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Lengwe National Park

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Lengwe National Park is a national park in Malawi located near the town of Chikwawa and about 40 miles southwest of Blantyre . Lengwe's topography is unusual for Malawi and consists of open deciduous forests and dense thickets. It is the home of the reclusive Nyala antelope.

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89-479: The climate of Lengwe is hot and dry, and the only source of consistent water is from the rain. Many man-made water holes have been constructed to attract and maintain the animal population. Jambo Africa Ltd operates Nyala Lodge for visitors. Animals that inhabit this park are the giraffe , spotted hyena , leopard , warthog , kudu , impala , Samango monkey , reedbuck , Cape buffalo and suni . Interesting birds, particularly shrikes , rollers , bee-eaters and

178-455: A mutation (the deletion of two nucleotides ) that inactives it. These changes are explained by the fact that its prey does not need to be subdued. Several groups of predatory fish have the ability to detect, track, and sometimes, as in the electric ray , to incapacitate their prey by sensing and generating electric fields . The electric organ is derived from modified nerve or muscle tissue. Physiological adaptations to predation include

267-452: A 2024 study found that, while males have thicker necks, females actually have proportionally longer ones, which is likely because of their greater need to find more food to sustain themselves and their dependent young. It has also been proposed that the neck serves to give the animal greater vigilance. Prey Predation is a biological interaction where one organism, the predator , kills and eats another organism, its prey . It

356-418: A better choice. If it chooses pursuit, its physical capabilities determine the mode of pursuit (e.g., ambush or chase). Having captured the prey, it may also need to expend energy handling it (e.g., killing it, removing any shell or spines, and ingesting it). Predators have a choice of search modes ranging from sit-and-wait to active or widely foraging . The sit-and-wait method is most suitable if

445-851: A broad range of taxa including arthropods. They are common among insects, including mantids, dragonflies , lacewings and scorpionflies . In some species such as the alderfly , only the larvae are predatory (the adults do not eat). Spiders are predatory, as well as other terrestrial invertebrates such as scorpions ; centipedes ; some mites , snails and slugs ; nematodes ; and planarian worms . In marine environments, most cnidarians (e.g., jellyfish , hydroids ), ctenophora (comb jellies), echinoderms (e.g., sea stars , sea urchins , sand dollars , and sea cucumbers ) and flatworms are predatory. Among crustaceans , lobsters , crabs , shrimps and barnacles are predators, and in turn crustaceans are preyed on by nearly all cephalopods (including octopuses , squid and cuttlefish ). Seed predation

534-416: A difficult time giving birth to young with the same neck proportions as adults. The giraffe's head and neck are held up by large muscles and a nuchal ligament , which are anchored by long thoracic vertebrae spines, giving them a hump. The giraffe's neck vertebrae have ball and socket joints . The point of articulation between the cervical and thoracic vertebrae of giraffes is shifted to lie between

623-466: A disproportionate lengthening of the cervical vertebrae , not from the addition of more vertebrae. Each cervical vertebra is over 28 cm (11 in) long. They comprise 52–54 per cent of the length of the giraffe's vertebral column , compared with the 27–33 percent typical of similar large ungulates, including the giraffe's closest living relative, the okapi . This elongation largely takes place after birth, perhaps because giraffe mothers would have

712-492: A dry savannah emerged across eastern and northern Africa and western India. Some researchers have hypothesised that this new habitat, coupled with a different diet, including acacia species, may have exposed giraffe ancestors to toxins that caused higher mutation rates and a higher rate of evolution. The coat patterns of modern giraffes may also have coincided with these habitat changes. Asian giraffes are hypothesised to have had more okapi-like colourations. The giraffe genome

801-466: A food shortage. Another theory, the sexual selection hypothesis, proposes that long necks evolved as a secondary sexual characteristic , giving males an advantage in "necking" contests (see below) to establish dominance and obtain access to sexually receptive females. In support of this theory, some studies have stated that necks are longer and heavier for males than females of the same age, and that males do not employ other forms of combat. However,

890-582: A food trap, mechanical stimulation, and electrical impulses to eventually catch and consume its prey. Some carnivorous fungi catch nematodes using either active traps in the form of constricting rings, or passive traps with adhesive structures. Many species of protozoa ( eukaryotes ) and bacteria ( prokaryotes ) prey on other microorganisms; the feeding mode is evidently ancient, and evolved many times in both groups. Among freshwater and marine zooplankton , whether single-celled or multi-cellular, predatory grazing on phytoplankton and smaller zooplankton

979-464: A giraffe's skull. However, as males age, their skulls become heavier and more club-like, helping them become more dominant in combat. The occipital condyles at the bottom of the skull allow the animal to tip its head over 90 degrees and grab food on the branches directly above them with the tongue. With eyes located on the sides of the head, the giraffe has a broad visual field from its great height. Compared to other ungulates , giraffe vision

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1068-471: A huge gulp of water and filtering it through their feathery baleen plates. Pursuit predators may be social , like the lion and wolf that hunt in groups, or solitary. Once the predator has captured the prey, it has to handle it: very carefully if the prey is dangerous to eat, such as if it possesses sharp or poisonous spines, as in many prey fish. Some catfish such as the Ictaluridae have spines on

1157-725: A kill, and the coyote can be either solitary or social. Other solitary predators include the northern pike, wolf spiders and all the thousands of species of solitary wasps among arthropods, and many microorganisms and zooplankton . Under the pressure of natural selection , predators have evolved a variety of physical adaptations for detecting, catching, killing, and digesting prey. These include speed, agility, stealth, sharp senses, claws, teeth, filters, and suitable digestive systems. For detecting prey , predators have well-developed vision , smell , or hearing . Predators as diverse as owls and jumping spiders have forward-facing eyes, providing accurate binocular vision over

1246-539: A long distance, sometimes for hours at a time. The method is used by human hunter-gatherers and by canids such as African wild dogs and domestic hounds. The African wild dog is an extreme persistence predator, tiring out individual prey by following them for many miles at relatively low speed. A specialised form of pursuit predation is the lunge feeding of baleen whales . These very large marine predators feed on plankton , especially krill , diving and actively swimming into concentrations of plankton, and then taking

1335-588: A medium-sized, lightly built body. Giraffokeryx appeared 15–12 mya on the Indian subcontinent and resembled an okapi or a small giraffe, and had a longer neck and similar ossicones . Giraffokeryx may have shared a clade with more massively built giraffids like Sivatherium and Bramatherium . Giraffids like Palaeotragus , Shansitherium and Samotherium appeared 14 mya and lived throughout Africa and Eurasia. These animals had broader skulls with reduced frontal cavities. Paleotragus resembled

1424-445: A patch and decide whether to spend time searching for prey in it. This may involve some knowledge of the preferences of the prey; for example, ladybirds can choose a patch of vegetation suitable for their aphid prey. To capture prey, predators have a spectrum of pursuit modes that range from overt chase ( pursuit predation ) to a sudden strike on nearby prey ( ambush predation ). Another strategy in between ambush and pursuit

1513-425: A powerful selective effect on prey, and the prey develop antipredator adaptations such as warning coloration , alarm calls and other signals , camouflage , mimicry of well-defended species, and defensive spines and chemicals. Sometimes predator and prey find themselves in an evolutionary arms race , a cycle of adaptations and counter-adaptations. Predation has been a major driver of evolution since at least

1602-785: A predator, while small prey might prove hard to find and in any case provide less of a reward. This has led to a correlation between the size of predators and their prey. Size may also act as a refuge for large prey. For example, adult elephants are relatively safe from predation by lions, but juveniles are vulnerable. Members of the cat family such as the snow leopard (treeless highlands), tiger (grassy plains, reed swamps), ocelot (forest), fishing cat (waterside thickets), and lion (open plains) are camouflaged with coloration and disruptive patterns suiting their habitats. In aggressive mimicry , certain predators, including insects and fishes, make use of coloration and behaviour to attract prey. Female Photuris fireflies , for example, copy

1691-421: A preferred target is scarce. When prey have a clumped (uneven) distribution, the optimal strategy for the predator is predicted to be more specialized as the prey are more conspicuous and can be found more quickly; this appears to be correct for predators of immobile prey, but is doubtful with mobile prey. In size-selective predation, predators select prey of a certain size. Large prey may prove troublesome for

1780-820: A relatively narrow field of view, whereas prey animals often have less acute all-round vision. Animals such as foxes can smell their prey even when it is concealed under 2 feet (60 cm) of snow or earth. Many predators have acute hearing, and some such as echolocating bats hunt exclusively by active or passive use of sound. Predators including big cats , birds of prey , and ants share powerful jaws, sharp teeth, or claws which they use to seize and kill their prey. Some predators such as snakes and fish-eating birds like herons and cormorants swallow their prey whole; some snakes can unhinge their jaws to allow them to swallow large prey, while fish-eating birds have long spear-like beaks that they use to stab and grip fast-moving and slippery prey. Fish and other predators have developed

1869-478: A significant amount of energy, to locate each food patch. For example, the black-browed albatross regularly makes foraging flights to a range of around 700 kilometres (430 miles), up to a maximum foraging range of 3,000 kilometres (1,860 miles) for breeding birds gathering food for their young. With static prey, some predators can learn suitable patch locations and return to them at intervals to feed. The optimal foraging strategy for search has been modelled using

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1958-406: A small animal, gulping the prey in an extremely rapid movement when it is within range. Many smaller predators such as the box jellyfish use venom to subdue their prey, and venom can also aid in digestion (as is the case for rattlesnakes and some spiders ). The marbled sea snake that has adapted to egg predation has atrophied venom glands, and the gene for its three finger toxin contains

2047-523: A smaller area. For example, when mixed flocks of birds forage, the birds in front flush out insects that are caught by the birds behind. Spinner dolphins form a circle around a school of fish and move inwards, concentrating the fish by a factor of 200. By hunting socially chimpanzees can catch colobus monkeys that would readily escape an individual hunter, while cooperating Harris hawks can trap rabbits. Predators of different species sometimes cooperate to catch prey. In coral reefs , when fish such as

2136-440: A variety of defences including the ability to hear the echolocation calls. Many pursuit predators that run on land, such as wolves, have evolved long limbs in response to the increased speed of their prey. Their adaptations have been characterized as an evolutionary arms race , an example of the coevolution of two species. In a gene centered view of evolution , the genes of predator and prey can be thought of as competing for

2225-477: Is ballistic interception , where a predator observes and predicts a prey's motion and then launches its attack accordingly. Ambush or sit-and-wait predators are carnivorous animals that capture prey by stealth or surprise. In animals, ambush predation is characterized by the predator's scanning the environment from a concealed position until a prey is spotted, and then rapidly executing a fixed surprise attack. Vertebrate ambush predators include frogs, fish such as

2314-659: Is a stub . You can help Misplaced Pages by expanding it . Giraffe See taxonomy The giraffe is a large African hoofed mammal belonging to the genus Giraffa . It is the tallest living terrestrial animal and the largest ruminant on Earth . Traditionally, giraffes have been thought of as one species , Giraffa camelopardalis , with nine subspecies . Most recently, researchers proposed dividing them into four extant species due to new research into their mitochondrial and nuclear DNA , and individual species can be distinguished by their fur coat patterns. Seven other extinct species of Giraffa are known from

2403-598: Is around 2.9 billion base pairs in length, compared to the 3.3 billion base pairs of the okapi. Of the proteins in giraffe and okapi genes, 19.4% are identical. The divergence of giraffe and okapi lineages dates to around 11.5 mya. A small group of regulatory genes in the giraffe appears responsible for the animal's height and associated circulatory adaptations. The International Union for Conservation of Nature (IUCN) currently recognises only one species of giraffe with nine subspecies. Carl Linnaeus originally classified living giraffes as one species in 1758. He gave it

2492-494: Is common, and found in many species of nanoflagellates , dinoflagellates , ciliates , rotifers , a diverse range of meroplankton animal larvae, and two groups of crustaceans, namely copepods and cladocerans . To feed, a predator must search for, pursue and kill its prey. These actions form a foraging cycle. The predator must decide where to look for prey based on its geographical distribution; and once it has located prey, it must assess whether to pursue it or to wait for

2581-404: Is correlated with calf survival. The skin under the blotches may regulate the animal's body temperature, being sites for complex blood vessel systems and large sweat glands . Spotless or solid-color giraffes are very rare, but have been observed. The fur may give the animal chemical defense, as its parasite repellents give it a characteristic scent. At least 11 main aromatic chemicals are in

2670-629: Is detected, the predator assesses whether to attack it. This may involve ambush or pursuit predation , sometimes after stalking the prey. If the attack is successful, the predator kills the prey, removes any inedible parts like the shell or spines, and eats it. Predators are adapted and often highly specialized for hunting, with acute senses such as vision , hearing , or smell . Many predatory animals , both vertebrate and invertebrate , have sharp claws or jaws to grip, kill, and cut up their prey. Other adaptations include stealth and aggressive mimicry that improve hunting efficiency. Predation has

2759-482: Is flexible and hairy to protect against sharp prickles. The upper jaw has a hard palate instead of front teeth. The molars and premolars are wide with low crowns on the surface. The giraffe has an extremely elongated neck, which can be up to 2.4 m (7 ft 10 in) in length. Along the neck is a mane made of short, erect hairs. The neck typically rests at an angle of 50–60 degrees, though juveniles are closer to 70 degrees. The long neck results from

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2848-477: Is leaves, fruits, and flowers of woody plants, primarily acacia species, which they browse at heights most other ground-based herbivores cannot reach. Lions , leopards , spotted hyenas , and African wild dogs may prey upon giraffes. Giraffes live in herds of related females and their offspring or bachelor herds of unrelated adult males but are gregarious and may gather in large groups. Males establish social hierarchies through "necking", combat bouts where

2937-439: Is more binocular and the eyes are larger with a greater retinal surface area. Giraffes may see in colour, and their senses of hearing and smell are sharp. The ears are movable. The nostrils are slit-shaped, possibly to withstand blowing sand. The giraffe's tongue is about 45 cm (18 in) long. It is black, perhaps to protect against sunburn, and can grasp foliage and delicately pick off leaves. The upper lip

3026-463: Is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the host ) and parasitoidism (which always does, eventually). It is distinct from scavenging on dead prey, though many predators also scavenge ; it overlaps with herbivory , as seed predators and destructive frugivores are predators. Predators may actively search for or pursue prey or wait for it, often concealed. When prey

3115-671: Is one of only two living genera of the family Giraffidae in the order Artiodactyla , the other being the okapi . They are ruminants of the clade Pecora , along with Antilocapridae ( pronghorns ), Cervidae (deer), Bovidae (cattle, antelope, goats and sheep) and Moschidae (musk deer). A 2019 genome study (cladogram below) finds that Giraffidae are a sister taxon to Antilocapridae, with an estimated split of over 20 million years ago. Tragulidae [REDACTED] Antilocapridae [REDACTED] Giraffidae [REDACTED] Cervidae [REDACTED] Bovidae [REDACTED] Moschidae [REDACTED] The family Giraffidae

3204-511: Is relatively short. The skin is mostly gray or tan, and can reach a thickness of 20 mm (0.79 in). The 80–100 cm (31–39 in) long tail ends in a long, dark tuft of hair and is used as a defense against insects. The coat has dark blotches or patches, which can be orange, chestnut , brown, or nearly black, surrounded by light hair, usually white or cream coloured. Male giraffes become darker as they grow old. The coat pattern has been claimed to serve as camouflage in

3293-514: Is restricted to mammals, birds, and insects but is found in almost all terrestrial ecosystems. Egg predation includes both specialist egg predators such as some colubrid snakes and generalists such as foxes and badgers that opportunistically take eggs when they find them. Some plants, like the pitcher plant , the Venus fly trap and the sundew , are carnivorous and consume insects . Methods of predation by plants varies greatly but often involves

3382-430: Is size. Prey that is too small may not be worth the trouble for the amount of energy it provides. Too large, and it may be too difficult to capture. For example, a mantid captures prey with its forelegs and they are optimized for grabbing prey of a certain size. Mantids are reluctant to attack prey that is far from that size. There is a positive correlation between the size of a predator and its prey. A predator may assess

3471-450: Is subsumed into G. camelopardalis camelopardalis . G. camelopardalis antiquorum (Kordofan giraffe) G. c. camelopardalis (Nubian giraffe) G. c. peralta (West African giraffe) (no subspecies) G. tippelskirchi tippelskirchi (Masai giraffe sensu stricto ) G. t. thornicrofti (Luangwa or Thornicroft's giraffe) G. giraffa angolensis (Angolan giraffe) G. g. giraffa (South African giraffe) The following table compares

3560-455: Is what is now referred to “Old Lengwe.” This did not contain the year-round water supply to the animals that the Shire had provided, 964 and 1970, four artificial waterholes were built throughout the area, and in 1970, Lengwe was given national park status. In 1975, the extension area was added, increasing the size to 887 square kilometers. Lengwe was expanded mainly to protect the catchment areas of

3649-451: The Cambrian period. At the most basic level, predators kill and eat other organisms. However, the concept of predation is broad, defined differently in different contexts, and includes a wide variety of feeding methods; moreover, some relationships that result in the prey's death are not necessarily called predation. A parasitoid , such as an ichneumon wasp , lays its eggs in or on its host;

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3738-531: The Pleistocene . Some biologists suggest the modern giraffes descended from G. jumae ; others find G. gracilis a more likely candidate. G. jumae was larger and more robust, while G. gracilis was smaller and more slender. The changes from extensive forests to more open habitats , which began 8 mya, are believed to be the main driver for the evolution of giraffes. During this time, tropical plants disappeared and were replaced by arid C4 plants , and

3827-598: The Yellow-spotted nicator . History In 1928, Lengwe was established as a Game Reserve to protect the large mammals found in the Lower Shire Valley, especially the nyala antelope and its habitat. This protection is important as the Lower Shire Valley is the farthest north where nyala can be found naturally. At that time, the protected area measured 520 square kilometers. Later, it was reduced to 120 square kilometers in order to provide land for agriculture. This 120-km

3916-465: The angel shark , the northern pike and the eastern frogfish . Among the many invertebrate ambush predators are trapdoor spiders and Australian Crab spiders on land and mantis shrimps in the sea. Ambush predators often construct a burrow in which to hide, improving concealment at the cost of reducing their field of vision. Some ambush predators also use lures to attract prey within striking range. The capturing movement has to be rapid to trap

4005-471: The binomial name Cervus camelopardalis . Mathurin Jacques Brisson coined the generic name Giraffa in 1762. During the 1900s, various taxonomies with two or three species were proposed. A 2007 study on the genetics of giraffes using mitochondrial DNA suggested at least six lineages could be recognised as species. A 2011 study using detailed analyses of the morphology of giraffes, and application of

4094-455: The common garter snake has developed a resistance to the toxin in the skin of the rough-skinned newt . Predators affect their ecosystems not only directly by eating their own prey, but by indirect means such as reducing predation by other species, or altering the foraging behaviour of a herbivore, as with the biodiversity effect of wolves on riverside vegetation or sea otters on kelp forests. This may explain population dynamics effects such as

4183-461: The fossil record. The giraffe's distinguishing characteristics are its extremely long neck and legs, horn-like ossicones , and spotted coat patterns. It is classified under the family Giraffidae , along with its closest extant relative, the okapi . Its scattered range extends from Chad in the north to South Africa in the south and from Niger in the west to Somalia in the east. Giraffes usually inhabit savannahs and woodlands . Their food source

4272-475: The grouper and coral trout spot prey that is inaccessible to them, they signal to giant moray eels , Napoleon wrasses or octopuses . These predators are able to access small crevices and flush out the prey. Killer whales have been known to help whalers hunt baleen whales . Social hunting allows predators to tackle a wider range of prey, but at the risk of competition for the captured food. Solitary predators have more chance of eating what they catch, at

4361-461: The marginal value theorem . Search patterns often appear random. One such is the Lévy walk , that tends to involve clusters of short steps with occasional long steps. It is a good fit to the behaviour of a wide variety of organisms including bacteria, honeybees, sharks and human hunter-gatherers. Having found prey, a predator must decide whether to pursue it or keep searching. The decision depends on

4450-454: The phylogenetic species concept , described eight species of living giraffes. A 2016 study also concluded that living giraffes consist of multiple species. The researchers suggested the existence of four species, which have not exchanged genetic information between each other for 1 to 2 million years. A 2020 study showed that depending on the method chosen, different taxonomic hypotheses recognizing from two to six species can be considered for

4539-542: The 1590s. The modern English form developed around 1600 from the French girafe . "Camelopard" / k ə ˈ m ɛ l ə ˌ p ɑːr d / is an archaic English name for the giraffe; it derives from the Ancient Greek καμηλοπάρδαλις ( kamēlopárdalis ), from κάμηλος ( kámēlos ), " camel ", and πάρδαλις ( párdalis ), " leopard ", referring to its camel-like shape and leopard-like colouration. The giraffe

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4628-454: The Shire, Mwanza, and Nkombedzi wa Fodya rivers. The extension also has the effect of protecting beautiful rock outcrops in the area and increasing the dispersal area for wildlife. The Park is managed by the Department of National Parks and Wildlife (DNPW. This Malawi location article is a stub . You can help Misplaced Pages by expanding it . This Africa protected areas related article

4717-709: The ability of predatory bacteria to digest the complex peptidoglycan polymer from the cell walls of the bacteria that they prey upon. Carnivorous vertebrates of all five major classes (fishes, amphibians, reptiles, birds, and mammals) have lower relative rates of sugar to amino acid transport than either herbivores or omnivores, presumably because they acquire plenty of amino acids from the animal proteins in their diet. To counter predation, prey have evolved defences for use at each stage of an attack. They can try to avoid detection, such as by using camouflage and mimicry . They can detect predators and warn others of their presence. If detected, they can try to avoid being

4806-728: The ability to crush or open the armoured shells of molluscs. Many predators are powerfully built and can catch and kill animals larger than themselves; this applies as much to small predators such as ants and shrews as to big and visibly muscular carnivores like the cougar and lion . Predators are often highly specialized in their diet and hunting behaviour; for example, the Eurasian lynx only hunts small ungulates . Others such as leopards are more opportunistic generalists, preying on at least 100 species. The specialists may be highly adapted to capturing their preferred prey, whereas generalists may be better able to switch to other prey when

4895-584: The assault. When animals eat seeds ( seed predation or granivory ) or eggs ( egg predation ), they are consuming entire living organisms, which by definition makes them predators. Scavengers , organisms that only eat organisms found already dead, are not predators, but many predators such as the jackal and the hyena scavenge when the opportunity arises. Among invertebrates, social wasps such as yellowjackets are both hunters and scavengers of other insects. While examples of predators among mammals and birds are well known, predators can be found in

4984-684: The back (dorsal) and belly (pectoral) which lock in the erect position; as the catfish thrashes about when captured, these could pierce the predator's mouth, possibly fatally. Some fish-eating birds like the osprey avoid the danger of spines by tearing up their prey before eating it. In social predation, a group of predators cooperates to kill prey. This makes it possible to kill creatures larger than those they could overpower singly; for example, hyenas , and wolves collaborate to catch and kill herbivores as large as buffalo, and lions even hunt elephants. It can also make prey more readily available through strategies like flushing of prey and herding it into

5073-432: The canopy. However, scientists disagree about just how much time giraffes spend feeding at levels beyond the reach of other browsers, and a 2010 study found that adult giraffes with longer necks actually suffered higher mortality rates under drought conditions than their shorter-necked counterparts. This study suggests that maintaining a longer neck requires more nutrients, which puts longer-necked giraffes at risk during

5162-416: The costs and benefits involved. A bird foraging for insects spends a lot of time searching but capturing and eating them is quick and easy, so the efficient strategy for the bird is to eat every palatable insect it finds. By contrast, a predator such as a lion or falcon finds its prey easily but capturing it requires a lot of effort. In that case, the predator is more selective. One of the factors to consider

5251-496: The different hypotheses for giraffe species. The description column shows the traditional nine subspecies in the one-species hypothesis. Also known as Baringo giraffe or Ugandan giraffe Also known as Niger giraffe or Nigerian giraffe Also known as Somali giraffe Also known as Namibian giraffe Also known as Cape giraffe Also known as Kilimanjaro giraffe Also known as Luangwa giraffe or Rhodesian giraffe The first extinct species to be described

5340-993: The eggs hatch into larvae, which eat the host, and it inevitably dies. Zoologists generally call this a form of parasitism , though conventionally parasites are thought not to kill their hosts. A predator can be defined to differ from a parasitoid in that it has many prey, captured over its lifetime, where a parasitoid's larva has just one, or at least has its food supply provisioned for it on just one occasion. There are other difficult and borderline cases. Micropredators are small animals that, like predators, feed entirely on other organisms; they include fleas and mosquitoes that consume blood from living animals, and aphids that consume sap from living plants. However, since they typically do not kill their hosts, they are now often thought of as parasites. Animals that graze on phytoplankton or mats of microbes are predators, as they consume and kill their food organisms, while herbivores that browse leaves are not, as their food plants usually survive

5429-472: The elongation of the neck, as it enabled giraffes to reach food that competitors could not. This advantage is real, as giraffes can and do feed up to 4.5 m (15 ft) high, while even quite large competitors, such as kudu, can feed up to only about 2 m (6 ft 7 in) high. There is also research suggesting that browsing competition is intense at lower levels, and giraffes feed more efficiently (gaining more leaf biomass with each mouthful) high in

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5518-404: The evolution of mimicry. Avoidance is not necessarily an evolutionary response as it is generally learned from bad experiences with prey. However, when the prey is capable of killing the predator (as can a coral snake with its venom), there is no opportunity for learning and avoidance must be inherited. Predators can also respond to dangerous prey with counter-adaptations. In western North America,

5607-420: The existence of four distinct species and seven subspecies. A 2024 study found a higher amount of ancient gene flow than expected between populations. The cladogram below shows the phylogenetic relationship between the four proposed species and seven subspecies based on a 2021 genome analysis. The eight lineages correspond to eight traditional subspecies in the one-species hypothesis. The Rothschild giraffe

5696-538: The first and second thoracic vertebrae (T1 and T2), unlike in most other ruminants, where the articulation is between the seventh cervical vertebra (C7) and T1. This allows C7 to contribute directly to increased neck length and has given rise to the suggestion that T1 is actually C8, and that giraffes have added an extra cervical vertebra. However, this proposition is not generally accepted, as T1 has other morphological features, such as an articulating rib , deemed diagnostic of thoracic vertebrae, and because exceptions to

5785-425: The fur, although indole and 3-methylindole are responsible for most of the smell. Because males have a stronger odour than females, it may also have a sexual function . Both sexes have prominent horn-like structures called ossicones , which can reach 13.5 cm (5.3 in). They are formed from ossified cartilage , covered in skin, and fused to the skull at the parietal bones . Being vascularised ,

5874-482: The genus Giraffa . That study also found that multi-species coalescent methods can lead to taxonomic over-splitting, as those methods delimit geographic structures rather than species. The three-species hypothesis, which recognises G . camelopardalis , G . giraffa , and G . tippelskirchi , is highly supported by phylogenetic analyses and also corroborated by most population genetic and multi-species coalescent analyses. A 2021 whole genome sequencing study suggests

5963-431: The larvae of coccinellid beetles (ladybirds) , alternate between actively searching and scanning the environment. Prey distributions are often clumped, and predators respond by looking for patches where prey is dense and then searching within patches. Where food is found in patches, such as rare shoals of fish in a nearly empty ocean, the search stage requires the predator to travel for a substantial time, and to expend

6052-486: The light and shade patterns of savannah woodlands. When standing among trees and bushes, they are hard to see at even a few metres distance. However, adult giraffes move about to gain the best view of an approaching predator, relying on their size and ability to defend themselves rather than on camouflage, which may be more important for calves. Each giraffe has a unique coat pattern. Calves inherit some coat pattern traits from their mothers, and variation in some spot traits

6141-429: The light signals of other species, thereby attracting male fireflies, which they capture and eat. Flower mantises are ambush predators; camouflaged as flowers, such as orchids , they attract prey and seize it when it is close enough. Frogfishes are extremely well camouflaged, and actively lure their prey to approach using an esca , a bait on the end of a rod-like appendage on the head, which they wave gently to mimic

6230-470: The mammalian limit of seven cervical vertebrae are generally characterised by increased neurological anomalies and maladies. There are several hypotheses regarding the evolutionary origin and maintenance of elongation in giraffe necks. Charles Darwin originally suggested the "competing browsers hypothesis", which has been challenged only recently. It suggests that competitive pressure from smaller browsers, like kudu , steenbok and impala , encouraged

6319-680: The neck is used as a weapon. Dominant males gain mating access to females, which bear sole responsibility for rearing the young. The giraffe has intrigued various ancient and modern cultures for its peculiar appearance and has often been featured in paintings, books, and cartoons. It is classified by the International Union for Conservation of Nature (IUCN) as vulnerable to extinction. It has been extirpated from many parts of its former range. Giraffes are still found in many national parks and game reserves , but estimates as of 2016 indicate there are approximately 97,500 members of Giraffa in

6408-452: The okapi and may have been its ancestor. Others find that the okapi lineage diverged earlier, before Giraffokeryx . Samotherium was a particularly important transitional fossil in the giraffe lineage, as the length and structure of its cervical vertebrae were between those of a modern giraffe and an okapi, and its neck posture was likely similar to the former's. Bohlinia , which first appeared in southeastern Europe and lived 9–7 mya,

6497-484: The ossicones may have a role in thermoregulation , and are used in combat between males. Appearance is a reliable guide to the sex or age of a giraffe: the ossicones of females and young are thin and display tufts of hair on top, whereas those of adult males tend to be bald and knobbed on top. A lump, which is more prominent in males, emerges in the middle of the skull. Males develop calcium deposits that form bumps on their skulls as they age. Multiple sinuses lighten

6586-434: The other hand, the fitness cost of a given lost dinner is unpredictable, as the predator may quickly find better prey. In addition, most predators are generalists, which reduces the impact of a given prey adaption on a predator. Since specialization is caused by predator-prey coevolution, the rarity of specialists may imply that predator-prey arms races are rare. It is difficult to determine whether given adaptations are truly

6675-538: The prey an opportunity to escape. Some frogs wait until snakes have begun their strike before jumping, reducing the time available to the snake to recalibrate its attack, and maximising the angular adjustment that the snake would need to make to intercept the frog in real time. Ballistic predators include insects such as dragonflies, and vertebrates such as archerfish (attacking with a jet of water), chameleons (attacking with their tongues), and some colubrid snakes . In pursuit predation, predators chase fleeing prey. If

6764-549: The prey are dense and mobile, and the predator has low energy requirements. Wide foraging expends more energy, and is used when prey is sedentary or sparsely distributed. There is a continuum of search modes with intervals between periods of movement ranging from seconds to months. Sharks, sunfish , Insectivorous birds and shrews are almost always moving while web-building spiders, aquatic invertebrates, praying mantises and kestrels rarely move. In between, plovers and other shorebirds , freshwater fish including crappies , and

6853-413: The prey flees in a straight line, capture depends only on the predator's being faster than the prey. If the prey manoeuvres by turning as it flees, the predator must react in real time to calculate and follow a new intercept path, such as by parallel navigation , as it closes on the prey. Many pursuit predators use camouflage to approach the prey as close as possible unobserved ( stalking ) before starting

6942-428: The prey's body. However, the "life-dinner" principle of Dawkins and Krebs predicts that this arms race is asymmetric: if a predator fails to catch its prey, it loses its dinner, while if it succeeds, the prey loses its life. The metaphor of an arms race implies ever-escalating advances in attack and defence. However, these adaptations come with a cost; for instance, longer legs have an increased risk of breaking, while

7031-433: The prey, given that the attack is not modifiable once launched. Ballistic interception is the strategy where a predator observes the movement of a prey, predicts its motion, works out an interception path, and then attacks the prey on that path. This differs from ambush predation in that the predator adjusts its attack according to how the prey is moving. Ballistic interception involves a brief period for planning, giving

7120-504: The price of increased expenditure of energy to catch it, and increased risk that the prey will escape. Ambush predators are often solitary to reduce the risk of becoming prey themselves. Of 245 terrestrial members of the Carnivora (the group that includes the cats, dogs, and bears), 177 are solitary; and 35 of the 37 wild cats are solitary, including the cougar and cheetah. However, the solitary cougar does allow other cougars to share in

7209-414: The pursuit. Pursuit predators include terrestrial mammals such as humans, African wild dogs, spotted hyenas and wolves; marine predators such as dolphins, orcas and many predatory fishes, such as tuna; predatory birds (raptors) such as falcons; and insects such as dragonflies . An extreme form of pursuit is endurance or persistence hunting , in which the predator tires out the prey by following it over

7298-470: The result of coevolution, where a prey adaptation gives rise to a predator adaptation that is countered by further adaptation in the prey. An alternative explanation is escalation , where predators are adapting to competitors, their own predators or dangerous prey. Apparent adaptations to predation may also have arisen for other reasons and then been co-opted for attack or defence. In some of the insects preyed on by bats, hearing evolved before bats appeared and

7387-417: The specialized tongue of the chameleon, with its ability to act like a projectile, is useless for lapping water, so the chameleon must drink dew off vegetation. The "life-dinner" principle has been criticized on multiple grounds. The extent of the asymmetry in natural selection depends in part on the heritability of the adaptive traits. Also, if a predator loses enough dinners, it too will lose its life. On

7476-684: The target of an attack, for example, by signalling that they are toxic or unpalatable , by signalling that a chase would be unprofitable, or by forming groups. If they become a target, they can try to fend off the attack with defences such as armour, quills , unpalatability, or mobbing; and they can often escape an attack in progress by startling the predator, playing dead , shedding body parts such as tails, or simply fleeing. Predators and prey are natural enemies, and many of their adaptations seem designed to counter each other. For example, bats have sophisticated echolocation systems to detect insects and other prey, and insects have developed

7565-668: The wild. More than 1,600 were kept in zoos in 2010. The name "giraffe" has its earliest known origins in the Arabic word zarāfah ( زرافة ), ultimately from Persian زُرنَاپَا ( zurnāpā ), a compound of زُرنَا ( zurnā , "flute, zurna") and پَا ( pā , "leg"). In early Modern English the spellings jarraf and ziraph were used, probably directly from the Arabic, and in Middle English jarraf and ziraph , gerfauntz . The Italian form giraffa arose in

7654-520: Was Giraffa sivalensis Falconer and Cautley 1843, a reevaluation of a vertebra that was initially described as a fossil of the living giraffe. While taxonomic opinion may be lacking on some names, the extinct species that have been published include: Fully grown giraffes stand 4.3–5.7 m (14–19 ft) tall, with males taller than females. The average weight is 1,192 kg (2,628 lb) for an adult male and 828 kg (1,825 lb) for an adult female. Despite its long neck and legs, its body

7743-668: Was likely a direct ancestor of the giraffe. Bohlinia closely resembled modern giraffes, having a long neck and legs and similar ossicones and dentition. Bohlinia colonised China and northern India and produced the Giraffa , which, around 7  million years ago , reached Africa. Climate changes led to the extinction of the Asian giraffes, while the African giraffes survived and radiated into new species. Living giraffes appear to have arisen around 1  million years ago in eastern Africa during

7832-627: Was once much more extensive, with over 10 fossil genera described. The elongation of the neck appears to have started early in the giraffe lineage . Comparisons between giraffes and their ancient relatives suggest vertebrae close to the skull lengthened earlier, followed by lengthening of vertebrae further down. One early giraffid ancestor was Canthumeryx , which has been dated variously to have lived 25 to 20 million years ago , 17–15 mya or 18–14.3 mya and whose deposits have been found in Libya. This animal resembled an antelope and had

7921-401: Was used to hear signals used for territorial defence and mating. Their hearing evolved in response to bat predation, but the only clear example of reciprocal adaptation in bats is stealth echolocation. A more symmetric arms race may occur when the prey are dangerous, having spines, quills, toxins or venom that can harm the predator. The predator can respond with avoidance, which in turn drives

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