140-416: The giant panda ( Ailuropoda melanoleuca ), also known as the panda bear or simply panda , is a bear species endemic to China . It is characterised by its white coat with black patches around the eyes, ears, legs and shoulders. Its body is rotund; adult individuals weigh 100 to 115 kg (220 to 254 lb) and are typically 1.2 to 1.9 m (3 ft 11 in to 6 ft 3 in) long. It
280-481: A WWF lawsuit , the United States Fish and Wildlife Service only allows US zoos to import a panda if the zoo can ensure China channels more than half of its loan fee into conservation efforts for giant pandas and their habitat. In May 2005, China offered a breeding pair to Taiwan . The issue became embroiled in cross-Strait relations – due to both the underlying symbolism and technical issues such as whether
420-499: A carnivoran , the giant panda's diet is primarily herbivorous , with approximately 99% of its diet consisting of bamboo. However, the giant panda still has the digestive system of a carnivore, as well as carnivore-specific genes, and thus derives little energy and little protein from the consumption of bamboo. The ability to break down cellulose and lignin is very weak, and their main source of nutrients comes from starch and hemicelluloses . The most important part of their bamboo diet
560-469: A basis of morphological and physiological facts as possible, and one in which "place is found for all observational and experimental data relating, even if indirectly, to the constitution, subdivision, origin, and behaviour of species and other taxonomic groups". Ideals can, it may be said, never be completely realized. They have, however, a great value of acting as permanent stimulants, and if we have some, even vague, ideal of an "omega" taxonomy we may progress
700-527: A biteforce of 1298.9 Newton (BFQ 151.4) at canine teeth and 1815.9 Newton (BFQ 141.8) at carnassial teeth. Adults measure around 1.2 to 1.9 m (3 ft 11 in to 6 ft 3 in) long, including a tail of about 10–15 cm (3.9–5.9 in), and 60 to 90 cm (24 to 35 in) tall at the shoulder . Males can weigh up to 160 kg (350 lb). Females are generally 10–20% smaller than males. They weigh between 70 kg (150 lb) and 125 kg (276 lb). The average weight for adults
840-724: A blatting sound; they achieve peaceful interactions through the emission of this sound. When in oestrus, a female emits a chirp. In hostile confrontations or during fights, the giant panda emits vocalizations such as a roar or growl. On the other hand, squeals typically indicate inferiority and submission in a dispute. Other vocalizations include honks and moans. Giant pandas heavily rely on olfactory communication to communicate with one another. Scent marks are used to spread these chemical cues and are placed on landmarks like rocks or trees. Chemical communication in giant pandas plays many roles in their social situations. Scent marks and odors are used to spread information about sexual status, whether
980-552: A chaotic and disorganized taxonomic literature. He not only introduced the standard of class, order, genus, and species, but also made it possible to identify plants and animals from his book, by using the smaller parts of the flower (known as the Linnaean system ). Plant and animal taxonomists regard Linnaeus' work as the "starting point" for valid names (at 1753 and 1758 respectively). Names published before these dates are referred to as "pre-Linnaean", and not considered valid (with
1120-429: A day through its diet. To prevent poisoning, they have evolved anti-toxic mechanisms to protect themselves. About 80% of the cyanide is metabolized to less toxic thiocyanate and discharged in urine, while the remaining 20% is detoxified by other minor pathways. During the shoot season (April–August), pandas store a large amount of food in preparation for the months succeeding this seasonal period, in which pandas live off
1260-465: A diet of bamboo leaves. The giant panda is a highly specialised animal with unique adaptations, and has lived in bamboo forests for millions of years. The average giant panda eats as much as 9 to 14 kg (20 to 31 lb) of bamboo shoots a day to compensate for the limited energy content of its diet. Ingestion of such a large quantity of material is possible and necessary because of the rapid passage of large amounts of indigestible plant material through
1400-443: A different sense, to mean the delimitation of species (not subspecies or taxa of other ranks), using whatever investigative techniques are available, and including sophisticated computational or laboratory techniques. Thus, Ernst Mayr in 1968 defined " beta taxonomy " as the classification of ranks higher than species. An understanding of the biological meaning of variation and of the evolutionary origin of groups of related species
1540-445: A disease caused by an obligate intracellular parasitic protozoan known as Toxoplasma gondii that infects most warm-blooded animals, including humans. They are likely susceptible to diseases from Baylisascaris schroederi , a parasitic nematode known to infect giant panda intestines. This nematode species is known to give pandas baylisascariasi , a deadly disease that kills more wild pandas than any other cause. Additionally,
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#17328455484111680-593: A fee of up to US$ 1,000,000 per year and with the provision that any cubs born during the loan are the property of China. As a result of this change in policy, nearly all the pandas in the world are owned by China, and pandas leased to foreign zoos and all cubs are eventually returned to China. As of 2022, Xin Xin at the Chapultepec Zoo in Mexico City, was the last living descendant of the gifted pandas. Since 1998, because of
1820-423: A female is in estrus or not, age, gender, individuality, dominance over territory, and choice of settlement. Giant pandas communicate by excreting volatile compounds, or scent marks, through the anogenital gland. Giant pandas have unique positions in which they will scent mark. Males deposit scent marks or urine by lifting their hind leg, rubbing their backside, or standing in order to rub the anogenital gland onto
1960-435: A female when choosing a mate. For example, age can be useful for a female to determine sexual maturity and sperm quality. Pandas are also able to determine when the signal was placed, further aiding in the quest to find a potential mate. However, chemical cues are not just used for communication between males and females, pandas can determine individuality from chemical signals. This allows them to be able to differentiate between
2100-449: A few Chinese provinces at elevations of up to 3,000 m (9,800 ft). It is solitary and gathers only in mating seasons. It relies on olfactory communication to communicate and uses scent marks as chemical cues and on landmarks like rocks or trees. Females rear cubs for an average of 18 to 24 months. The oldest known giant panda was 38 years old. As a result of farming, deforestation and infrastructural development,
2240-420: A fight. Giant pandas reach sexual maturity between the ages of four and eight, and may be reproductive until age 20. The mating season is between March and May, when a female goes into estrus , which lasts for two or three days and only occurs once a year. When mating, the female is in a crouching, head-down position as the male mounts her from behind. Copulation time ranges from 30 seconds to five minutes, but
2380-443: A landmark. Females, however, exercise squatting or simply rubbing their genitals onto a landmark. The season plays a major role in mediating chemical communication. Depending on the season, mainly whether it is breeding season or not, may influence which odors are prioritized. Chemical signals can have different functions in different seasons. During the non-breeding season, females prefer the odors of other females because reproduction
2520-451: A little way down the Greek alphabet. Some of us please ourselves by thinking we are now groping in a "beta" taxonomy. Turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy. Later authors have used the term in
2660-450: A mid-to-high density of livestock, as they depress the vegetation. The Tibetan Plateau is the only known area where both giant and red pandas can be found. Although sharing near-identical ecological niches, competition between the two species has rarely been observed. Nearly 50% of their respective distribution overlaps, and successful coexistence is achieved through distinct habitat selection. A captive female died from toxoplasmosis ,
2800-504: A notable renaissance, principally with respect to theoretical content. Part of the theoretical material has to do with evolutionary areas (topics e and f above), the rest relates especially to the problem of classification. Taxonomy is that part of Systematics concerned with topics (a) to (d) above. A whole set of terms including taxonomy, systematic biology, systematics , scientific classification, biological classification, and phylogenetics have at times had overlapping meanings – sometimes
2940-550: A potential partner or someone of the same sex, which could be a potential competitor. Chemical cues, or odors, play an important role in how a panda chooses their habitat. Pandas look for odors that tell them not only the identity of another panda, but if they should avoid them or not. Pandas tend to avoid their species for most of the year, breeding season being the brief time of major interaction. Chemical signaling allows for avoidance and competition . Pandas whose habitats are in similar locations will collectively leave scent marks in
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#17328455484113080-470: A single continuum, as per the scala naturae (the Natural Ladder). This, as well, was taken into consideration in the great chain of being. Advances were made by scholars such as Procopius , Timotheus of Gaza , Demetrios Pepagomenos , and Thomas Aquinas . Medieval thinkers used abstract philosophical and logical categorizations more suited to abstract philosophy than to pragmatic taxonomy. During
3220-478: A species, the giant panda must have at least two different species available in its range to avoid starvation. While primarily herbivorous, the giant panda still retains decidedly ursine teeth and will eat meat, fish, and eggs when available. In captivity, zoos typically maintain the giant panda's bamboo diet, though some will provide specially formulated biscuits or other dietary supplements. Pandas will travel between different habitats if they need to, so they can get
3360-652: A sub-area of systematics (definition 2), invert that relationship (definition 6), or appear to consider the two terms synonymous. There is some disagreement as to whether biological nomenclature is considered a part of taxonomy (definitions 1 and 2), or a part of systematics outside taxonomy. For example, definition 6 is paired with the following definition of systematics that places nomenclature outside taxonomy: In 1970, Michener et al. defined "systematic biology" and "taxonomy" (terms that are often confused and used interchangeably) in relation to one another as follows: Systematic biology (hereafter called simply systematics)
3500-524: A truly scientific attempt to classify organisms did not occur until the 18th century, with the possible exception of Aristotle, whose works hint at a taxonomy. Earlier works were primarily descriptive and focused on plants that were useful in agriculture or medicine. There are a number of stages in this scientific thinking. Early taxonomy was based on arbitrary criteria, the so-called "artificial systems", including Linnaeus 's system of sexual classification for plants (Linnaeus's 1735 classification of animals
3640-400: A unique location which is termed "scent stations". When pandas come across these scent stations, they are able to identify a specific panda and the scope of their habitat. This allows pandas to be able to pursue a potential mate or avoid a potential competitor. Pandas can assess an individual's dominance status, including their age and size, via odor cues and may choose to avoid a scent mark if
3780-447: Is sexually dimorphic , with males being typically 10 to 20% larger than females. A thumb is visible on its forepaw, which helps in holding bamboo in place for feeding. It has large molar teeth and expanded temporal fossa to meet its dietary requirements. It can digest starch and is mostly herbivorous with a diet consisting almost entirely of bamboo and bamboo shoots . The giant panda lives exclusively in six montane regions in
3920-403: Is 100 to 115 kg (220 to 254 lb). The giant panda's paw has a digit similar to a thumb and five fingers ; the thumb-like digit – actually a modified sesamoid bone – helps it to hold bamboo while eating. The giant panda's tail, measuring 10 to 15 cm (3.9 to 5.9 in), is the second-longest in the bear family, behind the sloth bear . Despite its taxonomic classification as
4060-555: Is a vulnerable species , threatened by continued habitat loss and fragmentation , and by a very low birthrate, both in the wild and in captivity . Its range is confined to a small portion on the western edge of its historical range, which stretched through southern and eastern China, northern Myanmar , and northern Vietnam . The species is scattered into more than 30 subpopulations of relatively few animals. Building of roads and human settlement near panda habitat, result in population declines. Diseases from domesticated pets and livestock
4200-497: Is a critical component of the taxonomic process. As a result, it informs the user as to what the relatives of the taxon are hypothesized to be. Biological classification uses taxonomic ranks, including among others (in order from most inclusive to least inclusive): Domain , Kingdom , Phylum , Class , Order , Family , Genus , Species , and Strain . The "definition" of a taxon is encapsulated by its description or its diagnosis or by both combined. There are no set rules governing
4340-400: Is a novel analysis of the variation patterns in a particular taxon . This analysis may be executed on the basis of any combination of the various available kinds of characters, such as morphological, anatomical , palynological , biochemical and genetic . A monograph or complete revision is a revision that is comprehensive for a taxon for the information given at a particular time, and for
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4480-458: Is a resource for fossils. Biological taxonomy is a sub-discipline of biology , and is generally practiced by biologists known as "taxonomists", though enthusiastic naturalists are also frequently involved in the publication of new taxa. Because taxonomy aims to describe and organize life , the work conducted by taxonomists is essential for the study of biodiversity and the resulting field of conservation biology . Biological classification
4620-405: Is a sex hormone found in both males and females; testosterone is the major androgen produced by males. Civetone and decanoic acid are chemicals found in female urine which promote behavioral responses in males; both chemicals are considered giant panda pheromones. Male pandas also secrete chemical signals that include information about their sexual reproductivity and age, which is beneficial for
4760-454: Is another threat. By 2100, it is estimated that the distribution of giant pandas will shrink by up to 100%, mainly due to the effects of climate change . The giant panda is listed on CITES Appendix I , meaning trade of their parts is prohibited and that they require this protection to avoid extinction . They have been protected and placed in category 1, by the 1988 Wildlife Protection Act . Bear Bears are carnivoran mammals of
4900-419: Is even more important for the second stage of taxonomic activity, the sorting of species into groups of relatives ("taxa") and their arrangement in a hierarchy of higher categories. This activity is what the term classification denotes; it is also referred to as "beta taxonomy". How species should be defined in a particular group of organisms gives rise to practical and theoretical problems that are referred to as
5040-524: Is from the Proto-Indo-European word *ǵʰwḗr- ~ *ǵʰwér "wild animal". This terminology for the animal originated as a taboo avoidance term: proto-Germanic tribes replaced their original word for bear— arkto —with this euphemistic expression out of fear that speaking the animal's true name might cause it to appear. According to author Ralph Keyes , this is the oldest known euphemism . Bear taxon names such as Arctoidea and Helarctos come from
5180-439: Is mainly attributed to the panda's avoidance of interspecific competition . The species has been located at elevations of 2,400 to 3,000 m (7,900 to 9,800 ft) above sea level. They frequent habitats with a healthy concentration of bamboos, typically old-growth forests , but may also venture into secondary forest habitats. The Daxiangling Mountain population inhabits both coniferous and broadleaf forests. Additionally,
5320-491: Is not their primary motivation. However, during breeding season, odors from the opposite sex will be more attractive. Because they are solitary mammals and their breeding season is so brief, female pandas secrete chemical cues in order to let males know their sexual status. The chemical cues female pandas secrete can be considered to be pheromones for sexual reproduction. Females deposit scent marks through their urine which induces an increase in androgen levels in males. Androgen
5460-427: Is similar to other subtropical mammals, and will instead move to elevations with warmer temperatures. Pandas rely primarily on spatial memory rather than visual memory . Though the panda is often assumed to be docile, it has been known to attack humans on rare occasions. Pandas have been known to cover themselves in horse manure to protect themselves against cold temperatures. The species communicates foremost through
5600-428: Is sometimes used in botany in place of phylum ), class , order , family , genus , and species . The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed a ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms. With advances in the theory, data and analytical technology of biological systematics,
5740-449: Is the scientific study of naming, defining ( circumscribing ) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given a taxonomic rank ; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain , kingdom , phylum ( division
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5880-400: Is the field that (a) provides scientific names for organisms, (b) describes them, (c) preserves collections of them, (d) provides classifications for the organisms, keys for their identification, and data on their distributions, (e) investigates their evolutionary histories, and (f) considers their environmental adaptations. This is a field with a long history that in recent years has experienced
6020-403: Is the result of powerful jaw muscles, which attach from the top of the head to the jaw. Large molars crush and grind fibrous plant material. The morphological characteristics of extinct relatives of the giant panda suggest that while the ancient giant panda was omnivorous 7 million years ago (mya), it only became herbivorous some 2–2.4 mya with the emergence of A. microta . Genome sequencing of
6160-577: Is the shoots, that are rich in starch and have up to 32% protein content. Accordingly, pandas have evolved a higher capability to digest starches than strict carnivores. Raw bamboo is toxic, containing cyanide compounds. Pandas' body tissues are less able than herbivores to detoxify cyanide, but their gut microbiomes are significantly enriched in putative genes coding for enzymes related to cyanide degradation, suggesting that they have cyanide-digesting gut microbes. It has been estimated that an adult panda absorbs 54.8–66.1 mg (0.846–1.020 gr) of cyanide
6300-560: The American black bear , a thriving bear species. In July 2009, Chinese scientists confirmed the birth of the first cub to be successfully conceived through artificial insemination using frozen sperm. The technique for freezing the sperm in liquid nitrogen was first developed in 1980 and the first birth was hailed as a solution to the dwindling availability of giant panda semen, which had led to inbreeding. Panda semen, which can be frozen for decades, could be shared between different zoos to save
6440-518: The Aristotelian system , with additions concerning the philosophical and existential order of creatures. This included concepts such as the great chain of being in the Western scholastic tradition, again deriving ultimately from Aristotle. The Aristotelian system did not classify plants or fungi , due to the lack of microscopes at the time, as his ideas were based on arranging the complete world in
6580-639: The Bering land bridge may have been possible during a major sea level low stand as early as the late Eocene (about 37 Mya) and continuing into the early Oligocene. European genera morphologically very similar to Allocyon , and to the much younger American Kolponomos (about 18 Mya), are known from the Oligocene, including Amphicticeps and Amphicynodon . There has been various morphological evidence linking amphicynodontines with pinnipeds , as both groups were semi-aquatic, otter-like mammals. In addition to
6720-576: The Empress Dowager Bo was buried with a panda skull in her vault. The grandson of Emperor Taizong of Tang is said to have given Japan two pandas and a sheet of panda skin as a sign of goodwill. Unlike many other animals in Ancient China , pandas were rarely thought to have medical uses. The few known uses include the Sichuan tribal peoples' use of panda urine to melt accidentally swallowed needles, and
6860-620: The Isthmus of Panama . Their earliest fossil representative is Plionarctos in North America (c. 10–2 Ma). This genus is probably the direct ancestor to the North American short-faced bears (genus Arctodus ), the South American short-faced bears ( Arctotherium ), and the spectacled bears, Tremarctos , represented by both an extinct North American species ( T. floridanus ), and
7000-575: The Neomura , the clade that groups together the Archaea and Eucarya , would have evolved from Bacteria, more precisely from Actinomycetota . His 2004 classification treated the archaeobacteria as part of a subkingdom of the kingdom Bacteria, i.e., he rejected the three-domain system entirely. Stefan Luketa in 2012 proposed a five "dominion" system, adding Prionobiota ( acellular and without nucleic acid ) and Virusobiota (acellular but with nucleic acid) to
7140-476: The Qinling Mountains and in the hilly province of Sichuan . Giant pandas are generally solitary. Each adult has a defined territory and a female is not tolerant of other females in her range. Social encounters occur primarily during the brief breeding season in which pandas in proximity to one another will gather. After mating, the male leaves the female alone to raise the cub. Pandas were thought to fall into
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#17328455484117280-503: The Renaissance and the Age of Enlightenment , categorizing organisms became more prevalent, and taxonomic works became ambitious enough to replace the ancient texts. This is sometimes credited to the development of sophisticated optical lenses, which allowed the morphology of organisms to be studied in much greater detail. One of the earliest authors to take advantage of this leap in technology
7420-680: The Western Han Dynasty in China, where the writer Sima Xiangru noted that the panda was the most treasured animal in the emperor's garden of exotic animals in the capital Chang'an (present Xi'an ). Not until the 1950s were pandas again recorded to have been exhibited in China's zoos. Chi Chi at the London Zoo became very popular. This influenced the World Wildlife Fund to use a panda as its symbol. A 2006 New York Times article outlined
7560-513: The crepuscular category, those who are active twice a day, at dawn and dusk; however, pandas may belong to a category all of their own, with activity peaks in the morning, afternoon and midnight. The low nutrition quality of bamboo means pandas need to eat more frequently, and due to their lack of major predators they can be active at any time of the day. Activity is highest in June and decreases in late summer to autumn with an increase from November through
7700-735: The family Ursidae ( / ˈ ɜːr s ɪ d iː , - d aɪ / ). They are classified as caniforms , or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout most of the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America , South America , and Eurasia . Common characteristics of modern bears include large bodies with stocky legs, long snouts, small rounded ears, shaggy hair, plantigrade paws with five nonretractile claws, and short tails. While
7840-533: The polar bear is mostly carnivorous , and the giant panda is mostly herbivorous , the remaining six species are omnivorous with varying diets. With the exception of courting individuals and mothers with their young, bears are typically solitary animals . They may be diurnal or nocturnal and have an excellent sense of smell . Despite their heavy build and awkward gait, they are adept runners, climbers, and swimmers. Bears use shelters, such as caves and logs, as their dens; most species occupy their dens during
7980-439: The species problem . The scientific work of deciding how to define species has been called microtaxonomy. By extension, macrotaxonomy is the study of groups at the higher taxonomic ranks subgenus and above, or simply in clades that include more than one taxon considered a species, expressed in terms of phylogenetic nomenclature . While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations,
8120-399: The spectacled bear ), and Ursinae (containing six species divided into one to three genera, depending on the authority). Nuclear chromosome analysis show that the karyotype of the six ursine bears is nearly identical, each having 74 chromosomes ( see Ursid hybrid ), whereas the giant panda has 42 chromosomes and the spectacled bear 52. These smaller numbers can be explained by
8260-541: The takin ( Budorcas taxicolor ). The takin and giant panda share a similar ecological niche , and they consume the same resources. When competition for food is fierce, pandas disperse to the outskirts of takin distribution. Other possible competitors include but is not limited to, the Eurasian wild pig ( Sus scrofa ), Chinese goral ( Naemorhedus griseus ) and the Asian black bear ( Ursus thibetanus ). Giant pandas avoid areas with
8400-461: The vertebrates ), as well as groups like the sharks and cetaceans , are commonly used. His student Theophrastus (Greece, 370–285 BC) carried on this tradition, mentioning some 500 plants and their uses in his Historia Plantarum . Several plant genera can be traced back to Theophrastus, such as Cornus , Crocus , and Narcissus . Taxonomy in the Middle Ages was largely based on
8540-488: The 1960s. In 1958, Julian Huxley used the term clade . Later, in 1960, Cain and Harrison introduced the term cladistic . The salient feature is arranging taxa in a hierarchical evolutionary tree , with the desideratum that all named taxa are monophyletic. A taxon is called monophyletic if it includes all the descendants of an ancestral form. Groups that have descendant groups removed from them are termed paraphyletic , while groups representing more than one branch from
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#17328455484118680-504: The 1970s, gifts of giant pandas to American and Japanese zoos formed an important part of the diplomacy of the People's Republic of China (PRC), as it marked some of the first cultural exchanges between China and the West. This practice has been termed "panda diplomacy". By 1984, however, pandas were no longer given as gifts. Instead, China began to offer pandas to other nations only on 10-year loans for
8820-466: The Asian bile bear market. The IUCN lists six bear species as vulnerable or endangered , and even least concern species, such as the brown bear , are at risk of extirpation in certain countries. The poaching and international trade of these most threatened populations are prohibited, but still ongoing. The English word "bear" comes from Old English bera and belongs to a family of names for
8960-523: The Chinese language has given the bear many different names, including mò ( 貘 , ancient Chinese name for giant panda), huāxióng ( 花熊 ; "spotted bear") and zhúxióng ( 竹熊 ; "bamboo bear"). The most popular names in China today are dàxióngmāo ( 大熊貓 ; lit. ' giant bear cat ' ), or simply xióngmāo ( 熊貓 ; lit. ' bear cat ' ). As with the word panda in English, xióngmāo ( 熊貓 )
9100-530: The Linnaean system has transformed into a system of modern biological classification intended to reflect the evolutionary relationships among organisms, both living and extinct. The exact definition of taxonomy varies from source to source, but the core of the discipline remains: the conception, naming, and classification of groups of organisms. As points of reference, recent definitions of taxonomy are presented below: The varied definitions either place taxonomy as
9240-487: The Origin of Species (1859) led to a new explanation for classifications, based on evolutionary relationships. This was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler (1883) and Engler (1886–1892). The advent of cladistic methodology in the 1970s led to classifications based on the sole criterion of monophyly , supported by the presence of synapomorphies . Since then,
9380-611: The Qinling population often selects evergreen broadleaf and conifer forests, while pandas in the Qionglai mountainous region exclusively select upland conifer forests. The remaining two populations, namely those occurring in the Liangshan and Xiaoxiangling mountains, predominantly occur in broadleaf evergreen and conifer forests. Giant pandas once roamed across Southeast Asia from Myanmar to northern Vietnam . Their range in China spanned much of
9520-427: The adapted wrist bone of the red panda , which is native to Nepal. In many older sources, the name "panda" or "common panda" refers to the red panda ( Ailurus fulgens ), which was described some 40 years earlier and over that period was the only animal known as a panda. The binomial name Ailuropoda melanoleuca means black and white (melanoleuca) cat-foot (ailuropoda). Since the earliest collection of Chinese writings,
9660-501: The ancient Greek ἄρκτος ( arktos ), meaning bear, as do the names " arctic " and " antarctic ", via the name of the constellation Ursa Major , the "Great Bear", prominent in the northern sky. Bear taxon names such as Ursidae and Ursus come from Latin Ursus/Ursa , he-bear/she-bear. The female first name " Ursula ", originally derived from a Christian saint 's name, means "little she-bear" (diminutive of Latin ursa ). In Switzerland,
9800-522: The animal and plant kingdoms toward the end of the 18th century, well before Charles Darwin's On the Origin of Species was published. The pattern of the "Natural System" did not entail a generating process, such as evolution, but may have implied it, inspiring early transmutationist thinkers. Among early works exploring the idea of a transmutation of species were Zoonomia in 1796 by Erasmus Darwin (Charles Darwin's grandfather), and Jean-Baptiste Lamarck 's Philosophie zoologique of 1809. The idea
9940-548: The basis of distinct cranial measurements, colour patterns, and population genetics . A detailed study of the giant panda's genetic history from 2012 confirms that the separation of the Qinling population occurred about 300,000 years ago, and reveals that the non-Qinling population further diverged into two groups, named the Minshan and the Qionglai - Daxiangling - Xiaoxiangling - Liangshan group respectively, about 2,800 years ago. Of
10080-756: The bear family with other carnivorans is shown in the following phylogenetic tree , which is based on the molecular phylogenetic analysis of six genes in Flynn (2005) with the musteloids updated following the multigene analysis of Law et al. (2018). Feliformia [REDACTED] Canidae [REDACTED] Ursidae [REDACTED] Pinnipedia [REDACTED] Mephitidae [REDACTED] Ailuridae [REDACTED] Procyonidae [REDACTED] Mustelidae [REDACTED] Note that although they are called "bears" in some languages, red pandas and raccoons and their close relatives are not bears, but rather musteloids . There are two phylogenetic hypotheses on
10220-463: The bear in Germanic languages , such as Swedish björn , also used as a first name. This form is conventionally said to be related to a Proto-Indo-European word for "brown", so that "bear" would mean "the brown one". However, Ringe notes that while this etymology is semantically plausible, a word meaning "brown" of this form cannot be found in Proto-Indo-European. He suggests instead that "bear"
10360-521: The brown bears (which includes the polar bear ). Modern brown bears evolved from U. minimus via Ursus etruscus , which itself is ancestral to the extinct Pleistocene cave bear . Species of Ursinae have migrated repeatedly into North America from Eurasia as early as 4 Mya during the early Pliocene. The polar bear is the most recently evolved species and descended from a population of brown bears that became isolated in northern latitudes by glaciation 400,000 years ago. The relationship of
10500-457: The correct name. Some linguists argue, in this construction, "bear" instead of "cat" is the base noun, making the name more grammatically and logically correct, which have led to the popular choice despite official writings. This name did not gain its popularity until 1988, when a private zoo in Tainan painted a sun bear black and white and created the Tainan fake panda incident . For many decades,
10640-459: The country, mainly in Sichuan , and also in neighbouring Shaanxi and Gansu . Successful habitat preservation has seen a rise in panda numbers, though loss of habitat due to human activities remains its biggest threat. In areas with a high concentration of medium-to-large-sized mammals—such as domestic cattle , a species known to degrade the landscape—the giant panda population is generally low. This
10780-564: The country. It has often served as China's national symbol , appeared on Chinese Gold Panda coins since 1982 and as one of the five Fuwa mascots of the 2008 Summer Olympics held in Beijing . The word panda was borrowed into English from French, but no conclusive explanation of the origin of the French word panda has been found. The closest candidate is the Nepali word ponya, possibly referring to
10920-542: The definition of taxa, but the naming and publication of new taxa is governed by sets of rules. In zoology , the nomenclature for the more commonly used ranks ( superfamily to subspecies ), is regulated by the International Code of Zoological Nomenclature ( ICZN Code ). In the fields of phycology , mycology , and botany , the naming of taxa is governed by the International Code of Nomenclature for algae, fungi, and plants ( ICN ). The initial description of
11060-428: The dietary switch was initiated prior to T1R1 loss-of-function. Pandas eat any of 25 bamboo species in the wild, with the most common including Fargesia dracocephala and Fargesia rufa . Only a few bamboo species are widespread at the high altitudes pandas now inhabit. Bamboo leaves contain the highest protein levels; stems have less. Because of the synchronous flowering, death, and regeneration of all bamboo within
11200-466: The early Pleistocene. By 3–4 Mya, the species Ursus minimus appears in the fossil record of Europe; apart from its size, it was nearly identical to today's Asian black bear . It is likely ancestral to all bears within Ursinae, perhaps aside from the sloth bear. Two lineages evolved from U. minimus : the black bears (including the sun bear , the Asian black bear, and the American black bear ); and
11340-558: The economics of keeping pandas, which costs five times more than keeping the next most expensive animal, an elephant. American zoos generally pay the Chinese government $ 1 million a year in fees, as part of a typical ten-year contract. San Diego's contract with China was to expire in 2008, but got a five-year extension at about half of the previous yearly cost. The last contract, with the Memphis Zoo in Memphis, Tennessee , ended in 2013. In
11480-417: The eight extant species in the bear family Ursidae, the giant panda's lineage branched off the earliest. Giant panda ( Ailuropoda melanoleuca ) [REDACTED] Spectacled bear ( Tremarctos ornatus ) [REDACTED] Ursinae (black, brown, sloth, sun, and polar bears) [REDACTED] The giant panda is endemic to China . It is found in small, fragmented populations in six mountainous regions in
11620-399: The entire world. Other (partial) revisions may be restricted in the sense that they may only use some of the available character sets or have a limited spatial scope. A revision results in a conformation of or new insights in the relationships between the subtaxa within the taxon under study, which may lead to a change in the classification of these subtaxa, the identification of new subtaxa, or
11760-489: The evidentiary basis has been expanded with data from molecular genetics that for the most part complements traditional morphology . Naming and classifying human surroundings likely began with the onset of language. Distinguishing poisonous plants from edible plants is integral to the survival of human communities. Medicinal plant illustrations show up in Egyptian wall paintings from c. 1500 BC , indicating that
11900-516: The exception of spiders published in Svenska Spindlar ). Even taxonomic names published by Linnaeus himself before these dates are considered pre-Linnaean. Modern taxonomy is heavily influenced by technology such as DNA sequencing , bioinformatics , databases , and imaging . A pattern of groups nested within groups was specified by Linnaeus' classifications of plants and animals, and these patterns began to be represented as dendrograms of
12040-1085: The extinct bear dogs of the family Amphicyonidae . Below is the cladogram based on McKenna and Bell (1997) classification: † Amphicyonidae [REDACTED] † Amphicynodontidae [REDACTED] Pinnipedia [REDACTED] † Hemicyoninae † Agriotheriinae † Ursavinae Ailuropodinae [REDACTED] Tremarctinae [REDACTED] Ursinae [REDACTED] Giant panda ( Ailuropoda melanoleuca ) [REDACTED] Spectacled bear ( Tremarctos ornatus ) [REDACTED] Sloth bear ( Melursus ursinus ) [REDACTED] Sun bear ( Helarctos malayanus ) [REDACTED] Asian black bear ( Ursus thibetanus ) [REDACTED] American black bear ( Ursus americanus ) [REDACTED] Polar bear ( Ursus maritimus ) [REDACTED] Brown bear ( Ursus arctos ) [REDACTED] Giant panda ( Ailuropoda melanoleuca ) [REDACTED] Taxonomy (biology) In biology , taxonomy (from Ancient Greek τάξις ( taxis ) 'arrangement' and -νομία ( -nomia ) ' method ')
12180-403: The eye patches might facilitate them identifying one another. The giant panda's thick, woolly coat keeps it warm in the cool forests of its habitat. The panda's skull shape is typical of durophagous carnivorans. It has evolved from previous ancestors to exhibit larger molars with increased complexity and expanded temporal fossa. A study revealed that a 117.5 kg (259 lb) giant panda had
12320-500: The eyes. The rest of the animal's coat is white. The bear's distinctive coloration appears to serve as camouflage in both winter and summer environments as they do not hibernate . The white areas serve as camouflage in snow, while the black shoulders and legs conceal them in shade. Studies in the wild have found that when viewed from a distance, the panda displays disruptive coloration , while up close, they rely more on blending in . The black ears may be used to display aggression, while
12460-486: The first modern groups tied to fossil ancestors was birds. Using the then newly discovered fossils of Archaeopteryx and Hesperornis , Thomas Henry Huxley pronounced that they had evolved from dinosaurs, a group formally named by Richard Owen in 1842. The resulting description, that of dinosaurs "giving rise to" or being "the ancestors of" birds, is the essential hallmark of evolutionary taxonomic thinking. As more and more fossil groups were found and recognized in
12600-511: The following March. Activity is also directly related to the amount of sunlight during colder days. There is a significant interaction of solar radiation, such that solar radiation has a stronger positive effect on activity levels of panda bears. Pandas communicate through vocalisation and scent marking such as clawing trees or spraying urine . They are able to climb and take shelter in hollow trees or rock crevices, but do not establish permanent dens. For this reason, pandas do not hibernate , which
12740-682: The formal naming of clades. Linnaean ranks are optional and have no formal standing under the PhyloCode , which is intended to coexist with the current, rank-based codes. While popularity of phylogenetic nomenclature has grown steadily in the last few decades, it remains to be seen whether a majority of systematists will eventually adopt the PhyloCode or continue using the current systems of nomenclature that have been employed (and modified, but arguably not as much as some systematists wish) for over 250 years. Well before Linnaeus, plants and animals were considered separate Kingdoms. Linnaeus used this as
12880-566: The fur and its mother's saliva . A month after birth, the colour pattern of the cub's fur is fully developed. Its fur is very soft and coarsens with age. The cub begins to crawl at 75 to 80 days; mothers play with their cubs by rolling and wrestling with them. The cubs can eat small quantities of bamboo after six months, though mother's milk remains the primary food source for most of the first year. Giant panda cubs weigh 45 kg (99 lb) at one year and live with their mothers until they are 18 months to two years old. The interval between births in
13020-560: The fusing of some chromosomes, and the banding patterns on these match those of the ursine species, but differ from those of procyonids, which supports the inclusion of these two species in Ursidae rather than in Procyonidae , where they had been placed by some earlier authorities. The earliest members of Ursidae belong to the extinct subfamily Amphicynodontinae, including Parictis (late Eocene to early middle Miocene , 38–18 Mya ) and
13160-454: The genus Ursavus during the early Oligocene (30–28 Mya); this genus proliferated into many species in Asia and is ancestral to all living bears. Species of Ursavus subsequently entered North America, together with Amphicynodon and Cephalogale , during the early Miocene (21–18 Mya). Members of the living lineages of bears diverged from Ursavus between 15 and 20 Mya, likely via
13300-494: The giant panda has been driven out of the lowland areas where it once lived. The wild population has increased again to 1,864 individuals as of March 2015. Since 2016, it has been listed as Vulnerable on the IUCN Red List . In July 2021, Chinese authorities also classified the giant panda as vulnerable. It is a conservation-reliant species . By 2007, the captive population comprised 239 giant pandas in China and another 27 outside
13440-447: The giant panda suggests that the dietary switch could have initiated from the loss of the sole umami taste receptor, encoded by the genes TAS1R1 and TAS1R3 (also known as T1R1 and T1R3), resulting from two frameshift mutations within the T1R1 exons. Umami taste corresponds to high levels of glutamate as found in meat and may have thus altered the food choice of the giant panda. Although
13580-466: The late 19th and early 20th centuries, palaeontologists worked to understand the history of animals through the ages by linking together known groups. With the modern evolutionary synthesis of the early 1940s, an essentially modern understanding of the evolution of the major groups was in place. As evolutionary taxonomy is based on Linnaean taxonomic ranks, the two terms are largely interchangeable in modern use. The cladistic method has emerged since
13720-562: The legendary fierce creature pixiu (貔貅) as referring to the giant panda is also common. During the reign of the Yongle Emperor (early 15th century), his relative from Kaifeng sent him a captured zouyu ( 騶虞 ), and another zouyu was sighted in Shandong . Zouyu is a legendary "righteous" animal, which, similarly to a qilin , only appears during the rule of a benevolent and sincere monarch. Pandas have been kept in zoos as early as
13860-646: The lone surviving representative of the Tremarctinae, the South American spectacled bear ( T. ornatus ). The subfamily Ursinae experienced a dramatic proliferation of taxa about 5.3–4.5 Mya, coincident with major environmental changes; the first members of the genus Ursus appeared around this time. The sloth bear is a modern survivor of one of the earliest lineages to diverge during this radiation event (5.3 Mya); it took on its peculiar morphology, related to its diet of termites and ants, no later than by
14000-418: The male first name "Urs" is especially popular, while the name of the canton and city of Bern is by legend derived from Bär , German for bear. The Germanic name Bernard (including Bernhardt and similar forms) means "bear-brave", "bear-hardy", or "bold bear". The Old English name Beowulf is a kenning , "bee-wolf", for bear, in turn meaning a brave warrior. The family Ursidae is one of nine families in
14140-423: The male may mount her repeatedly to ensure successful fertilisation. The gestation period is somewhere between 95 and 160 days - the variability is due to the fact that the fertilized egg may linger in the reproductive system for a while before implanting on the uterine wall. Giant pandas give birth to twins in about half of pregnancies. If twins are born, usually only one survives in the wild. The mother will select
14280-401: The merger of previous subtaxa. Taxonomic characters are the taxonomic attributes that can be used to provide the evidence from which relationships (the phylogeny ) between taxa are inferred. Kinds of taxonomic characters include: The term " alpha taxonomy " is primarily used to refer to the discipline of finding, describing, and naming taxa , particularly species. In earlier literature,
14420-461: The nutrients that they need and to balance their diet for reproduction. Although adult giant pandas have few natural predators other than humans, young cubs are vulnerable to attacks by snow leopards , yellow-throated martens , eagles, feral dogs, and the Asian black bear . Sub-adults weighing up to 50 kg (110 lb) may be vulnerable to predation by leopards . Giant pandas are sympatric with other large mammals and bamboo feeders, such as
14560-454: The panda's most distinctive features, its large size and round face, are adaptations to its bamboo diet. Anthropologist Russell Ciochon observed: "[much] like the vegetarian gorilla, the low body surface area to body volume [of the giant panda] is indicative of a lower metabolic rate. This lower metabolic rate and a more sedentary lifestyle allows the giant panda to subsist on nutrient poor resources such as bamboo." The giant panda's round face
14700-402: The population is threatened by canine distemper virus (CDV) , canine parvovirus , rotavirus , canine adenovirus , and canine coronavirus . Bacteria, such as Clostridium welchii , Proteus mirabilis , Klebsiella pneumoniae , and Escherichia coli , may also be lethal. The giant panda is a terrestrial animal and primarily spends its life roaming and feeding in the bamboo forests of
14840-434: The possibilities of closer co-operation with their cytological, ecological and genetics colleagues and to acknowledge that some revision or expansion, perhaps of a drastic nature, of their aims and methods, may be desirable ... Turrill (1935) has suggested that while accepting the older invaluable taxonomy, based on structure, and conveniently designated "alpha", it is possible to glimpse a far-distant taxonomy built upon as wide
14980-558: The precise taxonomic classification of the giant panda was under debate because it shares characteristics with both bears and raccoons . In 1985, molecular studies indicated that the giant panda is a true bear, part of the family Ursidae. These studies show it diverged about 19 million years ago from the common ancestor of the Ursidae ; it is the most basal member of this family and equidistant from all other extant bear species. Two subspecies of giant panda have been recognized on
15120-507: The pseudogenisation (conversion into a pseudogene ) of the umami taste receptor in Ailuropoda coincides with the dietary switch to herbivory, it is likely a result of, and not the reason for, the dietary change. The mutation time for the T1R1 gene in the giant panda is estimated to 4.2 mya while fossil evidence indicates bamboo consumption in the giant panda species at least 7 mya, signifying that although complete herbivory occurred around 2 mya,
15260-774: The rank of Order, although both exclude fossil representatives. A separate compilation (Ruggiero, 2014) covers extant taxa to the rank of Family. Other, database-driven treatments include the Encyclopedia of Life , the Global Biodiversity Information Facility , the NCBI taxonomy database , the Interim Register of Marine and Nonmarine Genera , the Open Tree of Life , and the Catalogue of Life . The Paleobiology Database
15400-628: The relationships among extant and fossil bear species. One is all species of bears are classified in seven subfamilies as adopted here and related articles: Amphicynodontinae , Hemicyoninae , Ursavinae , Agriotheriinae , Ailuropodinae , Tremarctinae , and Ursinae . Below is a cladogram of the subfamilies of bears after McLellan and Reiner (1992) and Qiu et al. . (2014): † Amphicynodontinae [REDACTED] † Hemicyoninae † Ursavinae † Agriotheriinae Ailuropodinae [REDACTED] Tremarctinae [REDACTED] Ursinae [REDACTED] The second alternative phylogenetic hypothesis
15540-407: The same, sometimes slightly different, but always related and intersecting. The broadest meaning of "taxonomy" is used here. The term itself was introduced in 1813 by de Candolle , in his Théorie élémentaire de la botanique . John Lindley provided an early definition of systematics in 1830, although he wrote of "systematic botany" rather than using the term "systematics". Europeans tend to use
15680-509: The short, straight digestive tract. It is also noted, however, that such rapid passage of digesta limits the potential of microbial digestion in the gastrointestinal tract, limiting alternative forms of digestion. Given this voluminous diet, the giant panda defecates up to 40 times a day. The limited energy input imposed on it by its diet has affected the panda's behavior. The giant panda tends to limit its social interactions and avoids steeply sloping terrain to limit its energy expenditures. Two of
15820-435: The signaler's competitive ability outweighs their own. A pandas' size can be conveyed through the height of the scent mark. Since larger animals can place higher scent marks, an elevated scent mark advertises a higher competitive ability. Age must also be taken into consideration when assessing a competitor's fighting ability. For example, a mature panda will be larger than a younger, immature panda and possess an advantage during
15960-530: The slightly younger Allocyon (early Oligocene , 34–30 Mya), both from North America. These animals looked very different from today's bears, being small and raccoon -like in overall appearance, with diets perhaps more similar to that of a badger . Parictis does not appear in Eurasia and Africa until the Miocene. It is unclear whether late-Eocene ursids were also present in Eurasia, although faunal exchange across
16100-422: The smallest baby of any placental mammal. It nurses from its mother's breast six to 14 times a day for up to 30 minutes at a time. For three to four hours, the mother may leave the den to feed, which leaves the cub defenseless. One to two weeks after birth, the cub's skin turns grey where its hair will eventually become black. Slight pink colour may appear on the cub's fur, as a result of a chemical reaction between
16240-455: The southeast region. By the Pleistocene , climate change affected panda populations, and the subsequent domination of modern humans led to large-scale habitat loss. In 2001, it was estimated that the range of the giant panda had declined by about 99% of its range in earlier millenniums. The giant panda has a body shape typical of bears. It has black fur on its ears, limbs, shoulders and around
16380-540: The species Ursavus elmensis . Based on genetic and morphological data, the Ailuropodinae (pandas) were the first to diverge from other living bears about 19 Mya, although no fossils of this group have been found before about 11 Mya. The New World short-faced bears (Tremarctinae) differentiated from Ursinae following a dispersal event into North America during the mid-Miocene (about 13 Mya). They invaded South America (≈2.5 or 1.2 Ma) following formation of
16520-560: The species. As of 2009, it is expected that zoos in destinations such as San Diego in the United States and Mexico City will be able to provide their own semen to inseminate more giant pandas. Attempts have also been made to reproduce giant pandas by interspecific pregnancy where cloned panda embryos were implanted into the uterus of an animal of another species. This has resulted in panda fetuses, but no live births. In Ancient China, people thought pandas to be rare and noble creatures –
16660-417: The stronger of the cubs, and the weaker cub will die due to starvation. The mother is thought to be unable to produce enough milk for two cubs since she does not store fat. The father has no part in helping raise the cub. When the cub is first born, it is pink, blind, and toothless, weighing only 90 to 130 g (3.2 to 4.6 oz), or about 1 / 800 of the mother's weight, proportionally
16800-434: The suborder Caniformia , or "doglike" carnivorans, within the order Carnivora . Bears' closest living relatives are the pinnipeds , canids , and musteloids (some scholars formerly argued that bears are directly derived from canids and should not be classified as a separate family). Modern bears comprise eight species in three subfamilies: Ailuropodinae (monotypic with the giant panda ), Tremarctinae (monotypic with
16940-509: The support of the pinniped–amphicynodontine clade, other morphological and some molecular evidence supports bears being the closest living relatives to pinnipeds. The raccoon-sized, dog-like Cephalogale is the oldest-known member of the subfamily Hemicyoninae , which first appeared during the middle Oligocene in Eurasia about 30 Mya. The subfamily includes the younger genera Phoberocyon (20–15 Mya), and Plithocyon (15–7 Mya). A Cephalogale -like species gave rise to
17080-472: The term had a different meaning, referring to morphological taxonomy, and the products of research through the end of the 19th century. William Bertram Turrill introduced the term "alpha taxonomy" in a series of papers published in 1935 and 1937 in which he discussed the philosophy and possible future directions of the discipline of taxonomy. ... there is an increasing desire amongst taxonomists to consider their problems from wider viewpoints, to investigate
17220-482: The terms "systematics" and "biosystematics" for the study of biodiversity as a whole, whereas North Americans tend to use "taxonomy" more frequently. However, taxonomy, and in particular alpha taxonomy , is more specifically the identification, description, and naming (i.e., nomenclature) of organisms, while "classification" focuses on placing organisms within hierarchical groups that show their relationships to other organisms. A taxonomic revision or taxonomic review
17360-505: The three-domain method is the separation of Archaea and Bacteria , previously grouped into the single kingdom Bacteria (a kingdom also sometimes called Monera ), with the Eukaryota for all organisms whose cells contain a nucleus . A small number of scientists include a sixth kingdom, Archaea, but do not accept the domain method. Thomas Cavalier-Smith , who published extensively on the classification of protists , in 2002 proposed that
17500-427: The top rank, dividing the physical world into the vegetable, animal and mineral kingdoms. As advances in microscopy made the classification of microorganisms possible, the number of kingdoms increased, five- and six-kingdom systems being the most common. Domains are a relatively new grouping. First proposed in 1977, Carl Woese 's three-domain system was not generally accepted until later. One main characteristic of
17640-436: The traditional three domains. Partial classifications exist for many individual groups of organisms and are revised and replaced as new information becomes available; however, comprehensive, published treatments of most or all life are rarer; recent examples are that of Adl et al., 2012 and 2019, which covers eukaryotes only with an emphasis on protists, and Ruggiero et al., 2015, covering both eukaryotes and prokaryotes to
17780-439: The transfer would be considered "domestic" or "international" or whether any true conservation purpose would be served by the exchange. A contest in 2006 to name the pandas was held in the mainland, resulting in the politically charged names Tuan Tuan and Yuan Yuan (from simplified Chinese : 团圆 ; traditional Chinese : 團圓 ; pinyin : tuanyuan ; lit. 'reunion', implying reunification ). China's offer
17920-514: The tree of life are called polyphyletic . Monophyletic groups are recognized and diagnosed on the basis of synapomorphies , shared derived character states. Cladistic classifications are compatible with traditional Linnean taxonomy and the Codes of Zoological and Botanical nomenclature , to a certain extent. An alternative system of nomenclature, the International Code of Phylogenetic Nomenclature or PhyloCode has been proposed, which regulates
18060-478: The use of panda pelts to control menstruation as described in the Qin dynasty encyclopedia Erya . The creature named mo (貘) mentioned in some ancient books has been interpreted as giant panda. The dictionary Shuowen Jiezi ( Eastern Han Dynasty ) says that the mo , from Shu (Sichuan), is bear-like, but yellow-and-black, although the older Erya describes mo simply as a "white leopard". The interpretation of
18200-638: The uses of different species were understood and that a basic taxonomy was in place. Organisms were first classified by Aristotle ( Greece , 384–322 BC) during his stay on the Island of Lesbos . He classified beings by their parts, or in modern terms attributes , such as having live birth, having four legs, laying eggs, having blood, or being warm-bodied. He divided all living things into two groups: plants and animals . Some of his groups of animals, such as Anhaima (animals without blood, translated as invertebrates ) and Enhaima (animals with blood, roughly
18340-540: The wild is generally two years. Initially, the primary method of breeding giant pandas in captivity was by artificial insemination , as they seemed to lose their interest in mating once they were captured. This led some scientists to trying methods such as showing them videos of giant pandas mating and giving the males sildenafil (commonly known as Viagra). In the 2000s, researchers started having success with captive breeding programs, and they have now determined giant pandas have comparable breeding to some populations of
18480-527: The winter for a long period of hibernation , up to 100 days. Bears have been hunted since prehistoric times for their meat and fur; they have also been used for bear-baiting and other forms of entertainment, such as being made to dance . With their powerful physical presence, they play a prominent role in the arts , mythology , and other cultural aspects of various human societies. In modern times, bears have come under pressure through encroachment on their habitats and illegal trade in bear parts, including
18620-486: Was Methodus Plantarum Nova (1682), in which he published details of over 18,000 plant species. At the time, his classifications were perhaps the most complex yet produced by any taxonomist, as he based his taxa on many combined characters. The next major taxonomic works were produced by Joseph Pitton de Tournefort (France, 1656–1708). His work from 1700, Institutiones Rei Herbariae , included more than 9000 species in 698 genera, which directly influenced Linnaeus, as it
18760-551: Was entitled " Systema Naturae " ("the System of Nature"), implying that he, at least, believed that it was more than an "artificial system"). Later came systems based on a more complete consideration of the characteristics of taxa, referred to as "natural systems", such as those of de Jussieu (1789), de Candolle (1813) and Bentham and Hooker (1862–1863). These classifications described empirical patterns and were pre- evolutionary in thinking. The publication of Charles Darwin 's On
18900-491: Was implemented by McKenna et al. (1997) to classify all the bear species into the superfamily Ursoidea , with Hemicyoninae and Agriotheriinae being classified in the family "Hemicyonidae". Amphicynodontinae under this classification were classified as stem- pinnipeds in the superfamily Phocoidea . In the McKenna and Bell classification both bears and pinnipeds are in a parvorder of carnivoran mammals known as Ursida , along with
19040-775: Was initially rejected by Chen Shui-bian , then President of Taiwan. However, when Ma Ying-jeou assumed the presidency in 2008, the offer was accepted and the pandas arrived in December of that year. In the 2020s, certain "celebrity pandas" have gained a cult following amongst internet users, with dedicated fan accounts existing to keep tabs on the animals. Known as "giant panda fever" or "panda-monium", individual pandas are known to get billions of views and engagements on social media, as well as product lines specifically emulating them. At Chengdu Research Base of Giant Panda Breeding , certain of these "celebrity pandas" are known to garner hours-long lines specifically to see them. The giant panda
19180-459: Was originally used to describe just the red panda, but dàxióngmāo ( 大熊貓 ) and xiǎoxióngmāo ( 小熊猫 ; lit. ' little bear cat ' ) were coined to differentiate between the species. In Taiwan , another popular name for panda is the inverted dàmāoxióng ( 大貓熊 ; lit. ' giant cat bear ' ), though many encyclopedias and dictionaries in Taiwan still use the "bear cat" form as
19320-597: Was popularized in the Anglophone world by the speculative but widely read Vestiges of the Natural History of Creation , published anonymously by Robert Chambers in 1844. With Darwin's theory, a general acceptance quickly appeared that a classification should reflect the Darwinian principle of common descent . Tree of life representations became popular in scientific works, with known fossil groups incorporated. One of
19460-532: Was the Italian physician Andrea Cesalpino (1519–1603), who has been called "the first taxonomist". His magnum opus De Plantis came out in 1583, and described more than 1500 plant species. Two large plant families that he first recognized are in use: the Asteraceae and Brassicaceae . In the 17th century John Ray ( England , 1627–1705) wrote many important taxonomic works. Arguably his greatest accomplishment
19600-429: Was the text he used as a young student. The Swedish botanist Carl Linnaeus (1707–1778) ushered in a new era of taxonomy. With his major works Systema Naturae 1st Edition in 1735, Species Plantarum in 1753, and Systema Naturae 10th Edition , he revolutionized modern taxonomy. His works implemented a standardized binomial naming system for animal and plant species, which proved to be an elegant solution to
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