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In ecology , a niche is the match of a species to a specific environmental condition. It describes how an organism or population responds to the distribution of resources and competitors (for example, by growing when resources are abundant, and when predators , parasites and pathogens are scarce) and how it in turn alters those same factors (for example, limiting access to resources by other organisms, acting as a food source for predators and a consumer of prey). "The type and number of variables comprising the dimensions of an environmental niche vary from one species to another [and] the relative importance of particular environmental variables for a species may vary according to the geographic and biotic contexts".

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63-490: [REDACTED] Look up niche in Wiktionary, the free dictionary. Niche may refer to: Science [ edit ] Developmental niche , a concept for understanding the cultural context of child development Ecological niche , a term describing the relational position of an organism's species Niche differentiation , in ecology, the process by which competing species use

126-409: A Gaussian might describe the frequency with which a species ate prey of a certain size, giving a more detailed niche description than simply specifying some median or average prey size. For such a bell-shaped distribution, the position , width and form of the niche correspond to the mean , standard deviation and the actual distribution itself. One advantage in using statistics is illustrated in

189-447: A storage effect . Species can differentiate their niche via a competition-predation trade-off if one species is a better competitor when predators are absent, and the other is better when predators are present. Defenses against predators, such as toxic compounds or hard shells, are often metabolically costly. As a result, species that produce such defenses are often poor competitors when predators are absent. Species can coexist through

252-532: A British Thoroughbred racehorse Niche (video game) , a 2017 video game Niche, a character in the manga Tegami Bachi Niche (footballer) , Bissau-Guinean footballer Network in Canadian History and Environment , a group of Canadian environmental historians Nishe , a B-Side from the 2000 single Unintended by English rock band, Muse Topics referred to by the same term [REDACTED] This disambiguation page lists articles associated with

315-727: A certain environment (have overlapping requirement niches) but fundamentally differ the ways that they use (or "impact") that environment. These requirements have repeatedly been violated by nonnative (i.e. introduced and invasive ) species, which often coexist with new species in their nonnative ranges, but do not appear to be constricted these requirements. In other words, contemporary niche theory predicts that species will be unable to invade new environments outside of their requirement (i.e. realized) niche, yet many examples of this are well-documented. Additionally, contemporary niche theory predicts that species will be unable to establish in environments where other species already consume resources in

378-427: A climatic perspective, to explain distribution and abundance. Current predictions on species responses to climate change strongly rely on projecting altered environmental conditions on species distributions. However, it is increasingly acknowledged that climate change also influences species interactions and an Eltonian perspective may be advantageous in explaining these processes. This perspective of niche allows for

441-482: A competition-predation trade-off if predators are more abundant when the less defended species is common, and less abundant if the well-defended species is common. This effect has been criticized as being weak, because theoretical models suggest that only two species within a community can coexist because of this mechanism. Two ecological paradigms deal with the problem. The first paradigm predominates in what may be called "classical" ecology. It assumes that niche space

504-412: A fundamental niche of the entire slope of a hillside, but its realized niche is only the top portion of the slope because species Y, which is a better competitor but cannot survive on the top portion of the slope, has excluded it from the lower portion of the slope. With this scenario, competition will continue indefinitely in the middle of the slope between these two species. Because of this, detection of

567-474: A given species), 'niche partitioning' (resource differentiation by coexisting species), and 'niche overlap' (overlap of resource use by different species). Statistics were introduced into the Hutchinson niche by Robert MacArthur and Richard Levins using the 'resource-utilization' niche employing histograms to describe the 'frequency of occurrence' as a function of a Hutchinson coordinate. So, for instance,

630-560: A group of Canadian environmental historians Nishe , a B-Side from the 2000 single Unintended by English rock band, Muse Topics referred to by the same term [REDACTED] This disambiguation page lists articles associated with the title Niche . If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Niche&oldid=1258203280 " Category : Disambiguation pages Hidden categories: Short description

693-455: A mammal-like niche. Island biogeography can help explain island species and associated unfilled niches. The ecological meaning of niche comes from the meaning of niche as a recess in a wall for a statue, which itself is probably derived from the Middle French word nicher , meaning to nest . The term was coined by the naturalist Roswell Hill Johnson but Joseph Grinnell was probably

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756-401: A market sector Niche blogging , a blog focused on a niche market Other uses [ edit ] Bassline (music genre) , a type of music related to UK garage also called Niche Niche (architecture) , an exedra or an apse that has been reduced in size A cell of a columbarium for a cremation urn Niche (company) , an Internet search and review service Niche (horse) ,

819-459: A niche specific to each species. Species can however share a 'mode of life' or 'autecological strategy' which are broader definitions of ecospace. For example, Australian grasslands species, though different from those of the Great Plains grasslands, exhibit similar modes of life. Once a niche is left vacant, other organisms can fill that position. For example, the niche that was left vacant by

882-487: A species' density declines, then the food it most depends on will become more abundant (since there are so few individuals to consume it). As a result, the remaining individuals will experience less competition for food. Although "resource" generally refers to food, species can partition other non-consumable objects, such as parts of the habitat. For example, warblers are thought to coexist because they nest in different parts of trees. Species can also partition habitat in

945-417: A term describing the relational position of an organism's species Niche differentiation , in ecology, the process by which competing species use the environment differently in a way that helps them to coexist Niche (protein structural motif) Stem-cell niche , the necessary cellular environment of a stem cell Business [ edit ] Niche market , a focused, targetable portion (subset) of

1008-495: A type of music related to UK garage also called Niche Niche (architecture) , an exedra or an apse that has been reduced in size A cell of a columbarium for a cremation urn Niche (company) , an Internet search and review service Niche (horse) , a British Thoroughbred racehorse Niche (video game) , a 2017 video game Niche, a character in the manga Tegami Bachi Niche (footballer) , Bissau-Guinean footballer Network in Canadian History and Environment ,

1071-451: A way that gives them access to different types of resources. As stated in the introduction, anole lizards appear to coexist because each uses different parts of the forests as perch locations. This likely gives them access to different species of insects. Research has determined that plants can recognize each other's root systems and differentiate between a clone, a plant grown from the same mother plants seeds, and other species. Based on

1134-511: Is absent or low, and therefore detection of niche differentiation is difficult or impossible. Below is a list of ways that species can partition their niche. This list is not exhaustive, but illustrates several classic examples. Resource partitioning is the phenomenon where two or more species divides out resources like food, space, resting sites etc. to coexist. For example, some lizard species appear to coexist because they consume insects of differing sizes. Alternatively, species can coexist on

1197-456: Is central to ecological biogeography , which focuses on spatial patterns of ecological communities. "Species distributions and their dynamics over time result from properties of the species, environmental variation..., and interactions between the two—in particular the abilities of some species, especially our own, to modify their environments and alter the range dynamics of many other species." Alteration of an ecological niche by its inhabitants

1260-548: Is constrained by different natural enemies, they will be able to coexist. Early work focused on specialist predators; however, more recent studies have shown that predators do not need to be pure specialists, they simply need to affect each prey species differently. The Janzen–Connell hypothesis represents a form of predator partitioning. Conditional differentiation (sometimes called temporal niche partitioning ) occurs when species differ in their competitive abilities based on varying environmental conditions. For example, in

1323-415: Is determined by the habitat in which a species lives and its accompanying behavioral adaptations . An Eltonian niche emphasizes that a species not only grows in and responds to an environment, it may also change the environment and its behavior as it grows. The Hutchinsonian niche uses mathematics and statistics to try to explain how species coexist within a given community. The concept of ecological niche

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1386-403: Is different from Wikidata All article disambiguation pages All disambiguation pages niche [REDACTED] Look up niche in Wiktionary, the free dictionary. Niche may refer to: Science [ edit ] Developmental niche , a concept for understanding the cultural context of child development Ecological niche ,

1449-474: Is greater than inter-specific (between species) competition. Since niche differentiation concentrates competition within-species, due to a decrease in between-species competition, the Lotka-Volterra model predicts that niche differentiation of any degree will result in coexistence. In reality, this still leaves the question of how much differentiation is needed for coexistence. A vague answer to this question

1512-489: Is largely saturated with individuals and species, leading to strong competition. Niches are restricted because "neighbouring" species, i.e., species with similar ecological characteristics such as similar habitats or food preferences, prevent expansion into other niches or even narrow niches down. This continual struggle for existence is an important assumption of natural selection introduced by Darwin as an explanation for evolution. The other paradigm assumes that niche space

1575-455: Is merely the product of the competitive exclusion principle. Also, because no species is out-competing any other species in the final community, the presence of niche differentiation will be difficult or impossible to detect. Finally, niche differentiation can arise as an evolutionary effect of competition. In this case, two competing species will evolve different patterns of resource use so as to avoid competition. Here too, current competition

1638-405: Is necessary for ecologists to be able to detect, measure, and quantify the niches of different coexisting and competing species. This is often done through a combination of detailed ecological studies, controlled experiments (to determine the strength of competition), and mathematical models . To understand the mechanisms of niche differentiation and competition, much data must be gathered on how

1701-419: Is that the more similar two species are, the more finely balanced the suitability of their environment must be in order to allow coexistence. There are limits to the amount of niche differentiation required for coexistence, and this can vary with the type of resource, the nature of the environment, and the amount of variation both within and between the species. To answer questions about niche differentiation, it

1764-425: Is the niche that is filled by birds of prey which eat small animals such as shrews and mice. In an oak wood this niche is filled by tawny owls , while in the open grassland it is occupied by kestrels . The existence of this carnivore niche is dependent on the further fact that mice form a definite herbivore niche in many different associations, although the actual species of mice may be quite different. Conceptually,

1827-462: Is the topic of niche construction . The majority of species exist in a standard ecological niche, sharing behaviors, adaptations, and functional traits similar to the other closely related species within the same broad taxonomic class, but there are exceptions. A premier example of a non-standard niche filling species is the flightless, ground-dwelling kiwi bird of New Zealand, which feeds on worms and other ground creatures, and lives its life in

1890-432: Is to a large degree vacant, i.e., that there are many vacant niches . It is based on many empirical studies and theoretical investigations especially of Kauffman 1993. Causes of vacant niches may be evolutionary contingencies or brief or long-lasting environmental disturbances. Both paradigms agree that species are never "universal" in the sense that they occupy all possible niches; they are always specialized, although

1953-546: The Sonoran Desert , some annual plants are more successful during wet years, while others are more successful during dry years. As a result, each species will have an advantage in some years, but not others. When environmental conditions are most favorable, individuals will tend to compete most strongly with member of the same species. For example, in a dry year, dry-adapted plants will tend to be most limited by other dry-adapted plants. This can help them to coexist through

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2016-473: The "impact niche" is defined as the combination of effects that a given consumer has on both a). the resources that it uses, and b). the other consumers in the ecosystem. Therefore, the impact niche is equivalent to the Eltonian niche since both concepts are defined by the impact of a given species on its environment. The range of environmental conditions where a species can successfully survive and reproduce (i.e.

2079-495: The Eltonian niche introduces the idea of a species' response to and effect on the environment. Unlike other niche concepts, it emphasizes that a species not only grows in and responds to an environment based on available resources, predators, and climatic conditions, but also changes the availability and behavior of those factors as it grows. In an extreme example, beavers require certain resources in order to survive and reproduce, but also construct dams that alter water flow in

2142-567: The Hutchinsonian definition of a realized niche) is also encompassed under contemporary niche theory, termed the "requirement niche". The requirement niche is bounded by both the availability of resources as well as the effects of coexisting consumers (e.g. competitors and predators). Contemporary niche theory provides three requirements that must be met in order for two species (consumers) to coexist: These requirements are interesting and controversial because they require any two species to share

2205-811: The behavior of the California thrasher is consistent with the chaparral habitat it lives in—it breeds and feeds in the underbrush and escapes from its predators by shuffling from underbrush to underbrush. Its 'niche' is defined by the felicitous complementing of the thrasher's behavior and physical traits (camouflaging color, short wings, strong legs) with this habitat. Grinnellian niches can be defined by non-interactive (abiotic) variables and environmental conditions on broad scales. Variables of interest in this niche class include average temperature, precipitation, solar radiation, and terrain aspect which have become increasingly accessible across spatial scales. Most literature has focused on Grinnellian niche constructs, often from

2268-441: The coordinate system." The niche concept was popularized by the zoologist G. Evelyn Hutchinson in 1957. Hutchinson inquired into the question of why there are so many types of organisms in any one habitat. His work inspired many others to develop models to explain how many and how similar coexisting species could be within a given community, and led to the concepts of 'niche breadth' (the variety of resources or habitats used by

2331-481: The ecological space occupied by a species) is subtly different from the "niche" as defined by Grinnell (an ecological role, that may or may not be actually filled by a species—see vacant niches ). A niche is a very specific segment of ecospace occupied by a single species. On the presumption that no two species are identical in all respects (called Hardin's 'axiom of inequality' ) and the competitive exclusion principle , some resource or adaptive dimension will provide

2394-404: The environment differently in a way that helps them to coexist Niche (protein structural motif) Stem-cell niche , the necessary cellular environment of a stem cell Business [ edit ] Niche market , a focused, targetable portion (subset) of a market sector Niche blogging , a blog focused on a niche market Other uses [ edit ] Bassline (music genre) ,

2457-796: The environment. As an example of niche partitioning, several anole lizards in the Caribbean islands share common diets—mainly insects. They avoid competition by occupying different physical locations. Although these lizards might occupy different locations, some species can be found inhabiting the same range, with up to 15 in certain areas. For example, some live on the ground while others are arboreal. Species who live in different areas compete less for food and other resources, which minimizes competition between species. However, species who live in similar areas typically compete with each other. The Lotka–Volterra equation states that two competing species can coexist when intra-specific (within species) competition

2520-463: The estimation of the competition coefficients. This postulate, however, can be misguided, as it ignores the impacts that the resources of each category have on the organism and the impacts that the organism has on the resources of each category. For instance, the resource in the overlap region can be non-limiting, in which case there is no competition for this resource despite niche overlap. An organism free of interference from other species could use

2583-574: The existence of both ecological equivalents and empty niches. An ecological equivalent to an organism is an organism from a different taxonomic group exhibiting similar adaptations in a similar habitat, an example being the different succulents found in American and African deserts, cactus and euphorbia , respectively. As another example, the anole lizards of the Greater Antilles are a rare example of convergent evolution , adaptive radiation , and

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2646-482: The existence of ecological equivalents: the anole lizards evolved in similar microhabitats independently of each other and resulted in the same ecomorphs across all four islands. In 1927 Charles Sutherland Elton , a British ecologist , defined a niche as follows: "The 'niche' of an animal means its place in the biotic environment, its relations to food and enemies ." Elton classified niches according to foraging activities ("food habits"): For instance there

2709-472: The exotic or invasive species . The mathematical representation of a species' fundamental niche in ecological space, and its subsequent projection back into geographic space, is the domain of niche modelling . Contemporary niche theory (also called "classic niche theory" in some contexts) is a framework that was originally designed to reconcile different definitions of niches (see Grinnellian, Eltonian, and Hutchinsonian definitions above), and to help explain

2772-427: The extinction of the tarpan has been filled by other animals (in particular a small horse breed, the konik ). Also, when plants and animals are introduced into a new environment, they have the potential to occupy or invade the niche or niches of native organisms, often outcompeting the indigenous species. Introduction of non-indigenous species to non-native habitats by humans often results in biological pollution by

2835-434: The figure, where it is clear that for the narrower distributions (top) there is no competition for prey between the extreme left and extreme right species, while for the broader distribution (bottom), niche overlap indicates competition can occur between all species. The resource-utilization approach postulates that not only can competition occur, but that it does occur, and that overlap in resource utilization directly enables

2898-502: The first to use it in a research program in 1917, in his paper "The niche relationships of the California Thrasher". The Grinnellian niche concept embodies the idea that the niche of a species is determined by the habitat in which it lives and its accompanying behavioral adaptations . In other words, the niche is the sum of the habitat requirements and behaviors that allow a species to persist and produce offspring. For example,

2961-433: The full range of conditions (biotic and abiotic) and resources in which it could survive and reproduce which is called its fundamental niche . However, as a result of pressure from, and interactions with, other organisms (i.e. inter-specific competition) species are usually forced to occupy a niche that is narrower than this, and to which they are mostly highly adapted ; this is termed the realized niche . Hutchinson used

3024-401: The idea of competition for resources as the primary mechanism driving ecology, but overemphasis upon this focus has proved to be a handicap for the niche concept. In particular, overemphasis upon a species' dependence upon resources has led to too little emphasis upon the effects of organisms on their environment, for instance, colonization and invasions. The term "adaptive zone" was coined by

3087-475: The narrow extent of focus, data sets characterizing Eltonian niches typically are in the form of detailed field studies of specific individual phenomena, as the dynamics of this class of niche are difficult to measure at a broad geographic scale. However, the Eltonian niche may be useful in the explanation of a species' endurance of global change. Because adjustments in biotic interactions inevitably change abiotic factors, Eltonian niches can be useful in describing

3150-402: The other to extinction. This rule also states that two species cannot occupy the same exact niche in a habitat and coexist together, at least in a stable manner. When two species differentiate their niches, they tend to compete less strongly, and are thus more likely to coexist. Species can differentiate their niches in many ways, such as by consuming different foods, or using different areas of

3213-561: The overall response of a species to new environments. The Hutchinsonian niche is an " n-dimensional hypervolume", where the dimensions are environmental conditions and resources , that define the requirements of an individual or a species to practice its way of life, more particularly, for its population to persist. The "hypervolume" defines the multi-dimensional space of resources (e.g., light, nutrients, structure, etc.) available to (and specifically used by) organisms, and "all species other than those under consideration are regarded as part of

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3276-441: The paleontologist George Gaylord Simpson to explain how a population could jump from one niche to another that suited it, jump to an 'adaptive zone', made available by virtue of some modification, or possibly a change in the food chain , that made the adaptive zone available to it without a discontinuity in its way of life because the group was 'pre-adapted' to the new ecological opportunity. Hutchinson's "niche" (a description of

3339-410: The past, several species inhabited an area, and all of these species had overlapping fundamental niches. However, through competitive exclusion, the less competitive species were eliminated, leaving only the species that were able to coexist (i.e. the most competitive species whose realized niches did not overlap). Again, this process does not include any evolutionary change of individual species, but it

3402-454: The presence of niche differentiation (through competition) will be relatively easy. Importantly, there is no evolutionary change of the individual species in this case; rather this is an ecological effect of species Y out-competing species X within the bounds of species Y's fundamental niche. Another way by which niche differentiation can arise is via the previous elimination of species without realized niches. This asserts that at some point in

3465-565: The river where the beaver lives. Thus, the beaver affects the biotic and abiotic conditions of other species that live in and near the watershed. In a more subtle case, competitors that consume resources at different rates can lead to cycles in resource density that differ between species. Not only do species grow differently with respect to resource density, but their own population growth can affect resource density over time . Eltonian niches focus on biotic interactions and consumer–resource dynamics (biotic variables) on local scales. Because of

3528-658: The root meristem in that direction, if the interaction is kin. Simonsen discusses how plants accomplish root communication with the addition of beneficial rhizobia and fungal networks and the potential for different genotypes of the kin plants, such as the legume M. Lupulina, and specific strains of nitrogen fixing bacteria and rhizomes can alter relationships between kin and non-kin competition. This means there could be specific subsets of genotypes in kin plants that selects well with specific strains that could outcompete other kin. What might seem like an instance in kin competition could just be different genotypes of organisms at play in

3591-403: The root secretions, also called exudates, plants can make this determination. The communication between plants starts with the secretions from plant roots into the rhizosphere. If another plant that is kin is entering this area the plant will take up exudates. The exudate, being several different compounds, will enter the plants root cell and attach to a receptor for that chemical halting growth of

3654-476: The same resources if each species is limited by different resources, or differently able to capture resources. Different types of phytoplankton can coexist when different species are differently limited by nitrogen, phosphorus, silicon, and light. In the Galapagos Islands , finches with small beaks are more able to consume small seeds, and finches with large beaks are more able to consume large seeds. If

3717-472: The same ways as the incoming species, however examples of this are also numerous. In ecology , niche differentiation (also known as niche segregation , niche separation and niche partitioning ) refers to the process by which competing species use the environment differently in a way that helps them to coexist. The competitive exclusion principle states that if two species with identical niches (ecological roles) compete , then one will inevitably drive

3780-457: The soil that increase the symbiotic efficiency. Predator partitioning occurs when species are attacked differently by different predators (or natural enemies more generally). For example, trees could differentiate their niche if they are consumed by different species of specialist herbivores , such as herbivorous insects. If a species density declines, so too will the density of its natural enemies, giving it an advantage. Thus, if each species

3843-463: The title Niche . If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Niche&oldid=1258203280 " Category : Disambiguation pages Hidden categories: Short description is different from Wikidata All article disambiguation pages All disambiguation pages Ecological niche A Grinnellian niche

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3906-809: The two species interact, how they use their resources, and the type of ecosystem in which they exist, among other factors. In addition, several mathematical models exist to quantify niche breadth, competition, and coexistence (Bastolla et al. 2005). However, regardless of methods used, niches and competition can be distinctly difficult to measure quantitatively, and this makes detection and demonstration of niche differentiation difficult and complex. Over time, two competing species can either coexist, through niche differentiation or other means, or compete until one species becomes locally extinct . Several theories exist for how niche differentiation arises or evolves given these two possible outcomes. Niche differentiation can arise from current competition. For instance, species X has

3969-442: The underlying processes that affect Lotka-Volterra relationships within an ecosystem. The framework centers around "consumer-resource models" which largely split a given ecosystem into resources (e.g. sunlight or available water in soil) and consumers (e.g. any living thing, including plants and animals), and attempts to define the scope of possible relationships that could exist between the two groups. In contemporary niche theory,

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