9-871: Orchideae is a tribe of orchids in the subfamily Orchidoideae . Historically, it was divided into 2 subtribes, Orchidinae and Habenariinae. The subtribe Orchidinae alone contains about 1,800 species. However, although some phylogenetic studies have established the monophyly of the subtribes, the generic boundaries are unclear, with many genera as traditionally circumscribed being paraphyletic or even polyphyletic . Species of genera such as Habenaria and Platanthera have been placed into both subtribes. A 2017 molecular phylogenetic study found that both subtribes did form clades , but did not formally recognize Habenariinae, because of missing genera and uncertainty over generic boundaries. The Asian species of Orchideae, in particular, have been subject to repeated changes of generic placement from 2012 onwards. As of 2017, Orchideae
18-459: A 2017 molecular phylogenetic study are listed below. As noted above, some generic boundaries remain uncertain; Habenaria in particular is not monophyletic . This Orchidoideae -related article is a stub . You can help Misplaced Pages by expanding it . Orchidoideae See text The Orchidoideae , or the orchidoid orchids, are a subfamily of the orchid family ( Orchidaceae ) that contains around 3630 species. Species typically have
27-598: A single ( monandrous ), fertile anther which is erect and basitonic . The subfamily Orchidoideae and the previously recognized subfamily Spiranthoideae are considered the closest allies in the natural group of the monandrous orchids because of several generally shared characters: Phylogeny of the Orchidoideae is volatile and still subject to change. Historically, the Orchidoideae have been partitioned into up to 6 tribes , including Orchideae , Diseae , Cranichideae , Chloraeeae , Diurideae , and Codonorchideae . However,
36-550: Is divided into 6 subtribes: Brownleeinae , Pachitinae , Huttonaeinae , Orchidinae, Coryciinae , and Disinae . Once divided into the Orchidinae and Habenariinae (a nonsystematic paraphyletic splitting), the Orchideae now comprise 6 subtribes. Along with previous molecular studies, Chase et al's 2015 analysis confirmed that Diseae was a polyphyletic grouping. Consequently, the former subtribes of Diseae have been redistributed within
45-475: The Orchideae (for instance, Satyrium is now classified in the Orchidinae). In his 2015 study Chase recommended further study of historically problematic genera in the former Diseae such as Huttonaea and Pachites . Further molecular analysis by Chen et al. in 2017 led to the following reorganization: Brownleeinae Pachitinae Huttonaeinae Orchidinae Coryciinae Disinae Genera included in
54-480: The Orchidinae and Habenariinae (a nonsystematic paraphyletic splitting), the Orchideae now comprise 6 subtribes. Along with previous molecular studies, Chase et al's 2015 analysis confirmed that Diseae was a polyphyletic grouping. Consequently, the former subtribes of Diseae have been redistributed within the Orchideae (for instance, Satyrium is now classified in the Orchidinae). In his 2015 study Chase recommended further study of historically problematic genera in
63-692: The former Chloraeeae are sister to the rest of the Cranichideae. As such, the Chloraeeae are now considered a subtribe of the Cranichideae. Moreover, based on morphological and genetic evidence, the genus Discyphus was elevated out of the Spiranthinae to form its own monophyletic subtribe, the Discyphinae . Chloraeinae Pterostylidinae Goodyerinae Galeottiellinae Manniellinae Spiranthinae Discyphinae Cranichidinae Once divided into
72-433: The former Diseae such as Huttonaea and Pachites . Further molecular analysis by Chen et al. in 2017 led to the following reorganization: Brownleeinae Pachitinae Huttonaeinae Orchidinae Coryciinae Disinae The Diurideae, a predominantly Australasian tribe, retain their previous subtribal organization and comprise 9 taxa: Discyphus Discyphus is a genus of flowering plants from
81-498: The most recent molecular phylogenetics analysis led by Chase et al. in 2015 and subsequently by Chen et al. in 2017 indicates that the Orchidoideae should be split into 4 tribes: the Orchideae, Cranichideae, Diurideae, and Codonorchideae. This monophyletic tribe consists of only one genus, Codonorchis , found in South America . This cosmopolitan tribe comprises 8 currently accepted subtribes. Molecular evidence suggests that
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