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68-566: The Laurales are an order of flowering plants . They are magnoliids , related to the Magnoliales . The order includes about 2500-2800 species from 85 to 90 genera, which comprise seven families of trees and shrubs . Most of the species are tropical and subtropical, though a few genera reach the temperate zone. The best known species in this order are those of the Lauraceae (for example bay laurel , cinnamon , avocado , and Sassafras ), and

136-1845: A molecular phylogeny of plants placed the flowering plants in their evolutionary context: Bryophytes [REDACTED] Lycophytes [REDACTED] Ferns [REDACTED] [REDACTED] [REDACTED] The main groups of living angiosperms are: Amborellales [REDACTED] 1 sp. New Caledonia shrub Nymphaeales [REDACTED] c. 80 spp. water lilies & allies Austrobaileyales [REDACTED] c. 100 spp. woody plants Magnoliids [REDACTED] c. 10,000 spp. 3-part flowers, 1-pore pollen, usu. branch-veined leaves Chloranthales [REDACTED] 77 spp. Woody, apetalous Monocots [REDACTED] c. 70,000 spp. 3-part flowers, 1 cotyledon , 1-pore pollen, usu. parallel-veined leaves   Ceratophyllales [REDACTED] c. 6 spp. aquatic plants Eudicots [REDACTED] c. 175,000 spp. 4- or 5-part flowers, 3-pore pollen, usu. branch-veined leaves Amborellales Melikyan, Bobrov & Zaytzeva 1999 Nymphaeales Salisbury ex von Berchtold & Presl 1820 Austrobaileyales Takhtajan ex Reveal 1992 Chloranthales Mart. 1835 Canellales Cronquist 1957 Piperales von Berchtold & Presl 1820 Magnoliales de Jussieu ex von Berchtold & Presl 1820 Laurales de Jussieu ex von Berchtold & Presl 1820 Acorales Link 1835 Alismatales Brown ex von Berchtold & Presl 1820 Petrosaviales Takhtajan 1997 Dioscoreales Brown 1835 Pandanales Brown ex von Berchtold & Presl 1820 Liliales Perleb 1826 Asparagales Link 1829 Arecales Bromhead 1840 Poales Small 1903 Zingiberales Grisebach 1854 Commelinales de Mirbel ex von Berchtold & Presl 1820 Xylem The most distinctive xylem cells are

204-442: A woody stem ), grasses and grass-like plants, a vast majority of broad-leaved trees , shrubs and vines , and most aquatic plants . Angiosperms are distinguished from the other major seed plant clade, the gymnosperms , by having flowers , xylem consisting of vessel elements instead of tracheids , endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from

272-488: A fabric with small spaces. In small passages, such as that between the plant cell walls (or in tracheids), a column of water behaves like rubber – when molecules evaporate from one end, they pull the molecules behind them along the channels. Therefore, transpiration alone provided the driving force for water transport in early plants. However, without dedicated transport vessels, the cohesion-tension mechanism cannot transport water more than about 2 cm, severely limiting

340-551: A film of surface moisture, enabling them to grow to much greater size. As a result of their independence from their surroundings, they lost their ability to survive desiccation – a costly trait to retain. During the Devonian, maximum xylem diameter increased with time, with the minimum diameter remaining pretty constant. By the middle Devonian, the tracheid diameter of some plant lineages ( Zosterophyllophytes ) had plateaued. Wider tracheids allow water to be transported faster, but

408-423: A great deal of resistance on flow; vessel members have perforated end walls, and are arranged in series to operate as if they were one continuous vessel. The function of end walls, which were the default state in the Devonian, was probably to avoid embolisms . An embolism is where an air bubble is created in a tracheid. This may happen as a result of freezing, or by gases dissolving out of solution. Once an embolism

476-400: A hundred meters from ground level to a tree 's highest branches. Transpirational pull requires that the vessels transporting the water be very small in diameter; otherwise, cavitation would break the water column. And as water evaporates from leaves, more is drawn up through the plant to replace it. When the water pressure within the xylem reaches extreme levels due to low water input from

544-455: A million times more conductive than the inter-cell method, giving the potential for transport over longer distances, and higher CO 2 diffusion rates. The earliest macrofossils to bear water-transport tubes are Silurian plants placed in the genus Cooksonia . The early Devonian pretracheophytes Aglaophyton and Horneophyton have structures very similar to the hydroids of modern mosses. Plants continued to innovate new ways of reducing

612-426: A plant's vascular system based on the classical research of Dixon-Joly (1894), Eugen Askenasy (1845–1903) (1895), and Dixon (1914,1924). Water is a polar molecule . When two water molecules approach one another, the slightly negatively charged oxygen atom of one forms a hydrogen bond with a slightly positively charged hydrogen atom in the other. This attractive force, along with other intermolecular forces ,

680-422: A range of mechanisms to contain the damage. Small pits link adjacent conduits to allow fluid to flow between them, but not air – although these pits, which prevent the spread of embolism, are also a major cause of them. These pitted surfaces further reduce the flow of water through the xylem by as much as 30%. The diversification of xylem strand shapes with tracheid network topologies increasingly resistant to

748-537: A robust internal structure that held long narrow channels for transporting water from the soil to all the different parts of the above-soil plant, especially to the parts where photosynthesis occurred. During the Silurian, CO 2 was readily available, so little water needed expending to acquire it. By the end of the Carboniferous, when CO 2 levels had lowered to something approaching today's, around 17 times more water

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816-498: A slightly different set of families (current placement, where different, in brackets): The extinct family Araripiaceae (containing only the genus Araripia ) is known from the Aptian of Brazil. Flowering plant Basal angiosperms Core angiosperms Flowering plants are plants that bear flowers and fruits , and form the clade Angiospermae ( / ˌ æ n dʒ i ə ˈ s p ər m iː / ). The term 'angiosperm'

884-446: A stem or root is elongating. Later, 'metaxylem' develops in the strands of xylem. Metaxylem vessels and cells are usually larger; the cells have thickenings typically either in the form of ladderlike transverse bars (scalariform) or continuous sheets except for holes or pits (pitted). Functionally, metaxylem completes its development after elongation ceases when the cells no longer need to grow in size. There are four primary patterns to

952-610: A total of 64 angiosperm orders and 416 families. The diversity of flowering plants is not evenly distributed. Nearly all species belong to the eudicot (75%), monocot (23%), and magnoliid (2%) clades. The remaining five clades contain a little over 250 species in total; i.e. less than 0.1% of flowering plant diversity, divided among nine families. The 25 most species-rich of 443 families, containing over 166,000 species between them in their APG circumscriptions, are: The botanical term "angiosperm", from Greek words angeíon ( ἀγγεῖον 'bottle, vessel') and spérma ( σπέρμα 'seed'),

1020-458: A vascular bundle will contain primary xylem only. The branching pattern exhibited by xylem follows Murray's law . Primary xylem is formed during primary growth from procambium . It includes protoxylem and metaxylem. Metaxylem develops after the protoxylem but before secondary xylem. Metaxylem has wider vessels and tracheids than protoxylem. Secondary xylem is formed during secondary growth from vascular cambium . Although secondary xylem

1088-415: A young vascular plant grows, one or more strands of primary xylem form in its stems and roots. The first xylem to develop is called 'protoxylem'. In appearance, protoxylem is usually distinguished by narrower vessels formed of smaller cells. Some of these cells have walls that contain thickenings in the form of rings or helices. Functionally, protoxylem can extend: the cells can grow in size and develop while

1156-858: Is starting to impact plants and is likely to cause many species to become extinct by 2100. Angiosperms are terrestrial vascular plants; like the gymnosperms, they have roots , stems , leaves , and seeds . They differ from other seed plants in several ways. The largest angiosperms are Eucalyptus gum trees of Australia, and Shorea faguetiana , dipterocarp rainforest trees of Southeast Asia, both of which can reach almost 100 metres (330 ft) in height. The smallest are Wolffia duckweeds which float on freshwater, each plant less than 2 millimetres (0.08 in) across. Considering their method of obtaining energy, some 99% of flowering plants are photosynthetic autotrophs , deriving their energy from sunlight and using it to create molecules such as sugars . The remainder are parasitic , whether on fungi like

1224-437: Is a lot lighter, thus cheaper to make, as vessels need to be much more reinforced to avoid cavitation. Xylem development can be described by four terms: centrarch, exarch, endarch and mesarch . As it develops in young plants, its nature changes from protoxylem to metaxylem (i.e. from first xylem to after xylem ). The patterns in which protoxylem and metaxylem are arranged are essential in studying plant morphology. As

1292-493: Is also found in members of the gymnosperm groups Gnetophyta and Ginkgophyta and to a lesser extent in members of the Cycadophyta , the two main groups in which secondary xylem can be found are: The xylem, vessels and tracheids of the roots, stems and leaves are interconnected to form a continuous system of water-conducting channels reaching all parts of the plants. The system transports water and soluble mineral nutrients from

1360-478: Is derived from the Greek words ἀγγεῖον / angeion ('container, vessel') and σπέρμα / sperma ('seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta . Angiosperms are by far the most diverse group of land plants with 64 orders , 416 families , approximately 13,000 known genera and 300,000 known species . They include all forbs (flowering plants without

1428-474: Is formed, it usually cannot be removed (but see later); the affected cell cannot pull water up, and is rendered useless. End walls excluded, the tracheids of prevascular plants were able to operate under the same hydraulic conductivity as those of the first vascular plant, Cooksonia . The size of tracheids is limited as they comprise a single cell; this limits their length, which in turn limits their maximum useful diameter to 80 μm. Conductivity grows with

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1496-597: Is lost much faster than CO 2 is absorbed, so plants need to replace it, and have developed systems to transport water from the moist soil to the site of photosynthesis. Early plants sucked water between the walls of their cells, then evolved the ability to control water loss (and CO 2 acquisition) through the use of stomata. Specialized water transport tissues soon evolved in the form of hydroids, tracheids, then secondary xylem, followed by an endodermis and ultimately vessels. The high CO 2 levels of Silurian-Devonian times, when plants were first colonizing land, meant that

1564-435: Is one of the principal factors responsible for the occurrence of surface tension in liquid water. It also allows plants to draw water from the root through the xylem to the leaf. Water is constantly lost through transpiration from the leaf. When one water molecule is lost another is pulled along by the processes of cohesion and tension. Transpiration pull, utilizing capillary action and the inherent surface tension of water,

1632-497: Is the primary mechanism of water movement in plants. However, it is not the only mechanism involved. Any use of water in leaves forces water to move into them. Transpiration in leaves creates tension (differential pressure) in the cell walls of mesophyll cells. Because of this tension, water is being pulled up from the roots into the leaves, helped by cohesion (the pull between individual water molecules, due to hydrogen bonds) and adhesion (the stickiness between water molecules and

1700-428: Is well worth plants' while to avoid cavitation occurring. For this reason, pits in tracheid walls have very small diameters, to prevent air entering and allowing bubbles to nucleate. Freeze-thaw cycles are a major cause of cavitation. Damage to a tracheid's wall almost inevitably leads to air leaking in and cavitation, hence the importance of many tracheids working in parallel. Once cavitation has occurred, plants have

1768-485: The Alismatales grow in marine environments, spreading with rhizomes that grow through the mud in sheltered coastal waters. Some specialised angiosperms are able to flourish in extremely acid or alkaline habitats. The sundews , many of which live in nutrient-poor acid bogs , are carnivorous plants , able to derive nutrients such as nitrate from the bodies of trapped insects. Other flowers such as Gentiana verna ,

1836-705: The Silurian (more than 400 million years ago), and trace fossils resembling individual xylem cells may be found in earlier Ordovician rocks. The earliest true and recognizable xylem consists of tracheids with a helical-annular reinforcing layer added to the cell wall . This is the only type of xylem found in the earliest vascular plants, and this type of cell continues to be found in the protoxylem (first-formed xylem) of all living groups of vascular plants. Several groups of plants later developed pitted tracheid cells independently through convergent evolution . In living plants, pitted tracheids do not appear in development until

1904-603: The hydrophilic cell walls of plants). This mechanism of water flow works because of water potential (water flows from high to low potential), and the rules of simple diffusion . Over the past century, there has been a great deal of research regarding the mechanism of xylem sap transport; today, most plant scientists continue to agree that the cohesion-tension theory best explains this process, but multiforce theories that hypothesize several alternative mechanisms have been suggested, including longitudinal cellular and xylem osmotic pressure gradients , axial potential gradients in

1972-460: The orchids for part or all of their life-cycle, or on other plants , either wholly like the broomrapes, Orobanche , or partially like the witchweeds, Striga . In terms of their environment, flowering plants are cosmopolitan, occupying a wide range of habitats on land, in fresh water and in the sea. On land, they are the dominant plant group in every habitat except for frigid moss-lichen tundra and coniferous forest . The seagrasses in

2040-883: The "Big Five" extinction events in Earth's history, only the Cretaceous–Paleogene extinction event had occurred while angiosperms dominated plant life on the planet. Today, the Holocene extinction affects all kingdoms of complex life on Earth, and conservation measures are necessary to protect plants in their habitats in the wild ( in situ ), or failing that, ex situ in seed banks or artificial habitats like botanic gardens . Otherwise, around 40% of plant species may become extinct due to human actions such as habitat destruction , introduction of invasive species , unsustainable logging , land clearing and overharvesting of medicinal or ornamental plants . Further, climate change

2108-402: The "next generation" of transport cell design, have a more rigid structure than hydroids, allowing them to cope with higher levels of water pressure. Tracheids may have a single evolutionary origin, possibly within the hornworts, uniting all tracheophytes (but they may have evolved more than once). Water transport requires regulation, and dynamic control is provided by stomata . By adjusting

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2176-517: The Laurales as a natural group was H. Hallier in 1905. He viewed them as being derived from the Magnoliales . During some or all of the 20th century, the Laurales generally included Amborella and the plants now classified in Austrobaileyales and Chloranthaceae . They were not removed until the advent of molecular data in the late 20th century; their previous inclusion made it harder to determine

2244-466: The amount of gas exchange, they can restrict the amount of water lost through transpiration. This is an important role where water supply is not constant, and indeed stomata appear to have evolved before tracheids, being present in the non-vascular hornworts. An endodermis probably evolved during the Silu-Devonian, but the first fossil evidence for such a structure is Carboniferous. This structure in

2312-497: The arrangement of protoxylem and metaxylem in stems and roots. The other three terms are used where there is more than one strand of primary xylem. In his book De plantis libri XVI (On Plants, in 16 books) (1583), the Italian physician and botanist Andrea Cesalpino proposed that plants draw water from soil not by magnetism ( ut magnes ferrum trahit , as magnetic iron attracts) nor by suction ( vacuum ), but by absorption, as occurs in

2380-458: The case of linen, sponges, or powders. The Italian biologist Marcello Malpighi was the first person to describe and illustrate xylem vessels, which he did in his book Anatome plantarum ... (1675). Although Malpighi believed that xylem contained only air, the British physician and botanist Nehemiah Grew , who was Malpighi's contemporary, believed that sap ascended both through the bark and through

2448-547: The common ancestor of all living gymnosperms before the end of the Carboniferous , over 300 million years ago. In the Cretaceous , angiosperms diversified explosively , becoming the dominant group of plants across the planet. Agriculture is almost entirely dependent on angiosperms, and a small number of flowering plant families supply nearly all plant-based food and livestock feed. Rice , maize and wheat provide half of

2516-405: The disc is sucked into the pore on that side, and blocks further flow. Other plants simply tolerate cavitation. For instance, oaks grow a ring of wide vessels at the start of each spring, none of which survive the winter frosts. Maples use root pressure each spring to force sap upwards from the roots, squeezing out any air bubbles. Growing to height also employed another trait of tracheids –

2584-635: The flowering plants as an unranked clade without a formal Latin name (angiosperms). A formal classification was published alongside the 2009 revision in which the flowering plants rank as the subclass Magnoliidae. From 1998, the Angiosperm Phylogeny Group (APG) has reclassified the angiosperms, with updates in the APG II system in 2003, the APG III system in 2009, and the APG IV system in 2016. In 2019,

2652-435: The fourth power of diameter, so increased diameter has huge rewards; vessel elements , consisting of a number of cells, joined at their ends, overcame this limit and allowed larger tubes to form, reaching diameters of up to 500 μm, and lengths of up to 10 m. Vessels first evolved during the dry, low CO 2 periods of the late Permian, in the horsetails, ferns and Selaginellales independently, and later appeared in

2720-470: The height of a plant increases and upwards transport of water by xylem is considered to limit the maximum height of trees. Three phenomena cause xylem sap to flow: The primary force that creates the capillary action movement of water upwards in plants is the adhesion between the water and the surface of the xylem conduits. Capillary action provides the force that establishes an equilibrium configuration, balancing gravity. When transpiration removes water at

2788-413: The key innovations that led to the success of the angiosperms . However, the occurrence of vessel elements is not restricted to angiosperms, and they are absent in some archaic or "basal" lineages of the angiosperms: (e.g., Amborellaceae , Tetracentraceae , Trochodendraceae , and Winteraceae ), and their secondary xylem is described by Arthur Cronquist as "primitively vesselless". Cronquist considered

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2856-405: The likelihood of cavitation. Cavitation occurs when a bubble of air forms within a vessel, breaking the bonds between chains of water molecules and preventing them from pulling more water up with their cohesive tension. A tracheid, once cavitated, cannot have its embolism removed and return to service (except in a few advanced angiosperms which have developed a mechanism of doing so). Therefore, it

2924-444: The long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called vessels . Xylem also contains two other type of cells: parenchyma and fibers . Xylem can be found: In transitional stages of plants with secondary growth , the first two categories are not mutually exclusive, although usually

2992-518: The manner of vines or lianas . The number of species of flowering plants is estimated to be in the range of 250,000 to 400,000. This compares to around 12,000 species of moss and 11,000 species of pteridophytes . The APG system seeks to determine the number of families , mostly by molecular phylogenetics . In the 2009 APG III there were 415 families. The 2016 APG IV added five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, for

3060-408: The maturation of the metaxylem (following the protoxylem ). In most plants, pitted tracheids function as the primary transport cells. The other type of vascular element, found in angiosperms, is the vessel element . Vessel elements are joined end to end to form vessels in which water flows unimpeded, as in a pipe. The presence of xylem vessels (also called trachea ) is considered to be one of

3128-403: The mid Cretaceous in angiosperms and gnetophytes. Vessels allow the same cross-sectional area of wood to transport around a hundred times more water than tracheids! This allowed plants to fill more of their stems with structural fibers, and also opened a new niche to vines , which could transport water without being as thick as the tree they grew on. Despite these advantages, tracheid-based wood

3196-433: The need for water was relatively low. As CO 2 was withdrawn from the atmosphere by plants, more water was lost in its capture, and more elegant transport mechanisms evolved. As water transport mechanisms, and waterproof cuticles, evolved, plants could survive without being continually covered by a film of water. This transition from poikilohydry to homoiohydry opened up new potential for colonization. Plants then needed

3264-511: The ornamental shrub Calycanthus of the Calycanthaceae. The earliest lauraceous fossils are from the early Cretaceous . It is possible that the ancient origin of this order is one of the reasons for its highly diverged morphology. Presently no single morphological property is known, which would unify all the members of Laurales. The presently accepted classification is based on molecular and genetic analysis. The first botanist to think of

3332-480: The overall transport rate depends also on the overall cross-sectional area of the xylem bundle itself. The increase in vascular bundle thickness further seems to correlate with the width of plant axes, and plant height; it is also closely related to the appearance of leaves and increased stomatal density, both of which would increase the demand for water. While wider tracheids with robust walls make it possible to achieve higher water transport tensions, this increases

3400-404: The relationships within the Laurales and between the Laurales and other groups. The following families are included in the Angiosperm Phylogeny Group system ( APG III ): Canellales Piperales Calycanthaceae Siparunaceae Atherospermataceae Gomortegaceae Hernandiaceae Monimiaceae Lauraceae Magnoliales Under the older Cronquist system , the Laurales included

3468-415: The resistance to flow within their cells, thereby increasing the efficiency of their water transport. Bands on the walls of tubes, in fact apparent from the early Silurian onwards, are an early improvisation to aid the easy flow of water. Banded tubes, as well as tubes with pitted ornamentation on their walls, were lignified and, when they form single celled conduits, are considered to be tracheids . These,

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3536-416: The roots (if, for example, the soil is dry), then the gases come out of solution and form a bubble – an embolism forms, which will spread quickly to other adjacent cells, unless bordered pits are present (these have a plug-like structure called a torus, that seals off the opening between adjacent cells and stops the embolism from spreading). Even after an embolism has occurred, plants are able to refill

3604-472: The roots covers the water transport tissue and regulates ion exchange (and prevents unwanted pathogens etc. from entering the water transport system). The endodermis can also provide an upwards pressure, forcing water out of the roots when transpiration is not enough of a driver. Once plants had evolved this level of controlled water transport, they were truly homoiohydric, able to extract water from their environment through root-like organs rather than relying on

3672-445: The roots throughout the plant. It is also used to replace water lost during transpiration and photosynthesis. Xylem sap consists mainly of water and inorganic ions, although it can also contain a number of organic chemicals as well. The transport is passive, not powered by energy spent by the tracheary elements themselves, which are dead by maturity and no longer have living contents. Transporting sap upwards becomes more difficult as

3740-470: The size of the earliest plants. This process demands a steady supply of water from one end, to maintain the chains; to avoid exhausting it, plants developed a waterproof cuticle . Early cuticle may not have had pores but did not cover the entire plant surface, so that gas exchange could continue. However, dehydration at times was inevitable; early plants cope with this by having a lot of water stored between their cell walls, and when it comes to it sticking out

3808-526: The spread of embolism likely facilitated increases in plant size and the colonization of drier habitats during the Devonian radiation . Conifers, by the Jurassic, developed bordered pits had valve-like structures to isolate cavitated elements. These torus-margo structures have an impermeable disc (torus) suspended by a permeable membrane (margo) between two adjacent pores. When a tracheid on one side depressurizes,

3876-568: The spring gentian, are adapted to the alkaline conditions found on calcium -rich chalk and limestone , which give rise to often dry topographies such as limestone pavement . As for their growth habit , the flowering plants range from small, soft herbaceous plants , often living as annuals or biennials that set seed and die after one growing season, to large perennial woody trees that may live for many centuries and grow to many metres in height. Some species grow tall without being self-supporting like trees by climbing on other plants in

3944-462: The support offered by their lignified walls. Defunct tracheids were retained to form a strong, woody stem, produced in most instances by a secondary xylem. However, in early plants, tracheids were too mechanically vulnerable, and retained a central position, with a layer of tough sclerenchyma on the outer rim of the stems. Even when tracheids do take a structural role, they are supported by sclerenchymatic tissue. Tracheids end with walls, which impose

4012-450: The top, the flow is needed to return to the equilibrium. Transpirational pull results from the evaporation of water from the surfaces of cells in the leaves . This evaporation causes the surface of the water to recess into the pores of the cell wall . By capillary action , the water forms concave menisci inside the pores. The high surface tension of water pulls the concavity outwards, generating enough force to lift water as high as

4080-543: The tough times by putting life "on hold" until more water is supplied. To be free from the constraints of small size and constant moisture that the parenchymatic transport system inflicted, plants needed a more efficient water transport system. During the early Silurian , they developed specialized cells, which were lignified (or bore similar chemical compounds) to avoid implosion; this process coincided with cell death, allowing their innards to be emptied and water to be passed through them. These wider, dead, empty cells were

4148-471: The vessels of Gnetum to be convergent with those of angiosperms. Whether the absence of vessels in basal angiosperms is a primitive condition is contested, the alternative hypothesis states that vessel elements originated in a precursor to the angiosperms and were subsequently lost. To photosynthesize, plants must absorb CO 2 from the atmosphere. However, this comes at a price: while stomata are open to allow CO 2 to enter, water can evaporate. Water

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4216-408: The vessels, and gel- and gas-bubble-supported interfacial gradients. Until recently, the differential pressure (suction) of transpirational pull could only be measured indirectly, by applying external pressure with a pressure bomb to counteract it. When the technology to perform direct measurements with a pressure probe was developed, there was initially some doubt about whether the classic theory

4284-554: The world's staple calorie intake, and all three plants are cereals from the Poaceae family (colloquially known as grasses). Other families provide important industrial plant products such as wood , paper and cotton , and supply numerous ingredients for beverages , sugar production , traditional medicine and modern pharmaceuticals . Flowering plants are also commonly grown for decorative purposes , with certain flowers playing significant cultural roles in many societies. Out of

4352-400: The xylem and restore the functionality. The cohesion-tension theory is a theory of intermolecular attraction that explains the process of water flow upwards (against the force of gravity ) through the xylem of plants. It was proposed in 1894 by John Joly and Henry Horatio Dixon . Despite numerous objections, this is the most widely accepted theory for the transport of water through

4420-406: The xylem. However, according to Grew, capillary action in the xylem would raise the sap by only a few inches; to raise the sap to the top of a tree, Grew proposed that the parenchymal cells become turgid and thereby not only squeeze the sap in the tracheids but force some sap from the parenchyma into the tracheids. In 1727, English clergyman and botanist Stephen Hales showed that transpiration by

4488-483: Was coined in the form "Angiospermae" by Paul Hermann in 1690, including only flowering plants whose seeds were enclosed in capsules. The term angiosperm fundamentally changed in meaning in 1827 with Robert Brown , when angiosperm came to mean a seed plant with enclosed ovules. In 1851, with Wilhelm Hofmeister 's work on embryo-sacs, Angiosperm came to have its modern meaning of all the flowering plants including Dicotyledons and Monocotyledons. The APG system treats

4556-468: Was correct, because some workers were unable to demonstrate negative pressures. More recent measurements do tend to validate the classic theory, for the most part. Xylem transport is driven by a combination of transpirational pull from above and root pressure from below, which makes the interpretation of measurements more complicated. Xylem appeared early in the history of terrestrial plant life. Fossil plants with anatomically preserved xylem are known from

4624-412: Was lost per unit of CO 2 uptake. However, even in these "easy" early days, water was at a premium, and had to be transported to parts of the plant from the wet soil to avoid desiccation . This early water transport took advantage of the cohesion-tension mechanism inherent in water. Water has a tendency to diffuse to areas that are drier, and this process is accelerated when water can be wicked along

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