A distinct population segment ( DPS ) is the smallest division of a taxonomic species permitted to be protected under the U.S. Endangered Species Act . Species , as defined in the Act for listing purposes, is a taxonomic species or subspecies of plant or animal, or in the case of vertebrate species, a distinct population segment.
37-623: Tympanuchus cupido pallidicinctus The lesser prairie-chicken ( Tympanuchus pallidicinctus ) is a species in the grouse family. Adult individuals are medium to large birds, striped white and brown like its near relative, the Greater prairie-chicken ( T. cupido ), though it is smaller and paler. Adults range from 15.0 to 16.1 inches (38 to 41 cm) in length and 22.1 to 28.7 ounces (630 to 810 g) in weight. Mature males sport yellow, comb-like feathers above each eye, as well as long head feathers that can be raised to show their pinkish-red Gular sacs . One
74-516: A group of birds from the order Galliformes , in the family Phasianidae . Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae ), a classification supported by mitochondrial DNA sequence studies, and applied by the American Ornithologists' Union , ITIS , International Ornithological Congress , and others. Grouse inhabit temperate and subarctic regions of
111-508: A male defends a territory which has resources that females need, like food and nest sites. These differences in male behavior in mating systems account for the evolution of body size in grouse. Males of territorial species were smaller than those of exploded lekking species, and males of typical lekking species were the largest overall. The male birds that exhibit lekking behavior, and have to compete with other males for females to choose them, have greater sexual dimorphism in size. This suggests
148-555: A minimum population of the birds. There is evidence suggesting that global warming may have a particularly detrimental influence by greatly reducing the size of the sagebrush ecosystem. Subfossil remains are known, e.g., from Rocky Arroyo in the Guadalupe Mountains , outside the species' current range but where more habitat existed in the less humid conditions in the outgoing last ice age . Range contraction apparently took place no later than about 8000 BC. On March 27, 2014,
185-458: A population or group of populations as a DPS was most recently articulated in a 1996 joint USFWS - NMFS policy (61 FR 4722: February 7, 1996): Three elements are considered in a decision regarding the status of a possible DPS as endangered or threatened under the Act. These are applied similarly for addition to the lists of endangered and threatened wildlife and plants, reclassification, and removal from
222-595: A portion of Texas, as endangered. On November 17, 2022, the USFWS published a final rule listing the Southern Distinct Population Segment (DPS) of the lesser prairie-chicken as endangered and the Northern DPS as threatened. Implementation of the rule was delayed, and it took effect on March 27, 2023. In 2015, Senator Jerry Moran (R-Kan) introduced an amendment to legislation authorizing construction of
259-420: Is a shallow depression or scrape on the ground—often in cover—with a scanty lining of plant material. The female lays one clutch , but may replace it if the eggs are lost. She begins to lay about a week after mating and lays one egg every day or two; the clutch comprises five to 12 eggs. The eggs have the shape of hen's eggs and are pale yellow, sparsely spotted with brown. On laying the second-last or last egg,
296-542: Is called sexual dimorphism . Male grouse tend to be larger than female grouse, which seems to hold true across all the species of grouse, with some difference within each species in terms of how drastic the size difference is. The hypothesis with the most supporting evidence for the evolution of sexual dimorphism in grouse is sexual selection. Sexual selection favors large males; stronger selection for larger size in males leads to greater size dimorphism. Female size will increase correspondingly as male size increases, and this
333-509: Is due to heredity (but not to the extent of the male size). This is because females that are smaller will still be able to reproduce without a substantial disadvantage, but this is not the case with males. The largest among the male grouse (commonly dubbed 'Biggrouse') attract the greatest numbers of females during their mating seasons. Male grouse display lekking behavior, which is when many males come together in one area and put on displays to attract females. Females selectively choose among
370-577: Is present on each side of the neck, which becomes inflated to be used in mating displays. About half of its current population lives in western Kansas , with the other half in the sandhills and prairies of western Oklahoma , the Texas Panhandle including the Llano Estacado , eastern New Mexico , and southeastern Colorado . They prefer environments containing sand sagebrush and shinnery oak , as well as various grasses and shrubs depending on
407-487: The IUCN due to its restricted and patchy range, it is vulnerable to habitat destruction , due to a combination of droughts, oil contamination , and conversion to farmland. The lesser prairie-chicken's habitat has been reduced by 85%, and their population has declined by about 97% since 1800, in part due to unrestricted hunting. Of the remaining patches of suitable habitat, only around 0.1% are sufficiently contiguous to sustain even
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#1732852592471444-796: The Keystone XL Pipeline that would overturn the listing. He disputed the listing as, "... another example of unnecessary intrusion into private lives and businesses by the federal government." His action was supported by the American Energy Alliance and opposed by the League of Conservation Voters. When the Senate voted on the Keystone bill, it did not get the 60 votes in favor that was required to pass. It got only 53 Republican and one Democratic Senator to vote in favor. The United States Department of
481-597: The Northern Hemisphere , from pine forests to moorland and mountainside , from 83°N ( rock ptarmigan in northern Greenland ) to 28°N ( Attwater's prairie chicken in Texas ). The turkeys are closely allied with grouse, but they have traditionally been excluded from Tetraonini, often placed in their own tribe, subfamily, or family; certain more modern treatments also exclude them. Later phylogenomic analyses demonstrated conclusively that they are sister to
518-490: The ptarmigans , have legs which are entirely covered in feathers; in winter the toes, too, have feathers or small scales on the sides, an adaptation for walking on snow and burrowing into it for shelter. Unlike many other galliforms, they typically have no spurs , although turkeys do possess very prominent spurs. Grouse feed mainly on vegetation—buds, catkins , leaves, and twigs—which typically accounts for over 95% of adults' food by weight. Thus, their diets vary greatly with
555-695: The red grouse and the ruffed grouse have benefited from habitat management. Most grouse species are listed by the IUCN as "least concern" or "near threatened", but the greater and lesser prairie chicken are listed as "vulnerable" and the Gunnison grouse is listed as "endangered". Some subspecies, such as Attwater's prairie chicken and the Cantabrian capercaillie , and some national and regional populations are also in danger. The wild turkey precipitously declined before returning to abundance, even in developed areas. The phenotypic difference between males and females
592-669: The Act, NMFS has relied on the evolutionarily significant unit (ESU) concept and previous developed a policy on the definition of species under the Act (56 FR 58612– 58618; November 20, 1991). The policy applies only to species of salmonids native to the Pacific. Under this policy, a stock of Pacific salmon is considered a DPS if it represents an evolutionarily significant unit (ESU) of a biological species. A stock must satisfy two criteria to be considered an ESU: (1) It must be substantially reproductively isolated from other conspecific population units; and (2) It must represent an important component in
629-569: The House did on July 27. The bill was passed by the Senate, but vetoed by Biden on September 27, 2023. This Galliformes article is a stub . You can help Misplaced Pages by expanding it . Grouse Pucrasia Meleagris Bonasa Tetrastes Centrocercus Dendragapus Tympanuchus Lagopus Falcipennis Canachites Tetrao Lyrurus and see text Tetraonidae Vigors , 1825 Tetraoninae Vigors, 1825 Grouse / ɡ r aʊ s / are
666-525: The Interior proposed creating a lesser prairie-chicken preserve as a national monument , but action was never taken action on the proposal. In May 2023, the U.S. Senate voted to strip the lesser prairie-chicken of its new listing using its authority under the Congressional Review Act . President Joe Biden 's administration threatened to veto the bill if the House of Representatives passed it, which
703-475: The area. Like its larger relative, it is known for its lekking behavior , particularly the “booming” call produced during mating displays. They typically prefer wide, open areas such as hills and ridges with short grass for their leks, however the use of man-made structures such as roads or oil platforms has also been observed. During the months of April and May, males will establish an area for themselves and potential mates, and will typically become territorial as
740-469: The classes of information that might bear on the biological and ecological importance of a discrete population segment. If a population segment is discrete and significant (i.e., it is a distinct population segment) its evaluation for endangered or threatened status will be based on the Act's definitions of those terms and a review of the factors enumerated in section 4(a)of the Act. It may be appropriate to assign different classifications to different DPSes of
777-481: The conservation of genetic diversity. In carrying out this examination, the Services [USFWS and NMFS] will consider available scientific evidence of the discrete population segment's importance to the taxon to which it belongs. This consideration may include, but is not limited to, the following: Because precise circumstances are likely to vary considerably from case to case, it is not possible to describe prospectively all
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#1732852592471814-650: The evolutionary legacy of the species. The 1996 joint policy considers the 1991 NMFS policy on Pacific salmon to be consistent with the joint policy, being a detailed extension of it. The joint policy further states that NMFS will continue to exercise the 1991 policy with respect to Pacific salmonids. Waples, R.S. 1991. Definition of 'species' under the Endangered Species Act: Application to Pacific salmon. U.S. Dept. Commerce, NOAA Tech Memo. NMFS, F/NWC-194. United States Fish and Wildlife Service, and National Marine Fisheries Service. 1996. Policy regarding
851-404: The female starts 21 to 28 days of incubation. Chicks hatch in dense, yellow-brown down and leave the nest immediately. They soon develop feathers and can fly shortly before they are two weeks old. The female (and the male in the willow grouse) stays with them and protects them until their first autumn, when they reach their mature weights (except in the male capercaillies ). They are sexually mature
888-577: The following spring, but often do not mate until later years. Grouse make up a considerable part of the vertebrate biomass in the Arctic and Subarctic. Their numbers may fall sharply in years of bad weather or high predator populations—significant grouse populations are a major food source for lynx , foxes , martens , and birds of prey . The three tundra species have maintained their former numbers. The prairie and forest species have declined greatly because of habitat loss, though popular game birds such as
925-456: The ground at dawn and dusk, which in some are given in leks . The displays feature males' brightly colored combs and in some species, brightly colored inflatable sacs on the sides of their necks. The males display their plumage , give vocalizations that vary widely between species, and may engage in other activities, such as drumming or fluttering their wings, rattling their tails, and making display flights. Occasionally, males fight. The nest
962-411: The hypothesis of sexual selection affecting male body size and also gives an explanation for why some species of grouse have a more drastic difference between male and female body size than others. Sexual size dimorphism can manifest itself differently between grouse and other birds. In some cases, the female is dominant over the male in breeding behavior, which can result in females that are larger than
999-459: The largest grouse species, attaining lengths of 130 cm (50 in) and weighing up to 10 kg (22 lb). Male grouse are larger than females, and can be twice as heavy in the western capercaillie (the largest of the traditional grouse). Like many other galliforms, males often sport incredibly elaborate ornamentation, such as crests, fan-tails, and inflatable, brightly colored patches of bare skin. Many grouse have feathered nostrils, and some species, such as
1036-530: The lesser prairie-chicken was listed as threatened under the Endangered Species Act but the listing was vacated in 2015 following a legal challenge. On June 1, 2021, the US Fish and Wildlife Service (USFWS) proposed splitting the species into two segments. The northern one, covering Oklahoma, Colorado, Kansas, and a portion of Texas, would be listed as threatened, and the southern one, covering New Mexico and
1073-460: The lists: A population segment of a vertebrate species may be considered discrete if it satisfies either one of the following conditions: If a population segment is considered discrete under one or more of the above conditions, its biological and ecological significance will then be considered in light of Congressional guidance (see Senate Report 151, 96th Congress, 1st Session) that the authority to list DPSes be used "...sparingly" while encouraging
1110-425: The males present for traits they find more appealing. Male grouse exhibit two types: typical lekking and exploded lekking. In typical lekking, males display in small areas defending a limited territory, and in exploded lekking, displaying males are covered over an expansive land area and share larger territories. Male grouse can also compete with one another for access to female grouse through territoriality, in which
1147-669: The males. Grouse are game , and hunters kill millions each year for food, sport, and other uses. In the United Kingdom, this takes the form of driven grouse shooting . The male black grouse 's tail feathers are a traditional ornament for hats in areas such as Scotland and the Alps . Folk dances from the Alps to the North American prairies imitate the displays of lekking males. Distinct Population Segment The criteria for designation of
Lesser prairie-chicken - Misplaced Pages Continue
1184-612: The most social, forming flocks of up to 100 in winter. All grouse spend most of their time on the ground, though when alarmed, they may take off in a flurry and go into a long glide. Most species stay within their breeding range all year, but make short seasonal movements; many individuals of the ptarmigan (called rock ptarmigan in the US) and willow grouse (called willow ptarmigan in the US) migrate hundreds of kilometers. In all but one species (the willow ptarmigan ), males are polygamous . Many species have elaborate courtship displays on
1221-684: The mother hen leaves to molt . The majority of the lesser prairie-chicken’s diet consists of foraged seeds, insects, and crops, as well as leaves, fruits, and forb buds . All of these are eaten year-round based on availability, however adolescents under 10 weeks tend to eat mostly insects, and adults will as well during warmer months. In fall, they will mostly shift to eating seeds, as well as leaves and flowers in winter months. When available, crops such as corn and grains will be eaten if left out. Generally, this diet provides them with enough water to survive, and they will rarely drink directly without cover unless absolutely necessary. Considered "vulnerable" by
1258-456: The same vertebrate taxon. The Fish and Wildlife Service's Listing and Recovery Priority Guidelines (48 FR 43098; September 21, 1983) generally afford DPSes the same consideration as subspecies, but when a subspecies and a DPS have the same numerical priority, the subspecies receives higher priority for listing. The Services will continue to generally accord subspecies higher priority than DPSes. In consideration of Pacific salmon for listing under
1295-667: The seasons. Hatchlings eat mostly insects and other invertebrates , gradually reducing their proportion of animal food to adult levels. Several of the forest-living species are notable for eating large quantities of conifer needles, which most other vertebrates refuse. To digest vegetable food, grouse have big crops and gizzards , eat grit to break up food, and have long intestines with well-developed caeca in which symbiotic bacteria digest cellulose . Forest species flock only in autumn and winter, though individuals tolerate each other when they meet. Prairie species are more social, and tundra species (ptarmigans, Lagopus ) are
1332-790: The spring season progresses. This process is repeated to a lesser degree in the fall, beginning as early as late August and reaching a height in October. Males in particular are likely to return to the same territory repeatedly. Females will begin nesting following mating, most frequently in May. Nests are dug about 5-10 cm into the ground, with a width of up to 20cm. These nests are lined with grasses, leaves, and feathers, and may be located within taller grasses or beneath brush and shrubs for shade and shelter. A single brood usually contains 11-14 eggs, hatching about 23 days after being laid. Chicks become fledged within 4 weeks, and after 8-10, will be left on their own as
1369-616: The traditionally-defined grouse, and they, along with the somewhat earlier-diverging koklass pheasant , may be treated as grouse (i.e., as basal members of the Tetraonini). This is reflected in some more recent circumscriptions. Like many other galliforms, grouse are generally heavily-built birds. The traditional grouse (excluding turkeys) range in length from 31 to 95 cm (12 to 37 + 1 ⁄ 2 in), and in weight from 0.3 to 6.5 kg ( 3 ⁄ 4 to 14 + 1 ⁄ 4 lb). If they are included, wild turkey toms are
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