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113-557: The coat of arms of the rione depicts a rampant griffon , a Greek mythological creature with the head of an eagle and the body of a lion . It was chosen as a symbol of pride and nobility. During antiquity, it belonged to the IX Augustan region called Circus Flaminius . In this area Domitianus built his stadium and an Odeon ( Odeum in Latin ), for musical and poetic competitions. Pompey too built there his curia . Around

226-417: A buildup of scent attracting the attention of predators, the lioness moves her cubs to a new den site several times a month, carrying them one-by-one by the nape of the neck. Usually, the mother does not integrate herself and her cubs back into the pride until the cubs are six to eight weeks old. Sometimes the introduction to pride life occurs earlier, particularly if other lionesses have given birth at about

339-540: A closer relationship with P. leo melanochaita . Lion samples from some parts of the Ethiopian Highlands cluster genetically with those from Cameroon and Chad, while lions from other areas of Ethiopia cluster with samples from East Africa. Researchers, therefore, assume Ethiopia is a contact zone between the two subspecies. Genome -wide data of a wild-born historical lion sample from Sudan showed that it clustered with P. l. leo in mtDNA-based phylogenies, but with

452-471: A constant lookout for circling vultures, which indicate the death or distress of an animal. Most carrion on which both hyenas and lions feed upon are killed by hyenas rather than lions. Carrion is thought to provide a large part of lion diet. Lions and spotted hyenas occupy a similar ecological niche and compete for prey and carrion; a review of data across several studies indicates a dietary overlap of 58.6%. Lions typically ignore hyenas unless they are on

565-425: A favoured position in the group, either stalking prey on the "wing", then attacking, or moving a smaller distance in the centre of the group and capturing prey fleeing from other lionesses. Males attached to prides do not usually participate in group hunting. Some evidence suggests, however, that males are just as successful as females; they are typically solo hunters who ambush prey in small bushland. They may join in

678-470: A few reported cases of old and wounded lions falling prey to wild dogs. Most lionesses reproduce by the time they are four years of age. Lions do not mate at a specific time of year and the females are polyestrous . Like those of other cats, the male lion's penis has spines that point backward. During withdrawal of the penis, the spines rake the walls of the female's vagina, which may cause ovulation . A lioness may mate with more than one male when she

791-548: A heavier mane. On average, Asiatic lions have sparser manes than African lions. This feature likely evolved to signal the fitness of males to females. Males with darker manes appear to have greater reproductive success and are more likely to remain in a pride for longer. They have longer and thicker hair and higher testosterone levels, but they are also more vulnerable to heat stress. The core body temperature does apparently not increase regardless of sex, season, feeding time, length and colour of mane, but only surface temperature

904-590: A high affinity to P. l. melanochaita . This result suggested that the taxonomic position of lions in Central Africa may require revision. Other lion subspecies or sister species to the modern lion existed in prehistoric times: The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago . Results of analyses differ in

1017-473: A key species sought for exhibition in zoological gardens across the world since the late 18th century. Cultural depictions of lions were prominent in Ancient Egypt , and depictions have occurred in virtually all ancient and medieval cultures in the lion's historic and current range. The English word lion is derived via Anglo-Norman liun from Latin leōnem (nominative: leō ), which in turn

1130-495: A kill or are being harassed, while the latter tend to visibly react to the presence of lions with or without the presence of food. In the Ngorongoro crater , lions subsist largely on kills stolen from hyenas, causing them to increase their kill rate. In Botswana's Chobe National Park, the situation is reversed as hyenas there frequently challenge lions and steal their kills, obtaining food from 63% of all lion kills. When confronted on

1243-505: A kill, hyenas may either leave or wait patiently at a distance of 30–100 m (98–328 ft) until the lions have finished. Hyenas may feed alongside lions and force them off a kill. The two species attack one another even when there is no food involved for no apparent reason. Lions can account for up to 71% of hyena deaths in Etosha National Park . Hyenas have adapted by frequently mobbing lions that enter their home ranges. When

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1356-519: A kill. Crippled lions and cubs may fall victim to hyenas and leopards or be trampled by buffalo or elephants. Careless lions may be maimed when hunting prey. Nile crocodiles may also kill and eat lions, evidenced by the occasional lion claw found in crocodile stomachs. Ticks commonly infest the ears, neck and groin regions of the lions. Adult forms of several tapeworm species of the genus Taenia have been isolated from lion intestines, having been ingested as larvae in antelope meat. Lions in

1469-558: A lion makes when sniffing chemical signals and involves an open mouth with bared teeth, raised muzzle, wrinkled nose, closed eyes and relaxed ears. Lions also use chemical and visual marking; males spray urine and scrape plots of ground and objects within the territory. The lion's repertoire of vocalisations is large; variations in intensity and pitch appear to be central to communication. Most lion vocalisations are variations of growling , snarling , meowing and roaring. Other sounds produced include puffing, bleating and humming. Roaring

1582-473: A longer evolutionary history than genetically less diverse lion samples from Asia and West and Central Africa. A whole genome-wide sequence of lion samples showed that samples from West Africa shared alleles with samples from Southern Africa, and samples from Central Africa shared alleles with samples from Asia. This phenomenon indicates that Central Africa was a melting pot of lion populations after they had become isolated, possibly migrating through corridors in

1695-399: A loose pride while females associate with up to 12 other females, forming a stronger pride together with their cubs. Female and male lions associate only when mating. Coalitions of males hold territory for a longer time than single lions. Males in coalitions of three or four individuals exhibit a pronounced hierarchy, in which one male dominates the others and mates more frequently. The lion

1808-449: A muscular, broad-chested body ; a short, rounded head; round ears; and a dark, hairy tuft at the tip of its tail. It is sexually dimorphic ; adult male lions are larger than females and have a prominent mane. It is a social species , forming groups called prides. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on medium-sized and large ungulates . The lion

1921-433: A nomad is a "range". Males associated with a pride patrol the fringes. Both males and females defend the pride against intruders, but the male lion is better-suited for this purpose due to its stockier, more powerful build. Some individuals consistently lead the defense against intruders, while others lag behind. Lions tend to assume specific roles in the pride; slower-moving individuals may provide other valuable services to

2034-544: A nomadic phase before gaining residence in a pride. A study undertaken in the Serengeti National Park revealed that nomadic coalitions gain residency at between 3.5 and 7.3 years of age. In Kruger National Park, dispersing male lions move more than 25 km (16 mi) away from their natal pride in search of their own territory. Female lions stay closer to their natal pride. Therefore, female lions in an area are more closely related to each other than male lions in

2147-449: A palace was jutting, its front was moved backwards to preserve it. Northward, the rione borders with Ponte (R. V), whose boundary is defined by Piazza delle Cinque Lune, Piazza di Tor Sanguigna, Largo Febo, Via di Santa Maria dell'Anima, Via di Tor Millina, Via della Pace, Piazza del Fico, Via del Corallo, Via del Governo Vecchio, Via dei Filippini and Vicolo Cellini. To the west and to the south, Parione borders with Regola (R. VII),

2260-481: A red chest and white wings". Aelian was the last source on the griffin to add fresh information on the griffin, and late writers (into medieval times) merely rehashed existing material on griffins, with the exception of the lore about their "agate eggs" which emerged at some indistinct time later on (cf. infra). The griffin has been associated with various deities (Apollo, Dionysus, Nemesis), in Greek mythography but here,

2373-425: A reed-bed, a cave, or some other sheltered area, usually away from the pride. She will often hunt alone while the cubs are still helpless, staying relatively close to the den. Lion cubs are born blind, their eyes opening around seven days after birth. They weigh 1.2–2.1 kg (2.6–4.6 lb) at birth and are almost helpless, beginning to crawl a day or two after birth and walking around three weeks of age. To avoid

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2486-413: A region of very rich soil but quite uninhabitable because griffins, a savage and tenacious breed of wild beasts, love.. the gold that is mined from deep within the earth there, and because they guard it with an amazing hostility to those who set foot there. The aforementioned Aelian ( Claudius Aelianus , d. 235 AD) added certain other embellishments, such as its reputation of "black plumage on its back with

2599-565: A short, rounded head, a reduced neck, and round ears; males have broader heads. The fur varies in colour from light buff to silvery grey, yellowish red, and dark brown. The colours of the underparts are generally lighter. A new-born lion has dark spots , which fade as the cub reaches adulthood, although faint spots may still be seen on the legs and underparts. The tail of all lions ends in a dark, hairy tuft that, in some lions, conceals an approximately 5 mm (0.20 in)-long, hard "spine" or "spur" composed of dermal papillae. The functions of

2712-429: A year old and begin to hunt effectively when nearing the age of two. Single lions are capable of bringing down zebra and wildebeest, while larger prey like buffalo and giraffe are riskier. In Chobe National Park , large prides have been observed hunting African bush elephants up to around 15 years old in exceptional cases, with the victims being calves, juveniles, and even subadults. In typical hunts, each lioness has

2825-404: Is in heat . Lions of both sexes may be involved in group homosexual and courtship activities. Males will also head-rub and roll around with each other before mounting each other. Generation length of the lion is about seven years. The average gestation period is around 110   days; the female gives birth to a litter of between one and four cubs in a secluded den, which may be a thicket,

2938-905: Is a generalist hypercarnivore and is considered to be both an apex and keystone predator due to its wide prey spectrum. Its prey consists mainly of medium-sized to large ungulates , particularly blue wildebeest , plains zebra , African buffalo , gemsbok and giraffe . It also frequently takes common warthog despite it being much smaller. In India, chital and sambar deer are the most common wild prey, while livestock contributes significantly to lion kills outside protected areas. It usually avoids fully grown adult elephants , rhinoceros and hippopotamus and small prey like dik-dik , hyraxes , hares and monkeys . Unusual prey include porcupines and small reptiles. Lions kill other predators but seldom consume them. Young lions first display stalking behaviour at around three months of age, although they do not participate in hunting until they are almost

3051-470: Is a stub . You can help Misplaced Pages by expanding it . Griffin The griffin , griffon , or gryphon ( Ancient Greek : γρύψ , romanized :  grýps ; Classical Latin : grȳps or grȳpus ; Late and Medieval Latin : gryphes , grypho etc.; Old French : griffon ) is a legendary creature with the body, tail, and back legs of a lion , and the head and wings of an eagle with its talons on

3164-543: Is a mixture of dry savannah forest and very dry, deciduous scrub forest . In Africa, the range of the lion originally spanned most of the central African rainforest zone and the Sahara desert. In the 1960s, it became extinct in North Africa, except in the southern part of Sudan. During the mid- Holocene , around 8,000-6,000 years ago, the range of lions expanded into Southeastern and Eastern Europe, partially re-occupying

3277-924: Is affected. Unlike in other felid species, female lions consistently interact with multiple males at once. Another hypothesis suggests that the mane also serves to protect the neck in fights, but this is disputed. During fights, including those involving maneless females and adolescents, the neck is not targeted as much as the face, back, and hindquarters. Injured lions also begin to lose their manes. Almost all male lions in Pendjari National Park are either maneless or have very short manes. Maneless lions have also been reported in Senegal, in Sudan 's Dinder National Park and in Tsavo East National Park , Kenya. Castrated lions often have little to no mane because

3390-465: Is an apex and keystone predator ; although some lions scavenge when opportunities occur and have been known to hunt humans , lions typically do not actively seek out and prey on humans. The lion inhabits grasslands , savannahs , and shrublands . It is usually more diurnal than other wild cats, but when persecuted, it adapts to being active at night and at twilight . During the Neolithic period,

3503-522: Is described as having a " visor " (i.e., beaks) made by Urartian craftsmen, similar to what is found on Greek protomes. Representations of griffin-like hybrids with four legs and a beaked head appeared in Ancient Egyptian art dating back to before 3000 BC. The oldest known depiction of a griffin-like animal in Egypt appears as a relief carving on slate on the cosmetic palette from Hierakonpolis ,

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3616-399: Is generally mutual and the recipient appears to express pleasure. The head and neck are the most common parts of the body licked; this behaviour may have arisen out of utility because lions cannot lick these areas themselves. Lions have an array of facial expressions and body postures that serve as visual gestures. A common facial expression is the "grimace face" or flehmen response , which

3729-521: Is no evidence for gene flow between the two lineages, indicating that they did not share the same geographic area. The Eurasian and American cave lions became extinct at the end of the last glacial period without mitochondrial descendants on other continents. The modern lion was probably widely distributed in Africa during the Middle Pleistocene and started to diverge in sub-Saharan Africa during

3842-459: Is not uniquely applied to the griffin beast, and tštš ( Teš-teš ) has also been used to denote the god Osiris elsewhere. Most statuary representations of griffins depict them with bird-like forelegs and talons , although in some older illustrations griffins have a lion's forelegs (see bronze figure, right); they generally have a lion's hindquarters. Its eagle's head is conventionally given prominent ears ; these are sometimes described as

3955-415: Is short and powerful; it attempts to catch prey with a fast rush and final leap, usually pulls it down by the rump, and kills with a clamping bite to the throat or muzzle . It can hold the prey's throat for up to 13 minutes, until the prey stops moving. It has a bite force from 1593.8 to 1768 Newtons at the canine tip and up 4167.6 Newtons at the carnassial notch. Lions typically consume prey at

4068-604: Is that these derive from the bumps (furrows) on a lion's snout. Another view regards the wart as deriving from the bumpy cockscomb on a rooster or other such fowls. Griffin-like animals were depicted on cylinder seals in Mesopotamia c. 3000 BC, perhaps as early as the Uruk period (4000–3100BC) and subsequent Proto-Elamite ( Jemdet Nasr ) period. An example of a winged lion with beaks, unearthed in Susa (cf. fig. right ) dates to

4181-586: Is the Tsavo lion pride that always has just one adult male. Prides act as fission–fusion societies , and members will split into subgroups that keep in contact with roars . Nomadic lions range widely and move around sporadically, either in pairs or alone. Pairs are more frequent among related males. A lion may switch lifestyles; nomads can become residents and vice versa. Interactions between prides and nomads tend to be hostile, although pride females in estrus allow nomadic males to approach them. Males spend years in

4294-615: Is the most social of all wild felid species, living in groups of related individuals with their offspring. Such a group is called a " pride ". Groups of male lions are called "coalitions". Females form the stable social unit in a pride and do not tolerate outside females. The majority of females remain in their birth prides while all males and some females will disperse . The average pride consists of around 15 lions, including several adult females and up to four males and their cubs of both sexes. Large prides, consisting of up to 30 individuals, have been observed. The sole exception to this pattern

4407-475: Is typically brownish and tinged with yellow, rust, and black hairs. Mutations in the genes microphthalmia-associated transcription factor and tyrosinase are possibly responsible for the colour of manes. It starts growing when lions enter adolescence, when testosterone levels increase, and reach their full size at around four years old. Cool ambient temperatures in European and North American zoos may result in

4520-426: Is unable to consume all of the kill, it rests for a few hours before continuing to eat. On hot days, the pride retreats to shade with one or two males standing guard. Lions defend their kills from scavengers such as vultures and hyenas. Lions scavenge on carrion when the opportunity arises, scavenging animals dead from natural causes such as disease or those that were killed by other predators. Scavenging lions keep

4633-537: The Achaemenian Persian Empire . Russian jewelry historian Elena Neva maintained that the Achaemenids considered the griffin "a protector from evil, witchcraft, and secret slander", but no writings exist from Achaemenid Persia to support her claim. R.L. Fox (1973) remarks that a "lion-griffin" attacks a stag in a pebble mosaic at Pella , from the 4th century BC, perhaps serving as an emblem of

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4746-503: The Lamassu , an Assyrian protective deity, often depicted with a bull or lion's body, eagle's wings, and human's head. Sumerian and Akkadian mythology feature the demon Anzu , half man and half bird, associated with the chief sky god Enlil . This was a divine storm-bird linked with the southern wind and the thunder clouds. Jewish mythology speaks of the Ziz , which resembles Anzu, as well as

4859-505: The Narmada River in central India. Lions spend much of their time resting; they are inactive for about twenty hours per day. Although lions can be active at any time, their activity generally peaks after dusk with a period of socialising, grooming, and defecating. Intermittent bursts of activity continue until dawn, when hunting most often takes place. They spend an average of two hours a day walking and fifty minutes eating. The lion

4972-478: The Nile Basin during the early Holocene . In zoos, lions have been bred with tigers to create hybrids for the curiosity of visitors or for scientific purpose. The liger is bigger than a lion and a tiger, whereas most tigons are relatively small compared to their parents because of reciprocal gene effects. The leopon is a hybrid between a lion and leopard. The lion is a muscular, broad-chested cat with

5085-880: The Two Dog Palette dated to the Early Dynastic Period , c.  3300–3100  BC. Griffin-type creatures combining raptor heads and mammalian bodies were depicted in the Levant , Syria , and Anatolia during the Middle Bronze Age , dated at about 1950–1550 BC. Griffin-type animals appeared in the art of ancient Crete in the MM III Period (1650–1600 BC) in Minoan chronology , found on sealings from Zakro and miniature frescos dated to this period. One early example of griffin-types in Minoan art occurs in

5198-582: The "curled tresses" that are the signature of Uratrian workmanship. Even the ornate crests on Minoan griffins (such as the fresco of the Throne Room, figure top of page) may be a development of these curled tresses. One prominent characteristic of the cauldron griffins is the "top-knob between the brows" (seemingly situated at the top of the head ). The top-knob feature has clear oriental origins. Jack Leonard Benson says these appendages were "topknots" subsequently rendered as "knobs" in later development of

5311-620: The 1200 the area was called Parione e S. Lorenzo in Damaso and the population kept on increasing until the 15th century, when the borough obtained a great importance thanks to the paving of Campo de' Fiori , that soon became an important economic center. Under Sixtus IV (1471–1484) the rione lost its chaotic look, typical of the Middle Ages , for a cleaner and tidier one, typical of the Renaissance . Recovering buildings, enlarging streets, building

5424-571: The 15th century BC frescoes of the Throne Room of the Bronze Age Palace of Knossos , as restored by Sir Arthur Evans . The griffin-like hybrid became a fixture of Aegean culture since the Late Bronze Age , but the animal called the gryps, gryphon, or griffin in Greek writings did not appear in Greek art until about 700 BC, or rather, it was "rediscovered" as artistic motif in

5537-699: The 4th millennium B.C., and is a unique example of a griffin-like animal with a male lion's mane . However, this monster then ceased to continue to be expressed after the Elamite culture. What the Sumerians of the Early Dynastic period portrayed instead were winged lions, and the lion-headed eagle ( Imdugud ). In the Akkadian Empire that succeeded Sumer, early examples (from early 3rd millennium BC ) of lions with bird heads appeared on cylinder seals, shown pulling

5650-609: The 8th to 7th centuries BC, adapting the style of griffin current in Neo-Hittite art. It became quite popular in the 6th and 5th centuries BC, when the Greeks first began to record accounts of the "gryps" creature from travelers to Asia, such as Aristeas of Proconnesus. A number of bronze griffin protomes on cauldrons have been unearthed in Greece (on Samos , and at Olympia , etc., cf. fig. right). Early Greek and early Etruscan (e.g.

5763-749: The Asiatic lion population. In 2017, the Cat Classification Task Force of the Cat Specialist Group revised lion taxonomy , and recognises two subspecies based on results of several phylogeographic studies on lion evolution , namely: However, there seems to be some degree of overlap between both groups in northern Central Africa. DNA analysis from a more recent study indicates that Central African lions are derived from both northern and southern lions, as they cluster with P. leo leo in mtDNA-based phylogenies whereas their genomic DNA indicates

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5876-526: The Barberini) examples of cauldron-griffins may have been of Syric-Urartian make, based on evidence (the "tendrils" or "tresses" motif was already touched upon, above), but "Vannic (Urartian) originals" have yet to be found (in the Orient). It has thus been controversially argued (by Ulf Jantzen  [ de ] ) that these attachments had always since the earliest times been crafted by Greek workshops, added to

5989-609: The Late Pleistocene. Lion populations in East and Southern Africa became separated from populations in West and North Africa when the equatorial rainforest expanded 183,500 to 81,800 years ago. They shared a common ancestor probably between 98,000 and 52,000 years ago. Due to the expansion of the Sahara between 83,100 and 26,600 years ago, lion populations in West and North Africa became separated. As

6102-740: The Mesopotamian lion-griffin. There is also the Armenian term Paskuč ( Armenian : պասկուչ ) that had been used to translate Greek gryp 'griffin' in the Septuagint , which H. P. Schmidt characterized as the counterpart of the simurgh. However, the cognate term Baškuč (glossed as 'griffin') also occurs in Middle Persian, attested in the Zoroastrian cosmological text Bundahishn XXIV (supposedly distinguishable from Sēnmurw which also appears in

6215-558: The Ngorongoro Crater were afflicted by an outbreak of stable fly ( Stomoxys calcitrans ) in 1962, resulting in lions becoming emaciated and covered in bloody, bare patches. Lions sought unsuccessfully to evade the biting flies by climbing trees or crawling into hyena burrows; many died or migrated and the local population dropped from 70 to 15 individuals. A more recent outbreak in 2001 killed six lions. Captive lions have been infected with canine distemper virus (CDV) since at least

6328-691: The adjacent Timbavati Private Game Reserve in eastern South Africa. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool . Nevertheless, 17 births have been recorded in five prides between 2007 and 2015. White lions are selected for breeding in captivity. They have reportedly been bred in camps in South Africa for use as trophies to be killed during canned hunts . African lions live in scattered populations across sub-Saharan Africa. The lion prefers grassy plains and savannahs , scrub bordering rivers, and open woodlands with bushes. It rarely enters closed forests. On Mount Elgon ,

6441-424: The age of two. When a new male lion takes over a pride, adolescents both male and female may be evicted. Lions may live 12–17 years in the wild. Although adult lions have no natural predators, evidence suggests most die violently from attacks by humans or other lions. Lions often inflict serious injuries on members of other prides they encounter in territorial disputes or members of the home pride when fighting at

6554-661: The ancient Greek Phoenix . The Bible mentions the Ziz in Psalms 50:11. This is also similar to a cherub . The cherub, or sphinx, was very popular in Phoenician iconography. In ancient Crete, griffins became very popular, and were portrayed in various media. A similar creature is the Minoan Genius . In the Hindu religion, Garuda is a large bird-like creature that serves as a mount ( vahana ) of

6667-544: The animal's expressive movements are highly developed. The most common peaceful, tactile gestures are head rubbing and social licking , which have been compared with the role of allogrooming among primates. Head rubbing, nuzzling the forehead, face and neck against another lion appears to be a form of greeting and is seen often after an animal has been apart from others or after a fight or confrontation. Males tend to rub other males, while cubs and females rub females. Social licking often occurs in tandem with head rubbing; it

6780-459: The ant into his description of griffins. Later, Pliny the Elder became the first to explicitly state the griffins as having wings and long ears. In one of the two passages, Pliny also located the "griffons" in Æthiopia . According to Adrienne Mayor , Pliny also wrote, "griffins were said to lay eggs in burrows on the ground and these nests contained gold nuggets ". Apollonius of Tyana , who

6893-499: The attack. They take advantage of factors that reduce visibility; many kills take place near some form of cover or at night. One study in 2018 recorded a lion running at a top speed of 74.1 km/h (46.0 mph). The lion accelerates at the start of the chase by 9.5 m/s² , whereas zebras, wildebeest and Thomson's gazelle accelerate by 5 m/s², 5.6 m/s² and 4.5 m/s², respectively; acceleration appears to be more important than steady displacement speed in lion hunts. The lion's attack

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7006-485: The boundary being marked by Via dei Banchi Vecchi, Via del Pellegrino, Via dei Cappellari, Campo de' Fiori , Via dei Giubbonari. Eastward, Parione borders with Sant'Eustachio (R. VIII), from which is separated by Via dei Chiavari, Largo dei Chiavari, Corso Vittorio Emanuele II , Corso del Rinascimento and Piazza delle Cinque Lune. 41°53′55″N 12°28′09″E  /  41.8985°N 12.4691°E  / 41.8985; 12.4691 This Lazio location article

7119-400: The cauldron Griffins. Benson's emphasis is that the Greeks attached a stylized "anorganic" topknot or an "inorganic" plug on the griffin's head (due to lack of information), while in contrast, a known oriental example (stone protomes from Nimrud ) is simple but more "plausible" (naturalistic), resembling a forelock. A cluster of "warts" between the eyes are also mentioned. One conjecture

7232-454: The cause of the decline is not fully understood, habitat loss and conflicts with humans are the greatest causes for concern. One of the most widely recognised animal symbols in human culture, the lion has been extensively depicted in sculptures and paintings, on national flags, and in literature and films. Lions have been kept in menageries since the time of the Roman Empire and have been

7345-522: The chariots for its rider, the weather god. The "lion-griffin" on Akkadian seals are also shown as fire-belching, and shaggy (at the neck) in particular examples. The bronzeworks of Luristan , the North and North West region of Iran in the Iron Age , include examples of Achaemenid art depicting both the "bird-griffin" and "lion-griffin" designs, such as are found on horse-bits . Bernard Goldman maintains

7458-411: The cubs varies—one male could patiently let the cubs play with his tail or his mane, while another may snarl and bat the cubs away. Pride lionesses often synchronise their reproductive cycles and communal rearing and suckling of the young, which suckle indiscriminately from any or all of the nursing females in the pride. The synchronisation of births is advantageous because the cubs grow to being roughly

7571-469: The deity Vishnu . It is also the name for the constellation Aquila . Local lore on the gryps or griffin was gathered by Aristeas of Proconnesus, a Greek who traveled to the Altai region between Mongolia and NW China in the 7th century BC. Although Aristeas's original poem was lost, the gryps lore preserved in secondhand accounts by the playwright Aeschylus (ca. 460 BC), and later his contemporary, Herodotus

7684-617: The front legs. Because the lion was traditionally considered the king of the beasts, and the eagle the king of the birds, by the Middle Ages, the griffin was thought to be an especially powerful and majestic creature. Since classical antiquity, griffins were known for guarding treasures and priceless possessions. In Greek and Roman texts, griffins and Arimaspians were associated with gold deposits of Central Asia. The earliest classical writings derive from Aristeas (7th cent. BC), preserved by Herodotus and Aeschylus (mid 5th century BC), but

7797-516: The god Apollo due to "syncretism between the two gods". At the Temple of Hera at Samos , a griffin-themed bronze "wine-cup" or "cauldron" had been installed, according to Herodotus. The vessel was attached griffin heads around the rim (like the protomes , described above): it was an Argolic or Argive krater , according to the text, standing on a tripod shaped like colossal figures. The notion that griffins lay stones or agate instead of eggs

7910-531: The gold which the griffins collected from various areas in the periphery (presumably including the Armaspi's territorial stream, the stream of Pluto "rolling with gold"). The equestrian Arimaspi would ride off with the loot, and the griffins would give pursuit. Aeschylus likened the gryps to "silent hounds of Zeus" That they are called dogs or hounds here has led to the conjecture that Aeschylus considered them wingless or flightless. Whereas Ctesias , had located

8023-450: The griffin's neck, carven on some of the Greek protomes. The tendril motif emerged at the beginning of the first millennium, BC., in various parts of the Orient. The "double spiral of hair running downwards from the base of the ear" is said to be a hallmark of Iranian (Uratrian) art. The Etruscan cauldron-griffins (e.g., from Barberini tomb  [ it ] , figure right ) also bear

8136-574: The griffins in India, and more explicitly classed them as beaked, four-legged birds. Herodotus also mentions elsewhere that there are gold-collecting ants in Kashmir , India, and this has been interpreted by modern scholars as "doublets or garbled versions" of the lore of gold-hoarding griffins. It appears that the accounts of griffins given by Pliny had been admixed with the lore of these gold-guarding ants of India, and later Aelian also inserted attributes of

8249-534: The group. Alternatively, there may be rewards associated with being a leader that fends off intruders; the rank of lionesses in the pride is reflected in these responses. The male or males associated with the pride must defend their relationship with the pride from outside males who may attempt to usurp them. Dominance hierarchies do not appear to exist among individuals of either sex in a pride. Asiatic lion prides differ in group composition. Male Asiatic lions are solitary or associate with up to three males, forming

8362-504: The historian. Herodotus explains (via Aristeas) that the gold-guarding griffin supposedly dwelled further north from the one-eyed Arimaspi people who robbed the gold from the fabulous creatures. Aristeas is said to have been informed through the Issedones people neighboring region to the Arimaspi, in the northern extremes (of Central Asia). Aeschylus also concurs that the Arimaspi robbed

8475-570: The hunting of large, slower-moving prey like buffalo; and even hunt them on their own. Moderately-sized hunting groups generally have higher success rates than lone females and larger groups. Lions are not particularly known for their stamina. For instance, a lioness's heart comprises only 0.57% of her body weight and a male's is about 0.45% of his body weight, whereas a hyena's heart comprises almost 1% of its body weight. Thus, lions run quickly only in short bursts at about 48–59 km/h (30–37 mph) and need to be close to their prey before starting

8588-528: The identifiable attested "accounts" presented in scholarship are largely not literary, but artistic, or numismatic . The griffin was naturally linked to Apollo, given the existence of the cultus of Hyperborean Apollo , with a cult center at the Greek colony of Olbia on the Black Sea . And even the main Temple of Apollo at Delphi featured a statue of the god flanked by griffins, or so it can be presumed based on

8701-401: The image of a griffin attacking a horse. Other Scythian artifacts show griffins attacking horses, stags, and goats. Griffins are typically shown attacking horses, deer, and humans in Greek art. Nomads were said to steal griffin-guarded gold according to Scythian oral traditions reported by Greek and Roman travelers. Several ancient mythological creatures are similar to the griffin. These include

8814-513: The kingdom of Macedon or a personal emblem of Antipater , one of Alexander 's successors. A golden frontal half of a griffin-like animal from the Ziwiye hoard (near Saqqez city) in Kurdistan Province , Iran resembles the western protomes in style. They were of Urartian workmanship (neither Assyrian or Scythian), though the hoard itself may have represented a Scythian burial. The animal

8927-495: The later lore that griffins deposited agate stone among the eggs in their nest. Pliny placed the griffins in Æthiopia , and Ctesias (5th century BC) in greater India . Scholars have observed that legends about the gold-digging ants of India may have contaminated griffin lore. In the Christian era, Isidore of Seville (7th century AD) wrote that griffins were a great enemy of horses. This notion may have readily developed from

9040-705: The lion has been recorded up to an elevation of 3,600 m (11,800 ft) and close to the snow line on Mount Kenya . Savannahs with an annual rainfall of 300 to 1,500 mm (12 to 59 in) make up the majority of lion habitat in Africa, estimated at 3,390,821 km (1,309,203 sq mi) at most, but remnant populations are also present in tropical moist forests in West Africa and montane forests in East Africa. The Asiatic lion now survives only in and around Gir National Park in Gujarat, western India. Its habitat

9153-801: The lion population in Kenya's Masai Mara National Reserve declined, the spotted hyena population increased rapidly. Lions tend to dominate cheetahs and leopards, steal their kills and kill their cubs and even adults when given the chance. Cheetahs often lose their kills to lions or other predators. A study in the Serengeti ecosystem revealed that lions killed at least 17 of 125 cheetah cubs born between 1987 and 1990. Cheetahs avoid their competitors by hunting at different times and habitats. Leopards, by contrast, do not appear to be motivated by an avoidance of lions, as they use heavy vegetation regardless of whether lions are present in an area and both cats are active around

9266-532: The lion ranged throughout Africa and Eurasia, from Southeast Europe to India, but it has been reduced to fragmented populations in sub-Saharan Africa and one population in western India . It has been listed as Vulnerable on the IUCN Red List since 1996 because populations in African countries have declined by about 43% since the early 1990s. Lion populations are untenable outside designated protected areas. Although

9379-562: The lion's ears, but are often elongated (more like a horse 's), and are sometimes feathered. The griffin of Greece, as depicted in cast bronze cauldron protomes (cf. below), has a squat face with short beaks that are open agape as if screaming, with the tongue showing. There is also a "top-knob" on its head or between the brows. There may also be so-called "tendrils", or curled "spiral-locks" depicted, presumably representing either hair/mane or feather/crest locks dangling down. Single- or double-streaked tendrils hang down both sides and behind

9492-551: The location of the hunt but sometimes drag large prey into cover. They tend to squabble over kills, particularly the males. Cubs suffer most when food is scarce but otherwise all pride members eat their fill, including old and crippled lions, which can live on leftovers. Large kills are shared more widely among pride members. An adult lioness requires an average of about 5 kg (11 lb) of meat per day while males require about 7 kg (15 lb). Lions gorge themselves and eat up to 30 kg (66 lb) in one session. If it

9605-481: The mid-1970s. CDV is spread by domestic dogs and other carnivores; a 1994 outbreak in Serengeti National Park resulted in many lions developing neurological symptoms such as seizures. During the outbreak, several lions died from pneumonia and encephalitis . Feline immunodeficiency virus and lentivirus also affect captive lions. When resting, lion socialisation occurs through a number of behaviours;

9718-471: The modern Persian language , the griffin has come to be called šērdāl ( Persian : شیردال ), meaning 'lion-eagle'. However, the practice of referring to ancient Iranian griffin objects or monuments as sherdal , is not followed by other current archaeological scholarship (e.g., here ). Possible Old or Middle Iranian names for the creature have been discussed. Middle Persian Sēnmurw in Sasanian culture

9831-457: The new bridge Ponte Sisto connecting Trastevere and Parione, improved the quality of the area. Thanks to this renewal, urbanisation increased between the 15th and the 16th century. In the same period, several artists were asked to renew the front of the greatest buildings: this habit was born in the northern Italy and was spreading in that period. In 1500 most of the commercial activity slowly moved from Campo de' Fiori to Piazza Navona , that

9944-434: The phylogenetic relationship of the lion; it was thought to form a sister group with the jaguar that diverged 3.46 to 1.22 million years ago , but also with the leopard that diverged 3.1 to 1.95 million years ago to 4.32 to 0.02 million years ago . Hybridisation between lion and snow leopard ancestors possibly continued until about 2.1 million years ago. The lion-leopard clade

10057-560: The physical descriptions are not very explicit. Thus even though they are sharp-beaked, their being likened to "unbarking hounds of Zeus" has led to the speculation they were seen as wingless. Pliny the Elder (1st century) was the first to explicitly state that griffins were winged and long eared. But Apollonius of Tyana wrote that griffins did not have true bird wings, but only membranous webbed feet that only gave them capability of short-distanced flight. Writers after Aelian (3rd century AD) did not add new material to griffin lore, except for

10170-667: The plain cauldrons imported from the Near East. Detractors (notably K. R. Maxwell-Hyslop ) believe that (early examples of ) the griffin-ornamented cauldron, in its entirely, were crafted in the East, though excavated finds from the Orient are scarce. In Central Asia , the griffin image was included in Scythian "animal style" artifacts of the 6th–4th centuries BC, but no writings explain their meaning. The Golden Pectoral from Tovsta Mohyla , interred in Scythian king's burial site, perhaps commissioned to Greek goldsmiths , who engraved

10283-445: The position that Luristan examples must be counted as developments of the "lion-griffin" type, even when it exhibits "stylization .. approaching the beak of a bird". The Luristan griffin-like creatures resemble and perhaps are descended from Assyrian creatures, possibly influenced by Mitannian animals, or perhaps there had been parallel development in both Assyrian and Elamite cultures. Bird-headed mammal images appeared in art of

10396-612: The previous males associated with a pride, the victors often kill any existing young cubs , perhaps because females do not become fertile and receptive until their cubs mature or die. Females often fiercely defend their cubs from a usurping male but are rarely successful unless a group of three or four mothers within a pride join forces against the male. Cubs also die from starvation and abandonment, and predation by leopards, hyenas and wild dogs. Male cubs are excluded from their maternal pride when they reach maturity at around two or three years of age, while some females may leave when they reach

10509-579: The rainforest decreased and thus gave rise to more open habitats, lions moved from West to Central Africa. Lions from North Africa dispersed to southern Europe and Asia between 38,800 and 8,300 years ago. Extinction of lions in southern Europe, North Africa and the Middle East interrupted gene flow between lion populations in Asia and Africa. Genetic evidence revealed numerous mutations in lion samples from East and Southern Africa, which indicates that this group has

10622-457: The range of the now extinct cave lion . In Hungary , the modern lion was present from about 4,500 to 3,200 years Before Present . In Ukraine , the modern lion was present from about 6,400 to 2,000 years Before Present. In Greece, it was common, as reported by Herodotus in 480 BC; it was considered rare by 300 BC and extirpated by AD 100. In Asia the lion once ranged in regions where climatic conditions supported an abundance of prey. It

10735-488: The removal of the gonads inhibits testosterone production. Rarely, both wild and captive lionesses have manes. Increased testosterone may be the cause of maned lionesses reported in northern Botswana. The white lion is a rare morph with a genetic condition called leucism , which is caused by a double recessive allele . It is not albino; it has normal pigmentation in the eyes and skin. White lions have occasionally been encountered in and around Kruger National Park and

10848-405: The representation struck on the tetradrachm coinage of Attica. Apollo rode a griffin to Hyperboria each winter, leaving Delphi, or so it was believed. Apollo riding griffin is known from multiple examples of red-figure pottery . And Apollo hitched griffins to his chariot according to Claudian . Dionysus was also depicted on a griffin-chariot or mounting griffin; the motif was borrowed from

10961-429: The same area. The evolution of sociability in lions was likely driven both by high population density and the clumped resources of savannah habitats. The larger the pride, the more high-quality territory they can defend; "hotspots" being near river confluences , where the cats have better access to water, prey and shelter (via vegetation). The area occupied by a pride is called a "pride area" whereas that occupied by

11074-420: The same size and have an equal chance of survival, and sucklings are not dominated by older cubs. Weaning occurs after six or seven months. Male lions reach maturity at about three years of age and at four to five years are capable of challenging and displacing adult males associated with another pride. They begin to age and weaken at between 10 and 15 years of age at the latest. When one or more new males oust

11187-570: The same text). Middle Persian Paškuč is also attested in Manichaean magical texts (Manichaean Middle Persian: pškwc ), and this must have meant a "griffin or a monster like a griffin" according to W. B. Henning . The griffin was given names which were descriptive epithets, such as tštš or tesh-tesh meaning "Tearer[-in-pieces]" inscribed on a griffin image found in a tomb at Deir El Bersha ; and sfr / srf "fiery one", attested at Beni Hasan . The descriptive epithet "Tearer"

11300-406: The same time of day. In addition, there is no evidence that lions affect leopard abundance. Leopards take refuge in trees, though lionesses occasionally attempt to climb up and retrieve their kills. Lions similarly dominate African wild dogs , taking their kills and dispatching pups or adult dogs. Population densities of wild dogs are low in areas where lions are more abundant. However, there are

11413-420: The same time. When first introduced to the rest of the pride, lion cubs lack confidence when confronted with adults other than their mother. They soon begin to immerse themselves in the pride life, however, playing among themselves or attempting to initiate play with the adults. Lionesses with cubs of their own are more likely to be tolerant of another lioness's cubs than lionesses without cubs. Male tolerance of

11526-422: The spur are unknown. The tuft is absent at birth and develops at around 5 + 1 ⁄ 2 months of age. It is readily identifiable at the age of seven months. Its skull is very similar to that of the tiger, although the frontal region is usually more depressed and flattened and has a slightly shorter postorbital region and broader nasal openings than those of the tiger. Due to the amount of skull variation in

11639-413: The tiger in size. The size and weight of adult lions vary across its range and habitats. Accounts of a few individuals that were larger than average exist from Africa and India. The male lion's mane is the most recognisable feature of the species. It may have evolved around 320,000–190,000 years ago. It grows downwards and backwards, covering most of the head, neck, shoulders, and chest. The mane

11752-465: The tradition that horseback-riding Arimaspians raided the griffin gold. The derivation of this word remains uncertain. It could be related to the Greek word γρυπός (grypos), meaning 'curved', or 'hooked'. Greek γρύφ (gryph) from γρύφ 'hook-nosed' is suggested. It could also have been an Anatolian loan word derived from a Semitic language; compare the Hebrew word for cherub כרוב kərúv . In

11865-407: The two species, usually only the structure of the lower jaw can be used as a reliable indicator of species. The skeletal muscles of the lion make up 58.8% of its body weight and represent the highest percentage of muscles among mammals. The lion has a high concentration of fast twitch muscle fibers, giving them quick bursts of speed but less stamina. Among felids, the lion is second only to

11978-400: Was a borrowing from Ancient Greek λέων léōn . The Hebrew word לָבִיא lavi may also be related. The generic name Panthera is traceable to the classical Latin word 'panthēra' and the ancient Greek word πάνθηρ 'panther'. Felis leo was the scientific name used by Carl Linnaeus in 1758, who described the lion in his work Systema Naturae . The genus name Panthera

12091-447: Was a fabulous composite creature, and Russian archaeologist Boris A. Litvinskij  [ ru ] argued for the possibility that the application of this term may extend to the griffin. The term Sēnmurw is recognized as the etymological ancestor of simurgh , which is generally regarded as a mythological bird (rather than a composite) in later medieval Persian literature, though some argue that this bird may have originated from

12204-593: Was coined by Lorenz Oken in 1816. Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished mostly by the size and colour of their manes and skins. In the 19th and 20th centuries, several lion type specimens were described and proposed as subspecies , with about a dozen recognised as valid taxa until 2017. Between 2008 and 2016, IUCN Red List assessors used only two subspecific names: P. l. leo for African lion populations, and P. l. persica for

12317-511: Was distributed in the Asian and African Palearctic since at least the early Pliocene . The earliest fossils recognisable as lions were found at Olduvai Gorge in Tanzania and are estimated to be up to 2 million years old. Estimates for the divergence time of the modern and cave lion lineages range from 529,000 to 392,000 years ago based on mutation rate per generation time of the modern lion. There

12430-416: Was introduced "at some in the evolution of griffin lore". Albertus Magnus (d. 1280) attributes to other writers the claim that "this bird places an ' eagle-stone ' ( echytem ) or agate ( gagatem ) among its eggs" to change the ambient temperature and enhance reproduction. Lion The lion ( Panthera leo ) is a large cat of the genus Panthera , native to Africa and India . It has

12543-596: Was nearly coeval with Pliny, gave a somewhat unique account of the griffin, claiming them to be lion-sized, and having no true wings, and instead had paws "webbed with red membranes", that gave them ability to makes leaps of flight of only a short distance. Pomponius Mela (fl. AD 43) wrote in his Book ii. 6: In Europe, constantly falling snow makes those places contiguous with the Riphaean Mountains .. so impassable that, in addition, they prevent those who deliberately travel here from seeing anything. After that comes

12656-857: Was present in the Caucasus until the 10th century. It lived in Palestine until the Middle Ages and in Southwest Asia until the late 19th century. By the late 19th century, it had been extirpated in most of Turkey. The last live lion in Iran was sighted in 1942, about 65 km (40 mi) northwest of Dezful , although the corpse of a lioness was found on the banks of the Karun river in Khuzestan province in 1944. It once ranged from Sind and Punjab in Pakistan to Bengal and

12769-414: Was the favorite place since it was wider. From 1574 to 1674, the appearance of Piazza Navona changed vastly, thanks to the work of Bernini , Borromini , and Bramante . From this period until Rome became capital of unified Italy in 1870 there were no major changes but the opening of Corso Vittorio Emanuele II , a great street having a flexuous track in order to avoid the already existing palaces. If

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