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91-753: MGEX has been the principal market for Hard Red Spring Wheat (HRSW) since 1881, offering futures and options contracts based on its unique commodity. HRSW is one of the highest- protein wheats. It is found in bagels , pizzas , high-quality breads and cereals , and some noodles and cookies . It is planted mostly in the U.S. Northern Plains and the Canadian Prairies . MGEX offers five financially settled agricultural index products: Hard Red Spring Wheat Index (HRSI), Hard Red Winter Wheat Index (HRWI), Soft Red Winter Wheat Index (SRWI), National Corn Index (NCI) and National Soybean Index (NSI). In an agreement with Data Transmission Network (now Telvent DTN),

182-617: A business-to-business electronic commerce and information services company in Omaha, Nebraska , MGEX has exclusive rights to DTNs agriculture and weather data bases, which the exchange uses to develop index products. Futures are traded exclusively electronically on the CME Globex platform. Options are traded side-by-side. Founded as the Minneapolis Chamber of Commerce in 1881, the MGEX has been

273-469: A supplement of an individual amino acid (methionine, lysine, threonine, or tryptophan) can be added to the feed. Protein content in foods is often measured in protein per serving rather than protein per calorie. For instance, the USDA lists 6 grams of protein per large whole egg (a 50-gram serving) rather than 84 mg of protein per calorie (71 calories total). For comparison, there are 2.8 grams of protein in

364-484: A body. A positive balance occurs when more nitrogen is consumed than is excreted, which indicates that some of the nitrogen is being used by the body to build proteins. A negative nitrogen balance occurs when more nitrogen is excreted than is consumed, which indicates that there is insufficient intake for the body to maintain its health. Graduate students at the University of Illinois were fed an artificial diet so that there

455-438: A combination of artificial and natural selection . This complexity and diversity of status has led to much confusion in the naming of wheats. Hexaploid species (6N) Tetraploid species (4N) Diploid species (2N) The wild species of wheat, along with the domesticated varieties einkorn , emmer and spelt , have hulls. This more primitive morphology (in evolutionary terms) consists of toughened glumes that tightly enclose

546-439: A day to have a healthy protein profile, and almost 6 kg to get enough calories. It is recommended that adult humans obtain between 10–35% of their 2000 calories a day as protein. Scientists had known since the early 20th century that rats could not survive on a diet whose only protein source was zein , which comes from maize (corn), but recovered if they were fed casein from cow's milk. This led William Cumming Rose to

637-530: A decline in climate resilience of wheat has been observed. Hunter-gatherers in West Asia harvested wild wheats for thousands of years before they were domesticated , perhaps as early as 21,000 BC, but they formed a minor component of their diets. In this phase of pre-domestication cultivation, early cultivars were spread around the region and slowly developed the traits that came to characterise their domesticated forms. Repeated harvesting and sowing of

728-408: A diet containing no protein and the nitrogen losses recorded. During the first week or more there is a rapid loss of labile proteins. Once the nitrogen losses stabilize, this baseline is determined to be the minimum required for maintenance. Then the test subjects were fed a measured amount of the food being tested. The difference between the nitrogen in that food and the nitrogen losses above baseline

819-409: A good crop of grain. The farmer may intervene while the crop is growing to add fertilizer , water by irrigation , or pesticides such as herbicides to kill broad-leaved weeds or insecticides to kill insect pests. The farmer may assess soil minerals, soil water, weed growth, or the arrival of pests to decide timely and cost-effective corrective actions, and crop ripeness and water content to select

910-461: A less reliable method based on grain size. Einkorn and emmer are considered two of the founder crops cultivated by the first farming societies in Neolithic West Asia. These communities also cultivated naked wheats ( T. aestivum and T. durum ) and a now-extinct domesticated form of Zanduri wheat ( T. timopheevii ), as well as a wide variety of other cereal and non-cereal crops. Wheat

1001-677: A low quality for human nutrition, according to the DIAAS protein quality evaluation method. Though they contain adequate amounts of the other essential amino acids, at least for adults, wheat proteins are deficient in the essential amino acid lysine . Because the proteins present in the wheat endosperm (gluten proteins) are particularly poor in lysine, white flours are more deficient in lysine compared with whole grains. Significant efforts in plant breeding are made to develop lysine-rich wheat varieties, without success, as of 2017 . Supplementation with proteins from other food sources (mainly legumes )

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1092-450: A major contribution to the nutritional welfare of the poor. Some wheat species are diploid , with two sets of chromosomes , but many are stable polyploids , with four sets of chromosomes ( tetraploid ) or six ( hexaploid ). Einkorn wheat ( Triticum monococcum ) is diploid (AA, two complements of seven chromosomes, 2n=14). Most tetraploid wheats (e.g. emmer and durum wheat) are derived from wild emmer , T. dicoccoides . Wild emmer

1183-475: A major contributor to increased grain output. More widely over the last 40 years, a massive increase in fertilizer use together with the increased availability of semi-dwarf varieties in developing countries, has greatly increased yields per hectare. In developing countries, use of (mainly nitrogenous) fertilizer increased 25-fold in this period. However, farming systems rely on much more than fertilizer and breeding to improve productivity. A good illustration of this

1274-577: A marketplace for producers, processors and millers for more than 125 years. The three Grain Exchange buildings in downtown Minneapolis are now listed on the National Register of Historic Places . In 1883, the Chamber of Commerce introduced its first futures contract: hard red spring wheat. By 1946 "Chamber of Commerce" had become synonymous with organizations devoted mainly to civic and social issues. In 1947,

1365-579: A serving of raw broccoli (100 grams) or 82 mg of protein per calorie (34 calories total), or the Daily Value of 47.67g of protein after eating 1,690g of raw broccoli a day at 574 cal. An egg contains 12.5g of protein per 100g, but 4 mg more protein per calorie, or the protein DV after 381g of egg, which is 545 cal. The ratio of essential amino acids (the quality of protein) is not taken into account, one would actually need to eat more than 3 kg of broccoli

1456-470: A source of straw , which could be used for fuel, wicker-making , or wattle and daub construction. Domestic wheat was quickly spread to regions where its wild ancestors did not grow naturally. Emmer was introduced to Cyprus as early as 8600 BC and einkorn c.  7500 BC ; emmer reached Greece by 6500 BC, Egypt shortly after 6000 BC, and Germany and Spain by 5000 BC. "The early Egyptians were developers of bread and

1547-426: A viable crop. When the use of seed drills replaced broadcasting sowing of seed in the 18th century, another great increase in productivity occurred. Yields of pure wheat per unit area increased as methods of crop rotation were applied to land that had long been in cultivation, and the use of fertilizers became widespread. Improved agricultural husbandry has more recently included pervasive automation , starting with

1638-439: A wheat grain or berry; botanically a fruit , it is often called a seed. The grains ripen to a golden yellow; a head of grain is called an ear. Leaves emerge from the shoot apical meristem in a telescoping fashion until the transition to reproduction i.e. flowering. The last leaf produced by a wheat plant is known as the flag leaf. It is denser and has a higher photosynthetic rate than other leaves, to supply carbohydrate to

1729-506: A year's supply. Wheat grain was stored after threshing , with the chaff removed. It was then processed into flour using ground stone mortars . Bread made from ground einkorn and the tubers of a form of club rush ( Bolboschoenus glaucus ) was made as early as 12,400 BC. At Çatalhöyük ( c.  7100–6000 BC ), both wholegrain wheat and flour was used to prepare bread, porridge and gruel . Apart from food, wheat may also have been important to Neolithic societies as

1820-431: Is 10% to 20% higher than adult levels and those for infants can be as much as 150% higher in the first year of life. Cysteine (or sulfur-containing amino acids), tyrosine (or aromatic amino acids), and arginine are always required by infants and growing children. Methionine and cysteine are grouped together because one of them can be synthesized from the other using the enzyme methionine S -methyltransferase and

1911-452: Is Australian wheat growing in the southern winter cropping zone, where, despite low rainfall (300 mm), wheat cropping is successful even with relatively little use of nitrogenous fertilizer. This is achieved by crop rotation with leguminous pastures. The inclusion of a canola crop in the rotations has boosted wheat yields by a further 25%. In these low rainfall areas, better use of available soil-water (and better control of soil erosion)

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2002-512: Is a rich source (20% or more of the Daily Value , DV) of multiple dietary minerals , such as manganese , phosphorus , magnesium , zinc , and iron (table). The B vitamins , niacin (36% DV), thiamine (33% DV), and vitamin B6 (23% DV), are present in significant amounts (table). Wheat is a significant source of vegetable proteins in human food, having a relatively high protein content compared to other major cereals. However, wheat proteins have

2093-400: Is achieved by retaining the stubble after harvesting and by minimizing tillage. Pests and diseases consume 21.47% of the world's wheat crop annually. There are many wheat diseases, mainly caused by fungi, bacteria, and viruses . Plant breeding to develop new disease-resistant varieties, and sound crop management practices are important for preventing disease. Fungicides, used to prevent

2184-455: Is also important to identify periods of higher risk from the climate. Farmers benefit from knowing when the 'flag leaf' (last leaf) appears, as this leaf represents about 75% of photosynthesis reactions during the grain filling period, and so should be preserved from disease or insect attacks to ensure a good yield. Several systems exist to identify crop stages, with the Feekes and Zadoks scales being

2275-405: Is an important source of carbohydrates . Globally, it is the leading source of vegetable proteins in human food, having a protein content of about 13%, which is relatively high compared to other major cereals but relatively low in protein quality (supplying essential amino acids ). When eaten as the whole grain , wheat is a source of multiple nutrients and dietary fiber . In a small part of

2366-402: Is commonly used to compensate for this deficiency, since the limitation of a single essential amino acid causes the others to break down and become excreted, which is especially important during growth. Consumed worldwide by billions of people, wheat is a significant food for human nutrition, particularly in the least developed countries where wheat products are primary foods. When eaten as

2457-418: Is greater than for all other crops combined. Global demand for wheat is increasing due to the unique viscoelastic and adhesive properties of gluten proteins, which facilitate the production of processed foods, whose consumption is increasing as a result of the worldwide industrialization process and westernization of diets . Wheat became a central agriculture endeavor in the worldwide British Empire in

2548-473: Is itself the result of a hybridization between two diploid wild grasses, T. urartu and a wild goatgrass such as Ae. speltoides . The hybridization that formed wild emmer (AABB, four complements of seven chromosomes in two groups, 4n=28) occurred in the wild, long before domestication, and was driven by natural selection . Hexaploid wheats evolved in farmers' fields as wild emmer hybridized with another goatgrass, Ae. squarrosa or Ae. tauschii , to make

2639-629: Is likely to be prioritised in drought-adapted crops, while stem non-structural carbohydrate is prioritised in varieties developed for countries where disease is a bigger issue. Depending on variety, wheat may be awned or not awned. Producing awns incurs a cost in grain number, but wheat awns photosynthesise more efficiently than their leaves with regards to water usage, so awns are much more frequent in varieties of wheat grown in hot drought-prone countries than those generally seen in temperate countries. For this reason, awned varieties could become more widely grown due to climate change . In Europe, however,

2730-458: Is more diverse, with domesticated varieties falling into two major groups: hulled or non-shattering, in which threshing separates the whole spikelet ; and free-threshing, where the individual grains are separated. Both varieties probably existed in prehistory, but over time free-threshing cultivars became more common. Wild emmer was first cultivated in the southern Levant , as early as 9600 BC. Genetic studies have found that, like einkorn, it

2821-399: Is normally planted after tilling the soil by ploughing and then harrowing to kill weeds and create an even surface. The seeds are then scattered on the surface, or drilled into the soil in rows. Winter wheat lies dormant during a winter freeze. It needs to develop to a height of 10 to 15 cm before the cold intervenes, so as to be able to survive the winter; it requires a period with

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2912-511: Is not the same as a wheat allergy . Other diseases triggered by eating wheat are non-coeliac gluten sensitivity (estimated to affect 0.5% to 13% of the general population ), gluten ataxia , and dermatitis herpetiformis . Certain short-chain carbohydrates present in wheat, known as FODMAPs (mainly fructose polymers ), may be the cause of non-coeliac gluten sensitivity. As of 2019 , reviews have concluded that FODMAPs only explain certain gastrointestinal symptoms, such as bloating , but not

3003-448: Is somewhat unclear, as some amino acids can be produced from others. The sulfur -containing amino acids, methionine and homocysteine , can be converted into each other but neither can be synthesized de novo in humans. Likewise, cysteine can be made from homocysteine but cannot be synthesized on its own. So, for convenience, sulfur-containing amino acids are sometimes considered a single pool of nutritionally equivalent amino acids as are

3094-411: Is synthesized by the urea cycle to meet the needs of an adult but perhaps not those of a growing child. Amino acids that must be obtained from the diet are called essential amino acids . Eukaryotes can synthesize some of the amino acids from other substrates . Consequently, only a subset of the amino acids used in protein synthesis are essential nutrients . Nonessential amino acids are produced in

3185-405: Is synthesized from NH 4 and glutamate, and asparagine is synthesized similarly. Proline and arginine are both derived from glutamate. Serine , formed from 3-phosphoglycerate , which comes from glycolysis , is the precursor of glycine and cysteine . Tyrosine is synthesized by the hydroxylation of phenylalanine , which is an essential amino acid. Estimating the daily requirement for

3276-493: Is valuable to impart viscoelastic functional qualities in dough , enabling the preparation of diverse processed foods such as breads, noodles, and pasta that facilitate wheat consumption. Raw red winter wheat is 13% water, 71% carbohydrates including 12% dietary fiber , 13% protein , and 2% fat (table). Some 75–80% of the protein content is as gluten . In a reference amount of 100 grams (3.5 oz), wheat provides 1,368 kilojoules (327 kilocalories) of food energy and

3367-426: The aromatic amino acid pair, phenylalanine and tyrosine . Likewise arginine , ornithine , and citrulline , which are interconvertible by the urea cycle , are considered a single group. If one of the essential amino acids is not available in the required quantities, protein synthesis will be inhibited, irrespective of the availability of the other amino acids. Protein deficiency has been shown to affect all of

3458-436: The extra-digestive symptoms that people with non-coeliac gluten sensitivity may develop. Other wheat proteins, amylase-trypsin inhibitors, have been identified as the possible activator of the innate immune system in coeliac disease and non-coeliac gluten sensitivity. These proteins are part of the plant's natural defense against insects and may cause intestinal inflammation in humans. In 2022, world wheat production

3549-853: The hexaploid wheats including bread wheat . A 2007 molecular phylogeny of the wheats gives the following not fully-resolved cladogram of major cultivated species; the large amount of hybridisation makes resolution difficult. Markings like "6N" indicate the degree of polyploidy of each species: Barley 2N, rye 2N/4N, and other cereals Triticum monococcum ( einkorn ) 2N × Aegilotriticum hybrids ( Aegilops x Triticum ) 6N Triticum timopheevii (zanduri wheat) and others 4N Triticum aestivum ( common or bread wheat ) 6N Triticum durum/turgidum ( durum wheat) 4N Triticum spelta ( spelt ) 6N Triticum turanicum ( khorasan wheat ) 4N Triticum dicoccum ( emmer ) 4N many other species During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids , have developed under

3640-427: The whole grain , wheat supplies multiple nutrients and dietary fiber recommended for children and adults. In genetically susceptible people, wheat gluten can trigger coeliac disease . Coeliac disease affects about 1% of the general population in developed countries . The only known effective treatment is a strict lifelong gluten-free diet . While coeliac disease is caused by a reaction to wheat proteins, it

3731-550: The "quality" or "value" of various kinds of protein. Measures include the biological value , net protein utilization , protein efficiency ratio , protein digestibility corrected amino acid score and the complete proteins concept. These concepts are important in the livestock industry , because the relative lack of one or more of the essential amino acids in animal feeds would have a limiting effect on growth and thus on feed conversion ratio . Thus, various feedstuffs may be fed in combination to increase net protein utilization, or

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3822-605: The 19th century, and remains of great importance in Australia, Canada and India. In Australia, with vast lands and a limited work force, expanded production depended on technological advances, especially regarding irrigation and machinery. By the 1840s there were 900 growers in South Australia . They used "Ridley's Stripper", a reaper-harvester perfected by John Ridley in 1843, to remove the heads of grain. In Canada, modern farm implements made large scale wheat farming possible from

3913-513: The 20th century, global wheat output expanded by about 5-fold, but until about 1955 most of this reflected increases in wheat crop area, with lesser (about 20%) increases in crop yields per unit area. After 1955 however, there was a ten-fold increase in the rate of wheat yield improvement per year, and this became the major factor allowing global wheat production to increase. Thus technological innovation and scientific crop management with synthetic nitrogen fertilizer , irrigation and wheat breeding were

4004-454: The 22nd amino acid), which is proteinogenic only in certain microorganisms, is not used by and therefore non-essential for most organisms, including humans. The limiting amino acid is the essential amino acid which is furthest from meeting nutritional requirements. This concept is important when determining the selection, number, and amount of foods to consume because even when total protein and all other essential amino acids are satisfied if

4095-459: The actual amino acid content of foods. The USDA used this technique in their own labs to determine the content of 7793 foods across 28 categories. The USDA published the final database in 2018 to the public. The limiting amino acid depends on the human requirements and there are currently two sets of human requirements from authoritative sources: one published by WHO and the other published by USDA . Various attempts have been made to express

4186-427: The amino acids arginine , cysteine , glutamine , glycine , proline and tyrosine are considered conditionally essential , which means that specific populations who do not synthesize it in adequate amounts, such as newborn infants and people with diseased livers who are unable to synthesize cysteine, must obtain one or more of these conditionally essential amino acids from their diet. For example, enough arginine

4277-653: The body's organs and many of its systems, for example affecting brain development in infants and young children; inhibiting upkeep of the immune system, increasing risk of infection; affecting gut mucosal function and permeability, thereby reducing absorption and increasing vulnerability to systemic disease ; and impacting kidney function. The physical signs of protein deficiency include edema , failure to thrive in infants and children, poor musculature, dull skin, and thin and fragile hair. Biochemical changes reflecting protein deficiency include low serum albumin and low serum transferrin . The amino acids that are essential in

4368-466: The body. The pathways for the synthesis of nonessential amino acids come from basic metabolic pathways. Glutamate dehydrogenase catalyzes the reductive amination of α-ketoglutarate to glutamate . A transamination reaction takes place in the synthesis of most amino acids. At this step, the chirality of the amino acid is established. Alanine and aspartate are synthesized by the transamination of pyruvate and oxaloacetate , respectively. Glutamine

4459-420: The catalyst methionine synthase . Phenylalanine and tyrosine are grouped together because tyrosine can be synthesized from phenylalanine using the enzyme phenylalanine hydroxylase . Historically, amino acid requirements were determined by calculating the balance between dietary nitrogen intake and nitrogen excreted in the liquid and solid wastes, because proteins represent the largest nitrogen content in

4550-414: The deepest of arable crops, extending as far down as 2 metres (6 ft 7 in). While the roots of a wheat plant are growing, the plant also accumulates an energy store in its stem, in the form of fructans , which helps the plant to yield under drought and disease pressure, but it has been observed that there is a trade-off between root growth and stem non-structural carbohydrate reserves. Root growth

4641-686: The developing ear. In temperate countries the flag leaf, along with the second and third highest leaf on the plant, supply the majority of carbohydrate in the grain and their condition is paramount to yield formation. Wheat is unusual among plants in having more stomata on the upper ( adaxial ) side of the leaf, than on the under ( abaxial ) side. It has been theorised that this might be an effect of it having been domesticated and cultivated longer than any other plant. Winter wheat generally produces up to 15 leaves per shoot and spring wheat up to 9 and winter crops may have up to 35 tillers (shoots) per plant (depending on cultivar). Wheat roots are among

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4732-585: The discovery of the essential amino acid threonine . Through manipulation of rodent diets, Rose was able to show that ten amino acids are essential for rats: lysine , tryptophan , histidine , phenylalanine , leucine , isoleucine , methionine , valine , and arginine , in addition to threonine. Rose's later work showed that eight amino acids are essential for adult human beings, with histidine also being essential for infants. Longer-term studies established histidine as also essential for adult humans. The distinction between essential and non-essential amino acids

4823-477: The domestic form comes after c.   8800 BC in southern Turkey, at Çayönü , Cafer Höyük , and possibly Nevalı Çori . Genetic evidence indicates that it was domesticated in multiple places independently. Wild emmer wheat ( T. turgidum subsp. dicoccoides ) is less widespread than einkorn, favouring the rocky basaltic and limestone soils found in the hilly flanks of the Fertile Crescent. It

4914-732: The earliest known wheat with sufficient gluten for yeasted breads was found in a granary at Assiros in Macedonia dated to 1350 BC. From the Middle East , wheat continued to spread across Europe and to the Americas in the Columbian exchange . In the British Isles, wheat straw ( thatch ) was used for roofing in the Bronze Age , and remained in common use until the late 19th century. White wheat bread

5005-507: The exchange was renamed the Minneapolis Grain Exchange. Today the exchange is most recognized by its logo and uses MGEX as first reference. On December 19, 2008, the Minneapolis Grain Exchange ceased operations of the open outcry trading floor, but continues daily operations for the electronic processing of financial transactions. Today, HRSW futures trade exclusively electronically and options trade side-by-side. In 2022, MGEX

5096-559: The farm and because of poor roads, inadequate storage technologies, inefficient supply chains and farmers' inability to bring the produce into retail markets dominated by small shopkeepers. Some 10% of total wheat production is lost at farm level, another 10% is lost because of poor storage and road networks, and additional amounts are lost at the retail level. In the Punjab region of the Indian subcontinent, as well as North China, irrigation has been

5187-456: The farmer have a detailed understanding of each stage of development in the growing plants. In particular, spring fertilizers , herbicides , fungicides , and growth regulators are typically applied only at specific stages of plant development. For example, it is currently recommended that the second application of nitrogen is best done when the ear (not visible at this stage) is about 1 cm in size (Z31 on Zadoks scale ). Knowledge of stages

5278-422: The general population, gluten – which comprises most of the protein in wheat – can trigger coeliac disease , noncoeliac gluten sensitivity , gluten ataxia , and dermatitis herpetiformis . Wheat is a stout grass of medium to tall height. Its stem is jointed and usually hollow, forming a straw. There can be many stems on one plant. It has long narrow leaves, their bases sheathing the stem, one above each joint. At

5369-594: The glumes are fragile and the rachis tough. On threshing, the chaff breaks up, releasing the grains. Wheat grain classes are named by colour, season, and hardness. The classes used in the United States are: Wheat is a staple cereal worldwide. Raw wheat berries can be ground into flour or, using hard durum wheat only, can be ground into semolina ; germinated and dried creating malt ; crushed or cut into cracked wheat; parboiled (or steamed), dried, crushed and de-branned into bulgur also known as groats . If

5460-562: The grains of wild grasses led to the creation of domestic strains, as mutant forms ('sports') of wheat were more amenable to cultivation. In domesticated wheat, grains are larger, and the seeds (inside the spikelets ) remain attached to the ear by a toughened rachis during harvesting. In wild strains, a more fragile rachis allows the ear to shatter easily, dispersing the spikelets. Selection for larger grains and non-shattering heads by farmers might not have been deliberately intended, but simply have occurred because these traits made gathering

5551-479: The grains, and (in domesticated wheats) a semi-brittle rachis that breaks easily on threshing. The result is that when threshed, the wheat ear breaks up into spikelets. To obtain the grain, further processing, such as milling or pounding, is needed to remove the hulls or husks. Hulled wheats are often stored as spikelets because the toughened glumes give good protection against pests of stored grain. In free-threshing (or naked) forms, such as durum wheat and common wheat,

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5642-486: The human diet were established in a series of experiments led by William Cumming Rose . The experiments involved elemental diets to healthy male graduate students. These diets consisted of corn starch , sucrose , butterfat without protein, corn oil , inorganic salts, the known vitamins , a large brown "candy" made of liver extract flavored with peppermint oil (to supply any unknown vitamins), and mixtures of highly purified individual amino acids. The main outcome measure

5733-406: The indispensable amino acids has proven to be difficult; these numbers have undergone considerable revision over the last 20 years. The following table lists the recommended daily amounts currently in use for essential amino acids in adult humans (unless specified otherwise), together with their standard one-letter abbreviations. The recommended daily intakes for children aged three years and older

5824-443: The infant or individuals in severe catabolic distress. These six are arginine , cysteine , glycine , glutamine , proline , and tyrosine . Six amino acids are non-essential ( dispensable ) in humans, meaning they can be synthesized in sufficient quantities in the body. These six are alanine , aspartic acid , asparagine , glutamic acid , serine , and selenocysteine (considered the 21st amino acid). Pyrrolysine (considered

5915-513: The late 1840s. By 1879, Saskatchewan was the center, followed by Alberta , Manitoba and Ontario , as the spread of railway lines allowed easy exports to Britain. By 1910, wheat made up 22% of Canada's exports, rising to 25% in 1930 despite the sharp decline in prices during the worldwide Great Depression . Efforts to expand wheat production in South Africa, Kenya and India were stymied by low yields and disease. However, by 2000 India had become

6006-464: The limiting amino acid is not satisfied then the meal is considered to be nutritionally limited by that amino acid. (*) Pyrrolysine , sometimes considered the "22nd amino acid", is not used by the human body. Of the twenty amino acids common to all life forms (not counting selenocysteine ), humans cannot synthesize nine: histidine , isoleucine , leucine , lysine , methionine , phenylalanine , threonine , tryptophan and valine . Additionally,

6097-476: The main drivers of wheat output growth in the second half of the century. There were some significant decreases in wheat crop area, for instance in North America. Better seed storage and germination ability (and hence a smaller requirement to retain harvested crop for next year's seed) is another 20th-century technological innovation. In Medieval England, farmers saved one-quarter of their wheat harvest as seed for

6188-568: The most widely used. Each scale is a standard system which describes successive stages reached by the crop during the agricultural season. For example, the stage of pollen formation from the mother cell, and the stages between anthesis and maturity, are susceptible to high temperatures, and this adverse effect is made worse by water stress. Technological advances in soil preparation and seed placement at planting time, use of crop rotation and fertilizers to improve plant growth, and advances in harvesting methods have all combined to promote wheat as

6279-488: The need to systematically remove vegetation or till the soil. They may also have exploited natural wetlands and floodplains to practice décrue farming , sowing seeds in the soil left behind by receding floodwater. It was harvested with stone-bladed sickles . The ease of storing wheat and other cereals led farming households to become gradually more reliant on it over time, especially after they developed individual storage facilities that were large enough to hold more than

6370-473: The next crop, leaving only three-quarters for food and feed consumption. By 1999, the global average seed use of wheat was about 6% of output. In the 21st century, rising temperatures associated with global warming are reducing wheat yield in several locations. Wheat is an annual crop. It can be planted in autumn and harvested in early summer as winter wheat in climates that are not too severe, or planted in spring and harvested in autumn as spring wheat. It

6461-496: The organism fast enough to supply its demand, and must therefore come from the diet. Of the 21 amino acids common to all life forms, the nine amino acids humans cannot synthesize are valine , isoleucine , leucine , methionine , phenylalanine , tryptophan , threonine , histidine , and lysine . Six other amino acids are considered conditionally essential in the human diet, meaning their synthesis can be limited under special pathophysiological conditions, such as prematurity in

6552-461: The raw wheat is broken into parts at the mill, as is usually done, the outer husk or bran can be used in several ways. Wheat is a major ingredient in such foods as bread , porridge , crackers , biscuits , muesli , pancakes , pasta , pies , pastries , pizza , semolina , cakes , cookies , muffins , rolls , doughnuts , gravy , beer , vodka , boza (a fermented beverage ), and breakfast cereals . In manufacturing wheat products, gluten

6643-452: The right moment to harvest. Harvesting involves reaping , cutting the stems to gather the crop; and threshing , breaking the ears to release the grain; both steps are carried out by a combine harvester . The grain is then dried so that it can be stored safe from mould fungi. Wheat normally needs between 110 and 130 days between sowing and harvest, depending upon climate, seed type, and soil conditions. Optimal crop management requires that

6734-507: The second largest producer of wheat in the world. In the 19th century the American wheat frontier moved rapidly westward. By the 1880s 70% of American exports went to British ports. The first successful grain elevator was built in Buffalo in 1842. The cost of transport fell rapidly. In 1869 it cost 37 cents to transport a bushel of wheat from Chicago to Liverpool . In 1905 it was 10 cents. In

6825-403: The second most-produced cereal after maize (known as corn in North America and Australia; wheat is often called corn in countries including Britain). Since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through the middle of the 21st century. Global demand for wheat is increasing because of the usefulness of gluten to the food industry. Wheat

6916-513: The seeds easier; nevertheless such 'incidental' selection was an important part of crop domestication . As the traits that improve wheat as a food source involve the loss of the plant's natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild. Wild einkorn wheat ( T. monococcum subsp. boeoticum ) grows across Southwest Asia in open parkland and steppe environments. It comprises three distinct races , only one of which, native to Southeast Anatolia ,

7007-511: The significant crop losses from fungal disease, can be a significant variable cost in wheat production. Estimates of the amount of wheat production lost owing to plant diseases vary between 10 and 25% in Missouri. A wide range of organisms infect wheat, of which the most important are viruses and fungi. Essential amino acid An essential amino acid , or indispensable amino acid , is an amino acid that cannot be synthesized from scratch by

7098-405: The temperature at or near freezing, its dormancy then being broken by the thaw or rise in temperature. Spring wheat does not undergo dormancy. Wheat requires a deep soil , preferably a loam with organic matter, and available minerals including soil nitrogen, phosphorus, and potassium. An acid and peaty soil is not suitable. Wheat needs some 30 to 38 cm of rain in the growing season to form

7189-514: The top of the stem is the flower head, containing some 20 to 100 flowers. Each flower contains both male and female parts. The flowers are wind-pollinated , with over 99% of pollination events being self-pollinations and the rest cross-pollinations . The flower is housed in a pair of small leaflike glumes . The two (male) stamens and (female) stigmas protrude outside the glumes. The flowers are grouped into spikelets , each with between two and six flowers. Each fertilised carpel develops into

7280-602: The use of threshing machines , and progressing to large and costly machines like the combine harvester which greatly increased productivity. At the same time, better varieties such as Norin 10 wheat , developed in Japan in the 1930s, or the dwarf wheat developed by Norman Borlaug in the Green Revolution , greatly increased yields. In addition to gaps in farming system technology and knowledge, some large wheat grain-producing countries have significant losses after harvest at

7371-531: The use of the oven and developed baking into one of the first large-scale food production industries." By 4000 BC, wheat had reached the British Isles and Scandinavia . Wheat likely appeared in China 's lower Yellow River around 2600 BC. The oldest evidence for hexaploid wheat has been confirmed through DNA analysis of wheat seeds, dating to around 6400–6200 BC, recovered from Çatalhöyük . As of 2023,

7462-537: Was nitrogen balance . Rose noted that the symptoms of nervousness, exhaustion, and dizziness were encountered to a greater or lesser extent whenever human subjects were deprived of an essential amino acid. Essential amino acid deficiency should be distinguished from protein-energy malnutrition , which can manifest as marasmus or kwashiorkor . Kwashiorkor was once attributed to pure protein deficiency in individuals who were consuming enough calories ("sugar baby syndrome"). However, this theory has been challenged by

7553-486: Was 808.4 million tonnes, led by China, India, and Russia which collectively provided 43.22% of the world total. As of 2019 , the largest exporters were Russia (32 million tonnes), United States (27), Canada (23) and France (20), while the largest importers were Indonesia (11 million tonnes), Egypt (10.4) and Turkey (10.0). In 2021, wheat was grown on 220.7 million hectares or 545 million acres worldwide, more than any other food crop. World trade in wheat

7644-444: Was a slightly positive nitrogen balance. Then one amino acid was omitted and the nitrogen balance recorded. If a positive balance continued, then that amino acid was deemed not essential. If a negative balance occurred, then that amino acid was slowly restored until a slightly positive nitrogen balance stabilized and the minimum amount recorded. A similar method was used to determine the protein content of foods. Test subjects were fed

7735-458: Was domesticated in southeastern Anatolia, but only once. The earliest secure archaeological evidence for domestic emmer comes from Çayönü, c.  8300–7600 BC , where distinctive scars on the spikelets indicated that they came from a hulled domestic variety. Slightly earlier finds have been reported from Tell Aswad in Syria, c.  8500–8200 BC , but these were identified using

7826-429: Was domesticated. The main feature that distinguishes domestic einkorn from wild is that its ears do not shatter without pressure, making it dependent on humans for dispersal and reproduction. It also tends to have wider grains. Wild einkorn was collected at sites such as Tell Abu Hureyra ( c.  10,700–9000 BC ) and Mureybet ( c.  9800–9300 BC ), but the earliest archaeological evidence for

7917-505: Was first cultivated in the regions of the Fertile Crescent around 9600 BC. Botanically, the wheat kernel is a caryopsis , a type of fruit . Wheat is grown on a larger area of land than any other food crop (220.7 million hectares or 545 million acres in 2021). World trade in wheat is greater than for all other crops combined. In 2021, world wheat production was 771 million tonnes (850 million short tons), making it

8008-585: Was historically a high status food, but during the nineteenth century it became in Britain an item of mass consumption, displacing oats , barley and rye from diets in the North of the country. It became "a sign of a high degree of culture". After 1860, the enormous expansion of wheat production in the United States flooded the world market, lowering prices by 40%, and (along with the expansion of potato growing) made

8099-428: Was owned by Miami International Holdings . Wheat#Major cultivated species of wheat Wheat is a grass widely cultivated for its seed , a cereal grain that is a staple food around the world. The many species of wheat together make up the genus Triticum ( / ˈ t r ɪ t ɪ k ə m / ); the most widely grown is common wheat ( T. aestivum ). The archaeological record suggests that wheat

8190-451: Was relatively uncommon for the first thousand years of the Neolithic (when barley predominated), but became a staple after around 8500 BC. Early wheat cultivation did not demand much labour. Initially, farmers took advantage of wheat's ability to establish itself in annual grasslands by enclosing fields against grazing animals and re-sowing stands after they had been harvested, without

8281-402: Was the amount the body retained to rebuild proteins. The amount of nitrogen retained divided by the total nitrogen intake is called net protein utilization . The amount of nitrogen retained divided by the (nitrogen intake minus nitrogen loss above baseline) is called biological value and is usually given as a percentage. Modern techniques make use of ion exchange chromatography to determine

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