight per minute and these are known as constructive waves as they tend to move material up the beach and have little erosive effect. Storm waves arrive on shore in rapid succession and are known as destructive waves as the swash moves beach material seawards. Under their influence, the sand and shingle on the beach is ground together and abraded. Around high tide,
13407-399: The source of the food supply which sustains most of the ocean ecosystem . Ocean photosynthesis also produces half of the oxygen in the Earth's atmosphere. Light can only penetrate a few hundred more meters; the rest of the deeper ocean is cold and dark (these zones are called mesopelagic and aphotic zones). The continental shelf is where the ocean meets dry land. It is more shallow, with
13530-412: The surface and is defined to be "the depth at which light intensity is only 1% of the surface value" (approximately 200 m in the open ocean). This is the zone where photosynthesis can occur. In this process plants and microscopic algae (free floating phytoplankton ) use light, water, carbon dioxide, and nutrients to produce organic matter. As a result, the photic zone is the most biodiverse and
13653-456: The system forms). As the world's ocean is the principal component of Earth's hydrosphere , it is integral to life on Earth, forms part of the carbon cycle and water cycle , and – as a huge heat reservoir – influences climate and weather patterns. The motions of the ocean surface, known as undulations or wind waves , are the partial and alternate rising and falling of the ocean surface. The series of mechanical waves that propagate along
13776-454: The systematic name Venus mercenaria ; this species is in the family Veneridae , the venus clams. Confusingly, the "ocean quahog" is a different species, Arctica islandica , which, although superficially similar in shape, is in a different family of bivalves: it is rounder than the hard clam, usually has a black periostracum , and there is no pallial sinus in the interior of the shell. The hard clam has many alternative common names . It
13899-451: The term ocean also refers to any of the large bodies of water into which the world ocean is conventionally divided. The following names describe five different areas of the ocean: Pacific , Atlantic , Indian , Antarctic/Southern , and Arctic . The ocean contains 97% of Earth's water and is the primary component of Earth's hydrosphere and is thereby essential to life on Earth. The ocean influences climate and weather patterns,
14022-466: The testes of over two-thirds of their crab hosts degenerate sufficiently for these male crabs to develop female secondary sex characteristics such as broader abdomens, smaller claws and egg-grasping appendages. Various species of helminth castrate their hosts (such as insects and snails). This may happen directly, whether mechanically by feeding on their gonads, or by secreting a chemical that destroys reproductive cells; or indirectly, whether by secreting
14145-650: The toughest meat and are used in such dishes as clam chowder , clam cakes , and stuffed clams , or are minced and mixed into dishes that use the smaller, more tender clams. Historically, Native Americans used the quahog as a component in wampum , the shell beads exchanged in the North American fur trade . The Narragansetts used the hard clam for food and ornaments. A population of hard clams exists in Southampton Water in Hampshire , England . Originally bred in
14268-403: The tropics, however effectively cheat by taking carbon from a fungus rather than exchanging it for minerals. They have much reduced roots, as they do not need to absorb water from the soil; their stems are slender with few vascular bundles , and their leaves are reduced to small scales, as they do not photosynthesize. Their seeds are small and numerous, so they appear to rely on being infected by
14391-506: The usual machinery of the cell such as enzymes , relying entirely on the host cell's ability to replicate DNA and synthesise proteins. Most viruses are bacteriophages , infecting bacteria. Parasitism is a major aspect of evolutionary ecology; for example, almost all free-living animals are host to at least one species of parasite. Vertebrates, the best-studied group, are hosts to between 75,000 and 300,000 species of helminths and an uncounted number of parasitic microorganisms. On average,
14514-434: The very clearest ocean water, and is similar to blue light scattering in the sky . Ocean water represents the largest body of water within the global water cycle (oceans contain 97% of Earth's water ). Evaporation from the ocean moves water into the atmosphere to later rain back down onto land and the ocean. Oceans have a significant effect on the biosphere . The ocean as a whole is thought to cover approximately 90% of
14637-558: The warm water outflows at Southampton Power Station for use as eel bait, the population became self-sustaining and can now be found in Southampton Water and has also spread to Portsmouth Harbour and Langstone Harbour . The term " red tide " refers to an accumulation of a toxin, such as saxitoxin , produced by marine algae. Filter-feeding shellfish are affected, such as clams, oysters, and mussels. As they filter microorganisms, clams ingest K. brevis algae. This algae accumulates in
14760-585: The way that the CHV1 virus helps to control the damage that chestnut blight , Cryphonectria parasitica , does to American chestnut trees, and in the way that bacteriophages can limit bacterial infections. It is likely, though little researched, that most pathogenic microparasites have hyperparasites which may prove widely useful in both agriculture and medicine. Social parasites take advantage of interspecific interactions between members of eusocial animals such as ants , termites , and bumblebees . Examples include
14883-471: The world's most important food crops. Orobanche also threatens a wide range of other important crops, including peas , chickpeas , tomatoes , carrots , and varieties of cabbage . Yield loss from Orobanche can be total; despite extensive research, no method of control has been entirely successful. Many plants and fungi exchange carbon and nutrients in mutualistic mycorrhizal relationships. Some 400 species of myco-heterotrophic plants, mostly in
15006-476: The world, and from the surface into the deep ocean. All this has impacts on the global climate system . Ocean water contains dissolved gases, including oxygen , carbon dioxide and nitrogen . An exchange of these gases occurs at the ocean's surface. The solubility of these gases depends on the temperature and salinity of the water. The carbon dioxide concentration in the atmosphere is rising due to CO 2 emissions , mainly from fossil fuel combustion. As
15129-750: Was recovered from the Isua Greenstone Belt and provides evidence that water existed on Earth 3.8 billion years ago. In the Nuvvuagittuq Greenstone Belt , Quebec , Canada, rocks dated at 3.8 billion years old by one study and 4.28 billion years old by another show evidence of the presence of water at these ages. If oceans existed earlier than this, any geological evidence either has yet to be discovered, or has since been destroyed by geological processes like crustal recycling . However, in August 2020, researchers reported that sufficient water to fill
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