57-418: Brazza's martin ( Phedinopsis brazzae ) is a passerine bird in the swallow family, Hirundinidae. It is 12 centimeters (4.7 in) long with grey-brown upperparts, heavily black-streaked white underparts, and a brownish tint to the breast plumage . The sexes are similar, but juvenile birds have more diffuse breast streaking and reddish-brown edges to the feathers of the back and wings. The song consists of
114-502: A tui -sized bird) and several bones of at least one species of saddleback -sized bird have recently been described. These date from the Early to Middle Miocene ( Awamoan to Lillburnian , 19–16 mya). In Europe, perching birds are not too uncommon in the fossil record from the Oligocene onward, belonging to several lineages: That suboscines expanded much beyond their region of origin
171-461: A clearer picture of passerine origins and evolution that reconciles molecular affinities, the constraints of morphology, and the specifics of the fossil record. The first passerines are now thought to have evolved in the Southern Hemisphere in the late Paleocene or early Eocene , around 50 million years ago. The initial diversification of passerines coincides with the separation of
228-471: A distinct super-family Certhioidea . This list is in taxonomic order, placing related families next to one another. The families listed are those recognised by the International Ornithologists' Union (IOC). The order and the division into infraorders, parvorders, and superfamilies follows the phylogenetic analysis published by Carl Oliveros and colleagues in 2019. The relationships between
285-469: A dozen and other species around five or six. The family Viduidae do not build their own nests, instead, they lay eggs in other birds' nests. The Passeriformes contain several groups of brood parasites such as the viduas , cuckoo-finches , and the cowbirds . The evolutionary history of the passerine families and the relationships among them remained rather mysterious until the late 20th century. In many cases, passerine families were grouped together on
342-399: A female and sang for about ten minutes. As it sang, the male leant towards the female, which caused the still-folded wings and tail to rise relative to the body. The Brazza's martin nests in the vertical banks of forested rivers from July to October, at the end of the dry season but before the river levels are high enough to cause flooding. A small colony of four pairs was recorded breeding near
399-512: A rocky outcrop on the side of a valley. The Brazza's martin nests singly or in loose colonies with sometimes widely separated burrows. The nest is a small heap of soft material such as feathers or dry grass at the end of a typically 50 cm (20 in) tunnel. The normal clutch is three white eggs. The eggs measure 18.5 mm × 11.25 mm (0.728 in × 0.443 in) and weigh 1.5 g (0.05 oz). The incubation and fledging times are unknown, although as with all hirundines
456-707: A series of short notes of increasing frequency, followed by a complex buzz that is sometimes completed by a number of clicks. The range of this species falls within the African countries of Angola , the Republic of the Congo , and the Democratic Republic of the Congo . Nesting in burrows in river banks, it lays a clutch of three white eggs. This bird feeds on flying insects, including termites , and may hunt over rivers or open savanna . It forms mixed flocks with other swallows, but
513-917: A serious impact, and this species seems under no immediate threat. Its ability to use degraded habitats also aids its survival. The Brazza's martin is not a protected species in Angola, the DRC, or the Republic of Congo. Passerine and see text A passerine ( / ˈ p æ s ə r aɪ n / ) is any bird of the order Passeriformes ( / ˈ p æ s ə r ɪ f ɔːr m iː z / ; from Latin passer 'sparrow' and formis '-shaped') which includes more than half of all bird species. Sometimes known as perching birds , passerines generally have an anisodactyl arrangement of their toes (three pointing forward and one back), which facilitates perching. With more than 140 families and some 6,500 identified species, Passeriformes
570-542: A somewhat darker brown head and white underparts heavily streaked with blackish-brown from the throat to vent. There is a brownish tint to the breast plumage. The square tail averages 46.8 mm (1.84 in) long and has white edges to the brown undertail coverts . The flight feathers are blackish-brown and the bill and legs are black. The eyes are dark brown and the black bill averages 8.5 mm (0.33 in) long. The sexes are similar, but juvenile birds have more diffuse breast streaking and reddish-brown or buff edges to
627-475: A species is a binomial or binomen, and comprises two Latin words, the first denoting the genus and the second denoting the species. The scientific name of a subspecies is formed slightly differently in the different nomenclature codes. In zoology, under the International Code of Zoological Nomenclature (ICZN), the scientific name of a subspecies is termed a trinomen , and comprises three words, namely
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#1732851277424684-426: A species. Botanists and mycologists have the choice of ranks lower than subspecies, such as variety (varietas) or form (forma), to recognize smaller differences between populations. In biological terms, rather than in relation to nomenclature, a polytypic species has two or more genetically and phenotypically divergent subspecies, races , or more generally speaking, populations that differ from each other so that
741-417: A subspecies. A common criterion for recognizing two distinct populations as subspecies rather than full species is the ability of them to interbreed even if some male offspring may be sterile. In the wild, subspecies do not interbreed due to geographic isolation or sexual selection . The differences between subspecies are usually less distinct than the differences between species. The scientific name of
798-438: Is a rank below species , used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology ), but that can successfully interbreed. Not all species have subspecies, but for those that do there must be at least two. Subspecies is abbreviated as subsp. or ssp. and the singular and plural forms are the same ("the subspecies is" or "the subspecies are"). In zoology , under
855-464: Is a subspecies or a full species, the species name may be written in parentheses. Thus Larus (argentatus) smithsonianus means the American herring gull ; the notation within the parentheses means that some consider it a subspecies of a larger herring gull species and therefore call it Larus argentatus smithsonianus , while others consider it a full species and therefore call it Larus smithsonianus (and
912-540: Is classed as Least Concern on the IUCN Red List . It is likely that this species is dug out for food by humans, but its small, dispersed colonies in firm soil suggest that it is a less rewarding target than the densely packed sandbank nests of species such as the African river martin and rosy bee-eater . Breeding colonies in river sandbars are liable to flooding, but neither natural causes nor hunting appears to be having
969-482: Is currently divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni , (suboscines) and Passeri (oscines or songbirds). The Passeri is now subdivided into two major groups recognized now as Corvides and Passerida respectively containing the large superfamilies Corvoidea and Meliphagoidea , as well as minor lineages, and the superfamilies Sylvioidea , Muscicapoidea , and Passeroidea but this arrangement has been found to be oversimplified. Since
1026-887: Is more scant before the Pleistocene, from which several still-existing families are documented. Apart from the indeterminable MACN -SC-1411 (Pinturas Early/Middle Miocene of Santa Cruz Province, Argentina), an extinct lineage of perching birds has been described from the Late Miocene of California, United States: the Palaeoscinidae with the single genus Palaeoscinis . "Palaeostruthus" eurius (Pliocene of Florida) probably belongs to an extant family, most likely passeroidean . Acanthisitti – New Zealand wrens (1 family containing 7 species, only 2 extant) Tyranni – suboscines (16 families containing 1,356 species) Passeri – oscines (125 families containing 5,158 species) The Passeriformes
1083-458: Is now known to breed in the south of the Democratic Republic of the Congo (DRC), the Republic of the Congo , and in northern Angola . There is one probable sighting from southeast Gabon . In the breeding season this martin is found near rivers with the steep banks that are needed for the nest burrows. Suitable habitat occurs along lowland tropical rivers like the Congo or rivers with sandbanks in
1140-523: Is proven by several fossils from Germany such as a presumed broadbill ( Eurylaimidae ) humerus fragment from the Early Miocene (roughly 20 mya) of Wintershof , Germany, the Late Oligocene carpometacarpus from France listed above, and Wieslochia , among others. Extant Passeri super-families were quite distinct by that time and are known since about 12–13 mya when modern genera were present in
1197-469: Is readily identified by its combination of brown upperparts, streaked underparts, and square tail. Although this little-known bird had been classified as data deficient by the International Union for Conservation of Nature (IUCN), it actually appears to be common and widespread, and it has been listed as a species of Least Concern since 2008. There may be some hunting of this martin for food, but
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#17328512774241254-442: Is the long-tailed widowbird . The chicks of passerines are altricial : blind, featherless, and helpless when hatched from their eggs. Hence, the chicks require extensive parental care. Most passerines lay colored eggs, in contrast with nonpasserines, most of whose eggs are white except in some ground-nesting groups such as Charadriiformes and nightjars , where camouflage is necessary, and in some parasitic cuckoos , which match
1311-459: Is the short-tailed pygmy tyrant , at 6.5 cm (2.6 in) and 4.2 g (0.15 oz). The foot of a passerine has three toes directed forward and one toe directed backward, called anisodactyl arrangement. The hind toe ( hallux ) is long and joins the leg at approximately the same level as the front toes. This arrangement enables passerine birds to easily perch upright on branches. The toes have no webbing or joining, but in some cotingas ,
1368-545: Is the largest order of birds and among the most diverse clades of terrestrial vertebrates , representing 60% of birds. Passerines are divided into three suborders : Acanthisitti (New Zealand wrens), Tyranni (composed mostly of South American suboscines), and Passeri (oscines or songbirds). Passerines originated in the Southern Hemisphere around 60 million years ago. Most passerines are insectivorous or omnivorous , and eat both insects and fruit or seeds. The terms "passerine" and "Passeriformes" are derived from
1425-413: Is variously described as heavy, or like that of a sand martin . Brazza's martin can be distinguished from most other swallows within its range by the heavy streaking on the underparts. Although the lesser striped swallow also has white underparts with dark streaking, it is larger, has a deeply forked tail and a very different plumage, with dark blue upperparts, a red rump and a chestnut head. Compared to
1482-652: The Corvida and numerous minor lineages make up songbird diversity today. Extensive biogeographical mixing happens, with northern forms returning to the south, southern forms moving north, and so on. Perching bird osteology , especially of the limb bones, is rather diagnostic. However, the early fossil record is poor because passerines are relatively small, and their delicate bones do not preserve well. Queensland Museum specimens F20688 ( carpometacarpus ) and F24685 ( tibiotarsus ) from Murgon, Queensland , are fossil bone fragments initially assigned to Passeriformes . However,
1539-559: The International Code of Zoological Nomenclature , the subspecies is the only taxonomic rank below that of species that can receive a name. In botany and mycology , under the International Code of Nomenclature for algae, fungi, and plants , other infraspecific ranks , such as variety , may be named. In bacteriology and virology , under standard bacterial nomenclature and virus nomenclature , there are recommendations but not strict requirements for recognizing other important infraspecific ranks. A taxonomist decides whether to recognize
1596-621: The Old World warblers and Old World babblers have turned out to be paraphyletic and are being rearranged. Several taxa turned out to represent highly distinct lineages, so new families had to be established, some of theirs – like the stitchbird of New Zealand and the Eurasian bearded reedling – monotypic with only one living species. In the Passeri alone, a number of minor lineages will eventually be recognized as distinct superfamilies. For example,
1653-631: The crows , do not sound musical to human beings. Some, such as the lyrebird , are accomplished mimics. The New Zealand wrens are tiny birds restricted to New Zealand , at least in modern times; they were long placed in Passeri. Most passerines are smaller than typical members of other avian orders. The heaviest and altogether largest passerines are the thick-billed raven and the larger races of common raven , each exceeding 1.5 kg (3.3 lb) and 70 cm (28 in). The superb lyrebird and some birds-of-paradise , due to very long tails or tail coverts, are longer overall. The smallest passerine
1710-445: The kinglets constitute a single genus with less than 10 species today but seem to have been among the first perching bird lineages to diverge as the group spread across Eurasia. No particularly close relatives of theirs have been found among comprehensive studies of the living Passeri, though they might be fairly close to some little-studied tropical Asian groups. Nuthatches , wrens , and their closest relatives are currently grouped in
1767-503: The scientific name of the house sparrow , Passer domesticus , and ultimately from the Latin term passer , which refers to sparrows and similar small birds. The order is divided into three suborders, Tyranni (suboscines), Passeri (oscines or songbirds), and the basal Acanthisitti . Oscines have the best control of their syrinx muscles among birds, producing a wide range of songs and other vocalizations, though some of them, such as
Brazza's martin - Misplaced Pages Continue
1824-402: The superb lyrebird has 16, and several spinetails in the family Furnariidae have 10, 8, or even 6, as is the case of Des Murs's wiretail . Species adapted to tree trunk climbing such as treecreepers and woodcreeper have stiff tail feathers that are used as props during climbing. Extremely long tails used as sexual ornaments are shown by species in different families. A well-known example
1881-417: The "core martins", including burrowing species like the sand martin ; the "nest-adopters", which are birds like the tree swallow that use natural cavities; and the "mud nest builders", such as the barn swallow , which build a nest from mud. The species nests in burrows and therefore belong to the "core martins" group. Brazza's martin is thought to be an early offshoot from the main swallow lineage, although
1938-524: The Brazza's martin was classified as data deficient . It was thought to have a much more restricted distribution range, but a 2007 paper presented evidence that extended its known range by 500 km (300 mi) to the north and 175 km (100 mi) southwards, a quadrupling of the area. It has a large range of 402,000 km (155,000 sq mi) although the total population is unknown. Its extensive range and apparently stable population means this bird
1995-479: The Congo sand martin or Congo martin , Riparia congica . Brazza's martin is a member of the swallow family of birds, and is classed as a member of the Hirundininae subfamily, which comprises all swallows and martins except the very distinctive river martins . DNA sequence studies suggest that there are three major groupings within the Hirundininae, broadly correlating with the type of nest built. These groups are
2052-586: The German ornithologist Hans Edmund Wolters . The genus name, Phedinopsis , combines the genus Phedina with the Ancient Greek opsis meaning "appearance". The species name commemorates Italian-born French explorer Pierre Savorgnan de Brazza , later to become governor-general of the French Congo , who collected the type specimen. This species was often called the "Congo martin", but this invites confusion with
2109-511: The Mascarene martin, this species is smaller, has a plainer back and finer dashing on the throat and chest, but there is no range overlap. The distribution of the Brazza's martin was initially poorly known, and until 1922 the type specimen in the Paris Museum was the only recorded example of this species. A Father Callewaert then collected 20 martins near Luluabourg (now Kananga ), and this bird
2166-596: The basis of morphological similarities that, it is now believed, are the result of convergent evolution , not a close genetic relationship. For example, the wrens of the Americas and Eurasia , those of Australia , and those of New Zealand look superficially similar and behave in similar ways, yet belong to three far-flung branches of the passerine family tree; they are as unrelated as it is possible to be while remaining Passeriformes. Advances in molecular biology and improved paleobiogeographical data gradually are revealing
2223-449: The binomen followed by the name of the subspecies. For example, the binomen for the leopard is Panthera pardus . The trinomen Panthera pardus fusca denotes a subspecies, the Indian leopard . All components of the trinomen are written in italics. In botany , subspecies is one of many ranks below that of species, such as variety , subvariety , form , and subform. To identify the rank,
2280-403: The chicks are altricial , hatching naked and blind. As with other swallows, Brazza's martin feeds on flying insects, including termites , and may hunt over rivers or open savanna. It can occur in single-species flocks or with other swallows including barn swallows, lesser striped swallows, or rock martins . There has been little ornithological research in this part of Africa, and until 2008
2337-502: The corvoidean and basal songbirds. The modern diversity of Passerida genera is known mostly from the Late Miocene onward and into the Pliocene (about 10–2 mya). Pleistocene and early Holocene lagerstätten (<1.8 mya) yield numerous extant species, and many yield almost nothing but extant species or their chronospecies and paleosubspecies. In the Americas , the fossil record
Brazza's martin - Misplaced Pages Continue
2394-404: The families in the suborder Tyranni (suboscines) were all well determined but some of the nodes in Passeri (oscines or songbirds) were unclear owing to the rapid splitting of the lineages. Infraorder Eurylaimides : Old World suboscines Infraorder Tyrannides : New World suboscines Parvorder Furnariida Parvorder Tyrannida Relationships between living Passeriformes families based on
2451-446: The feathers of the back and wings. This bird has no subspecies . The song consists of a series of short notes increasing in frequency which are followed by a complex buzz and sometimes completed by a number of clicks. The song becomes increasingly loud, although the final clicks are quite soft. The song is similar to that of the banded martin, and does not resemble the Mascarene martin, suggesting unresolved taxonomic problems. The flight
2508-504: The highlands of Angola. The highland locations have wide grassy riverways running through miombo woodlands , whereas the Congo Basin is tropical forest with over 200 cm (80 in) of rain a year. The lowland habitats are a patchwork of dry, seasonally flooded and permanently wet woodland, and seasonally flooded savanna , and the swamp forests contain trees such as Symphonia globulifera , raffia palms and Mitragyna species, and
2565-649: The material is too fragmentary and their affinities have been questioned. Several more recent fossils from the Oligocene of Europe, such as Wieslochia , Jamna , Resoviaornis , and Crosnoornis , are more complete and definitely represent early passeriforms, and have been found to belong to a variety of modern and extinct lineages. From the Bathans Formation at the Manuherikia River in Otago , New Zealand, MNZ S42815 (a distal right tarsometatarsus of
2622-524: The mid-2000s, studies have investigated the phylogeny of the Passeriformes and found that many families from Australasia traditionally included in the Corvoidea actually represent more basal lineages within oscines. Likewise, the traditional three-superfamily arrangement within the Passeri has turned out to be far more complex and will require changes in classification. Major " wastebin " families such as
2679-509: The passerine host's egg. The vinous-throated parrotbill has two egg colors, white and blue, to deter the brood parasitic common cuckoo . Clutches vary considerably in size: some larger passerines of Australia such as lyrebirds and scrub-robins lay only a single egg, most smaller passerines in warmer climates lay between two and five, while in the higher latitudes of the Northern Hemisphere, hole-nesting species like tits can lay up to
2736-943: The phylogenetic analysis of Oliveros et al (2019). Some terminals have been renamed to reflect families recognised by the IOC but not in that study. The IOC families Alcippeidae and Teretistridae were not sampled in this study. Acanthisittidae (New Zealand wrens) Eurylaimidae (eurylaimid broadbills) Philepittidae (asites) Calyptomenidae (African and green broadbills) Pittidae (pittas) Sapayoidae (sapayoa) Melanopareiidae (crescent chests) Conopophagidae (gnateaters) Thamnophilidae (antbirds) Grallariidae (antpittas) Rhinocryptidae (tapaculos) Formicariidae (antthrushes) Scleruridae (leaftossers) Dendrocolaptidae (woodcreepers) Furnariidae (ovenbirds) Pipridae (manakins) Cotingidae (cotingas) Tityridae (tityras, becards) Subspecies In biological classification , subspecies ( pl. : subspecies)
2793-467: The rank of variety are taken to be names of subspecies (see International Code of Nomenclature of Prokaryotes ). As in botany, subspecies is conventionally abbreviated as "subsp.", and is used in the scientific name: Bacillus subtilis subsp. spizizenii . In zoological nomenclature , when a species is split into subspecies, the originally described population is retained as the "nominotypical subspecies" or "nominate subspecies", which repeats
2850-433: The riverbanks are often lined with arrowroot . This martin seems to be able to adapt to open savanna habitats containing Hymenocardia acida , in which it will roost overnight when not breeding, and is therefore not heavily dependent on the neighbouring forests as long as the riverine breeding sites survive. In a presumed example of courtship behaviour , a male Brazza's martin perched about 30 cm (1 ft) away from
2907-418: The same name as the species. For example, Motacilla alba alba (often abbreviated M. a. alba ) is the nominotypical subspecies of the white wagtail ( Motacilla alba ). The subspecies name that repeats the species name is referred to in botanical nomenclature as the subspecies " autonym ", and the subspecific taxon as the "autonymous subspecies". When zoologists disagree over whether a certain population
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#17328512774242964-509: The second and third toes are united at their basal third. The leg of passerine birds contains an additional special adaptation for perching. A tendon in the rear of the leg running from the underside of the toes to the muscle behind the tibiotarsus will automatically be pulled and tighten when the leg bends, causing the foot to curl and become stiff when the bird lands on a branch. This enables passerines to sleep while perching without falling off. Most passerine birds have 12 tail feathers but
3021-911: The southern continents in the early Eocene . The New Zealand wrens are the first to become isolated in Zealandia , and the second split involved the origin of the Tyranni in South America and the Passeri in the Australian continent . The Passeri experienced a great radiation of forms in Australia. A major branch of the Passeri, the parvorder Passerida , dispersed into Eurasia and Africa about 40 million years ago, where they experienced further radiation of new lineages. This eventually led to three major Passerida lineages comprising about 4,000 species, which in addition to
3078-509: The species does not appear to be facing any serious short-term threats. The Brazza's martin was first described in 1886 under the binomial name Phedina brazzae by the French zoologist Émile Oustalet from a specimen obtained at Nganchu in the Ngabé District of what is now the Republic of Congo. This martin is now the only species placed in the genus Phedinopsis that was introduced in 1971 by
3135-468: The striped plumage suggests a distant relationship with several streaked African Hirundo species. Brazza's martin was previously included in the genus Phedina , although it is now included in its own genus Phedinopsis due to the significant differences in vocalisations and nest type from its relative. The Brazza's martin is 12 cm (4.7 in) long with wings averaging 100.5 mm (3.96 in). This small hirundine has grey-brown upperparts with
3192-556: The subspecific name must be preceded by "subspecies" (which can be abbreviated to "subsp." or "ssp."), as in Schoenoplectus californicus subsp. tatora . In bacteriology , the only rank below species that is regulated explicitly by the code of nomenclature is subspecies , but infrasubspecific taxa are extremely important in bacteriology; Appendix 10 of the code lays out some recommendations that are intended to encourage uniformity in describing such taxa. Names published before 1992 in
3249-425: The user of the notation is not taking a position). A subspecies is a taxonomic rank below species – the only such rank recognized in the zoological code, and one of three main ranks below species in the botanical code. When geographically separate populations of a species exhibit recognizable phenotypic differences, biologists may identify these as separate subspecies; a subspecies is a recognized local variant of
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