An ecoregion ( ecological region ) is an ecologically and geographically defined area that is smaller than a bioregion , which in turn is smaller than a biogeographic realm . Ecoregions cover relatively large areas of land or water, and contain characteristic, geographically distinct assemblages of natural communities and species . The biodiversity of flora , fauna and ecosystems that characterise an ecoregion tends to be distinct from that of other ecoregions. In theory, biodiversity or conservation ecoregions are relatively large areas of land or water where the probability of encountering different species and communities at any given point remains relatively constant, within an acceptable range of variation (largely undefined at this point). Ecoregions are also known as "ecozones" ("ecological zones"), although that term may also refer to biogeographic realms .
51-537: The Knysna–Amatole montane forests ecoregion , of the tropical and subtropical moist broadleaf forests biome , is in South Africa . It covers an Afromontane area of 3,100 square kilometres (1,200 sq mi) in the Eastern Cape and Western Cape provinces. The ecoregion, which is South Africa's smallest in area, covers two separate enclaves. The ecoregion has a subtropical/warm-temperate climate (Cfb in
102-594: A biogeographical classification system of ecoregions for the United States in a map published in 1976. He subsequently expanded the system to include the rest of North America in 1981, and the world in 1989. The Bailey system, based on climate, is divided into four domains (polar, humid temperate, dry, and humid tropical), with further divisions based on other climate characteristics (subarctic, warm temperate, hot temperate, and subtropical; marine and continental; lowland and mountain). A team of biologists convened by
153-472: A certain vegetation form. Both include many biomes in fact. To divide the world into a few ecological zones is difficult, notably because of the small-scale variations that exist everywhere on earth and because of the gradual changeover from one biome to the other. Their boundaries must therefore be drawn arbitrarily and their characterization made according to the average conditions that predominate in them. A 1978 study on North American grasslands found
204-415: A given ecoregion are more similar to each other than to those of surrounding ecoregions and together form a conservation unit. Freshwater systems include rivers , streams , lakes , and wetlands . Freshwater ecoregions are distinct from terrestrial ecoregions, which identify biotic communities of the land, and marine ecoregions, which are biotic communities of the oceans. A map of Freshwater Ecoregions of
255-469: A holistic, "weight-of-evidence" approach where the importance of various factors may vary. An example of the algorithmic approach is Robert Bailey 's work for the U.S. Forest Service, which uses a hierarchical classification that first divides land areas into very large regions based on climatic factors, and subdivides these regions, based first on dominant potential vegetation, and then by geomorphology and soil characteristics. The weight-of-evidence approach
306-425: A positive logistic correlation between evapotranspiration in mm/yr and above-ground net primary production in g/m /yr. The general results from the study were that precipitation and water use led to above-ground primary production, while solar irradiation and temperature lead to below-ground primary production (roots), and temperature and water lead to cool and warm season growth habit. These findings help explain
357-509: A set of ecoregions identified by WWF whose conservation would achieve the goal of saving a broad diversity of the Earth's ecosystems, includes a number of areas highlighted for their freshwater biodiversity values. The Global 200 preceded Freshwater Ecoregions of the World and incorporated information from regional freshwater ecoregional assessments that had been completed at that time. Sources related to
408-455: A variety of habitats . While a biome can cover small areas, a microbiome is a mix of organisms that coexist in a defined space on a much smaller scale. For example, the human microbiome is the collection of bacteria, viruses, and other microorganisms that are present on or in a human body. A biota is the total collection of organisms of a geographic region or a time period, from local geographic scales and instantaneous temporal scales all
459-433: A way to recognize the irreversible coupling of human and ecological systems at global scales and manage Earth's biosphere and anthropogenic biomes. Major anthropogenic biomes: The endolithic biome, consisting entirely of microscopic life in rock pores and cracks, kilometers beneath the surface, has only recently been discovered, and does not fit well into most classification schemes. Anthropogenic climate change has
510-407: A web application developed by Resolve and Google Earth Engine. An ecoregion is a "recurring pattern of ecosystems associated with characteristic combinations of soil and landform that characterise that region". Omernik (2004) elaborates on this by defining ecoregions as: "areas within which there is spatial coincidence in characteristics of geographical phenomena associated with differences in
561-503: Is woody plant encroachment , which can change grass savanna into shrub savanna. Average temperatures have risen more than twice the usual amount in both arctic and mountainous biomes, which leads to the conclusion that arctic and mountainous biomes are currently the most vulnerable to climate change. South American terrestrial biomes have been predicted to go through the same temperature trends as arctic and mountainous biomes. With its annual average temperature continuing to increase,
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#1732854982940612-426: Is analogous to that used for terrestrial ecoregions. Major habitat types are identified: polar, temperate shelves and seas, temperate upwelling, tropical upwelling, tropical coral, pelagic (trades and westerlies), abyssal, and hadal (ocean trench). These correspond to the terrestrial biomes . The Global 200 classification of marine ecoregions is not developed to the same level of detail and comprehensiveness as that of
663-596: Is exemplified by James Omernik's work for the United States Environmental Protection Agency , subsequently adopted (with modification) for North America by the Commission for Environmental Cooperation . The intended purpose of ecoregion delineation may affect the method used. For example, the WWF ecoregions were developed to aid in biodiversity conservation planning, and place a greater emphasis than
714-400: Is used when applied to plant communities only, while biome is used when concerned with both plants and animals. Whittaker's convention of biome-type or formation-type is a broader method to categorize similar communities. Whittaker used what he called "gradient analysis" of ecocline patterns to relate communities to climate on a worldwide scale. Whittaker considered four main ecoclines in
765-599: The Himalayas and the Sahara . The boundaries of ecoregions are often not as decisive or well recognized, and are subject to greater disagreement. Ecoregions are classified by biome type, which are the major global plant communities determined by rainfall and climate. Forests, grasslands (including savanna and shrubland), and deserts (including xeric shrublands ) are distinguished by climate ( tropical and subtropical vs. temperate and boreal climates) and, for forests, by whether
816-964: The Köppen climate classification ). Rainfall occurs year-round, and ranges from 525 mm to 1,220 mm per year in the Knysna forest, and from 750 mm to 1,500 mm in the Amatole forests. The trees are of tropical and afromontane origin, and include ironwood ( Olea capensis ), stinkwood ( Ocotea bullata ), Outeniqua yellowwood ( Afrocarpus falcatus ), real yellowwood ( Podocarpus latifolius ), Cape holly ( Ilex mitis ), white pear ( Apodytes dimidiata ), Cape beech ( Rapanea melanophloeos ), bastard saffron ( Cassine peragua ), Cape plane ( Ochna arborea var. arborea ), assegai tree ( Curtisia dentata ), kamassi ( Gonioma kamassi ), white alder ( Platylophus trifoliatus ), and red alder ( Cunonia capensis ). The forests were home to African elephants , but
867-594: The World Wildlife Fund (WWF) developed a scheme that divided the world's land area into biogeographic realms (called "ecozones" in a BBC scheme), and these into ecoregions (Olson & Dinerstein, 1998, etc.). Each ecoregion is characterized by a main biome (also called major habitat type). This classification is used to define the Global 200 list of ecoregions identified by the WWF as priorities for conservation. For
918-550: The terrestrial ecoregions , there is a specific EcoID, format XXnnNN (XX is the biogeographic realm , nn is the biome number, NN is the individual number). The applicability of the realms scheme above - based on Udvardy (1975)—to most freshwater taxa is unresolved. According to the WWF, the following are classified as freshwater biomes: Biomes of the coastal and continental shelf areas ( neritic zone ): Example: Pruvot (1896) zones or "systems": Longhurst (1998) biomes : Other marine habitat types (not covered yet by
969-450: The "ecoregion" as a unit of analysis. The " Global 200 " is the list of ecoregions identified by WWF as priorities for conservation . Terrestrial ecoregions are land ecoregions, as distinct from freshwater and marine ecoregions. In this context, terrestrial is used to mean "of land" (soil and rock), rather than the more general sense "of Earth " (which includes land and oceans). WWF (World Wildlife Fund) ecologists currently divide
1020-866: The 18th century, and the Amatole forests since the 20th century. Since 1939 the forests have mostly been within protected areas and are recovering well, although managed timber harvesting is allowed. Ecoregion Three caveats are appropriate for all bio-geographic mapping approaches. Firstly, no single bio-geographic framework is optimal for all taxa. Ecoregions reflect the best compromise for as many taxa as possible. Secondly, ecoregion boundaries rarely form abrupt edges; rather, ecotones and mosaic habitats bound them. Thirdly, most ecoregions contain habitats that differ from their assigned biome . Biogeographic provinces may originate due to various barriers, including physical (plate tectonics, topographic highs), climatic (latitudinal variation, seasonal range) and ocean chemical related (salinity, oxygen levels). The history of
1071-558: The Earth. The use of the term ecoregion is an outgrowth of a surge of interest in ecosystems and their functioning. In particular, there is awareness of issues relating to spatial scale in the study and management of landscapes . It is widely recognized that interlinked ecosystems combine to form a whole that is "greater than the sum of its parts". There are many attempts to respond to ecosystems in an integrated way to achieve "multi-functional" landscapes, and various interest groups from agricultural researchers to conservationists are using
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#17328549829401122-639: The Global 200/WWF scheme): Humans have altered global patterns of biodiversity and ecosystem processes. As a result, vegetation forms predicted by conventional biome systems can no longer be observed across much of Earth's land surface as they have been replaced by crop and rangelands or cities. Anthropogenic biomes provide an alternative view of the terrestrial biosphere based on global patterns of sustained direct human interaction with ecosystems, including agriculture , human settlements , urbanization , forestry and other uses of land . Anthropogenic biomes offer
1173-547: The Omernik or Bailey systems on floral and faunal differences between regions. The WWF classification defines an ecoregion as: A large area of land or water that contains a geographically distinct assemblage of natural communities that: According to WWF, the boundaries of an ecoregion approximate the original extent of the natural communities prior to any major recent disruptions or changes. WWF has identified 867 terrestrial ecoregions, and approximately 450 freshwater ecoregions across
1224-472: The WWC scheme: Others: Biome A biome ( / ˈ b aɪ . oʊ m / ) is a distinct geographical region with specific climate , vegetation , and animal life . It consists of a biological community that has formed in response to its physical environment and regional climate . Biomes may span more than one continent. A biome encompasses multiple ecosystems within its boundaries. It can also comprise
1275-514: The WWF concept prioritizes biogeography, that is, the distribution of distinct species assemblages. In 2017, an updated terrestrial ecoregions dataset was released in the paper "An Ecoregion-Based Approach to Protecting Half the Terrestrial Realm" led by E. Dinerstein with 48 co-authors. Using recent advances in satellite imagery the ecoregion perimeters were refined and the total number reduced to 846 (and later 844), which can be explored on
1326-519: The World (TEOW), led by D. Olsen, E. Dinerstein, E. Wikramanayake, and N. Burgess. While the two approaches are related, the Bailey ecoregions (nested in four levels) give more importance to ecological criteria and climate zones, while the WWF ecoregions give more importance to biogeography, that is, the distribution of distinct species assemblages. The TEOW framework originally delineated 867 terrestrial ecoregions nested into 14 major biomes, contained with
1377-439: The World , edited by David W. Goodall , provides a comprehensive coverage of the major "ecosystem types or biomes" on Earth: The eponymously named Heinrich Walter classification scheme considers the seasonality of temperature and precipitation. The system, also assessing precipitation and temperature, finds nine major biome types, with the important climate traits and vegetation types . The boundaries of each biome correlate to
1428-783: The World, released in 2008, has 426 ecoregions covering virtually the entire non-marine surface of the earth. World Wildlife Fund (WWF) identifies twelve major habitat types of freshwater ecoregions: Large lakes, large river deltas, polar freshwaters, montane freshwaters, temperate coastal rivers, temperate floodplain rivers and wetlands, temperate upland rivers, tropical and subtropical coastal rivers, tropical and subtropical floodplain rivers and wetlands, tropical and subtropical upland rivers, xeric freshwaters and endorheic basins, and oceanic islands. The freshwater major habitat types reflect groupings of ecoregions with similar biological, chemical, and physical characteristics and are roughly equivalent to biomes for terrestrial systems. The Global 200 ,
1479-467: The animal element and the exclusion of the taxonomic element of species composition . In 1935, Tansley added the climatic and soil aspects to the idea, calling it ecosystem . The International Biological Program (1964–74) projects popularized the concept of biome. However, in some contexts, the term biome is used in a different manner. In German literature, particularly in the Walter terminology,
1530-480: The authors was to support global biodiversity conservation by providing a "fourfold increase in resolution over that of the 198 biotic provinces of Dasmann (1974) and the 193 units of Udvardy (1975)." In 2007, a comparable set of Marine Ecoregions of the World (MEOW) was published, led by M. Spalding, and in 2008 a set of Freshwater Ecoregions of the World (FEOW) was published, led by R. Abell. Bailey's ecoregion concept prioritizes ecological criteria and climate, while
1581-493: The biological effects of temperature and rainfall on vegetation under the assumption that these two abiotic factors are the largest determinants of the types of vegetation found in a habitat. Holdridge uses the four axes to define 30 so-called "humidity provinces", which are clearly visible in his diagram. While this scheme largely ignores soil and sun exposure, Holdridge acknowledged that these were important. The principal biome-types by Allee (1949): The principal biomes of
Knysna–Amatole montane forests - Misplaced Pages Continue
1632-677: The broad latitudinal divisions of polar, temperate, and tropical seas, with subdivisions based on ocean basins (except for the southern hemisphere temperate oceans, which are based on continents). Major marine biogeographic realms, analogous to the eight terrestrial biogeographic realms , represent large regions of the ocean basins: Arctic , Temperate Northern Atlantic , Temperate Northern Pacific , Tropical Atlantic , Western Indo-Pacific , Central Indo-Pacific , Eastern Indo-Pacific , Tropical Eastern Pacific , Temperate South America , Temperate Southern Africa , Temperate Australasia , and Southern Ocean . A similar system of identifying areas of
1683-533: The categories used in Holdridge's bioclassification scheme (see below), which were then later simplified by Whittaker. The number of classification schemes and the variety of determinants used in those schemes, however, should be taken as strong indicators that biomes do not fit perfectly into the classification schemes created. In 1947, the American botanist and climatologist Leslie Holdridge classified climates based on
1734-449: The conditions of moisture and cold stress that are strong determinants of plant form, and therefore the vegetation that defines the region. Extreme conditions, such as flooding in a swamp, can create different kinds of communities within the same biome. Schultz (1988, 2005) defined nine ecozones (his concept of ecozone is more similar to the concept of biome than to the concept of ecozone of BBC): Robert G. Bailey nearly developed
1785-495: The crowned eagle ( Stephanoaetus coronatus ) and the African wood owl ( Strix woodfordii ). Reptiles include the endemic Knysna dwarf chameleon ( Bradypodion damaranum ). Thomas Henry Duthie was the first appointed Supervisor of Crown Forests and Lands. Despite the small size of the ecoregion, the Knysna and Amatole forests are South Africa's largest individual forests. The Knysna forest has been exploited for valuable timber since
1836-401: The delineation of ecoregions an imperfect science. Another complication is that environmental conditions across an ecoregion boundary may change very gradually, e.g. the prairie-forest transition in the midwestern United States, making it difficult to identify an exact dividing boundary. Such transition zones are called ecotones . Ecoregions can be categorized using an algorithmic approach or
1887-405: The early 1970s, the term 'ecoregion' was introduced (short for ecological region), and R.G. Bailey published the first comprehensive map of U.S. ecoregions in 1976. The term was used widely in scholarly literature in the 1980s and 1990s, and in 2001 scientists at the U.S. conservation organization World Wildlife Fund (WWF) codified and published the first global-scale map of Terrestrial Ecoregions of
1938-501: The land surface of the Earth into eight biogeographical realms containing 867 smaller terrestrial ecoregions (see list ). The WWF effort is a synthesis of many previous efforts to define and classify ecoregions. The eight realms follow the major floral and faunal boundaries, identified by botanists and zoologists, that separate the world's major plant and animal communities. Realm boundaries generally follow continental boundaries, or major barriers to plant and animal distribution, like
1989-664: The last count (2017-2019) suggested that only one adult female, age 45 years, survives. In addition African leopard , bushbuck , blue duiker , bushpig and other mammals are resident. The density and assortment of birds is not exceptionally high, but the buff-spotted flufftail ( Sarothrura elegans ), Narina trogon ( Apaloderma narina ), Knysna lourie ( Tauraco corythaix ), seven species of cuckoo, Knysna woodpecker ( Campethera notata ), Knysna warbler ( Bradypterus sylvaticus ), chorister robin-chat ( Cossypha dichroa ), Cape batis ( Batis capensis ) and forest canary ( Serinus scotops ) occur, while birds of prey found here include
2040-463: The oceans for conservation purposes is the system of large marine ecosystems (LMEs), developed by the US National Oceanic and Atmospheric Administration (NOAA). A freshwater ecoregion is a large area encompassing one or more freshwater systems that contains a distinct assemblage of natural freshwater communities and species. The freshwater species, dynamics, and environmental conditions within
2091-410: The potential to greatly alter the distribution of Earth's biomes. Meaning, biomes around the world could change so much that they would be at risk of becoming new biomes entirely. More specifically, between 54% and 22% of global land area will experience climates that correspond to other biomes. 3.6% of land area will experience climates that are completely new or unusual. An example of a biome shift
Knysna–Amatole montane forests - Misplaced Pages Continue
2142-415: The quality, health, and integrity of ecosystems ". "Characteristics of geographical phenomena" may include geology , physiography , vegetation, climate, hydrology , terrestrial and aquatic fauna , and soils, and may or may not include the impacts of human activity (e.g. land use patterns, vegetation changes). There is significant, but not absolute, spatial correlation among these characteristics, making
2193-420: The term biome is sometimes used as a synonym of biogeographic province , an area based on species composition (the term floristic province being used when plant species are considered), or also as synonym of the "morphoclimatic and phytogeographical domain" of Ab'Sáber , a geographic space with subcontinental dimensions, with the predominance of similar geomorphologic and climatic characteristics, and of
2244-408: The term is somewhat vague. It has been used in many contexts: forest classifications (Loucks, 1962), biome classifications (Bailey, 1976, 2014), biogeographic classifications ( WWF / Global 200 scheme of Olson & Dinerstein, 1998), etc. The phrase "ecological region" was widely used throughout the 20th century by biologists and zoologists to define specific geographic areas in research. In
2295-468: The term is used similarly as biotope (a concrete geographical unit), while the biome definition used in this article is used as an international, non-regional, terminology—irrespectively of the continent in which an area is present, it takes the same biome name—and corresponds to his "zonobiome", "orobiome" and "pedobiome" (biomes determined by climate zone, altitude or soil). In the Brazilian literature,
2346-454: The terrestrial ecoregions; only the priority conservation areas are listed. See Global 200 Marine ecoregions for a full list of marine ecoregions. In 2007, TNC and WWF refined and expanded this scheme to provide a system of comprehensive near shore (to 200 meters depth) Marine Ecoregions of the World (MEOW). The 232 individual marine ecoregions are grouped into 62 marine provinces , which in turn group into 12 marine realms , which represent
2397-602: The terrestrial realm. Along these gradients, Whittaker noted several trends that allowed him to qualitatively establish biome-types: Whittaker summed the effects of gradients (3) and (4) to get an overall temperature gradient and combined this with a gradient (2), the moisture gradient, to express the above conclusions in what is known as the Whittaker classification scheme. The scheme graphs average annual precipitation (x-axis) versus average annual temperature (y-axis) to classify biome-types. The multi-authored series Ecosystems of
2448-913: The trees are predominantly conifers ( gymnosperms ), or whether they are predominantly broadleaf ( Angiosperms ) and mixed (broadleaf and conifer). Biome types like Mediterranean forests, woodlands, and scrub ; tundra ; and mangroves host very distinct ecological communities, and are recognized as distinct biome types as well. Marine ecoregions are: "Areas of relatively homogeneous species composition , clearly distinct from adjacent systems….In ecological terms, these are strongly cohesive units, sufficiently large to encompass ecological or life history processes for most sedentary species." They have been defined by The Nature Conservancy (TNC) and World Wildlife Fund (WWF) to aid in conservation activities for marine ecosystems . Forty-three priority marine ecoregions were delineated as part of WWF's Global 200 efforts. The scheme used to designate and classify marine ecoregions
2499-410: The way up to whole-planet and whole-timescale spatiotemporal scales. The biotas of the Earth make up the biosphere . The term was suggested in 1916 by Clements , originally as a synonym for biotic community of Möbius (1877). Later, it gained its current definition, based on earlier concepts of phytophysiognomy , formation and vegetation (used in opposition to flora ), with the inclusion of
2550-476: The world by Kendeigh (1961): Whittaker classified biomes using two abiotic factors: precipitation and temperature. His scheme can be seen as a simplification of Holdridge's; more readily accessible, but missing Holdridge's greater specificity. Whittaker based his approach on theoretical assertions and empirical sampling. He had previously compiled a review of biome classifications. Whittaker's distinction between biome and formation can be simplified: formation
2601-486: The world's 8 major biogeographical realms. Subsequent regional papers by the co-authors covering Africa, Indo-Pacific, and Latin America differentiate between ecoregions and bioregions, referring to the latter as "geographic clusters of ecoregions that may span several habitat types, but have strong biogeographic affinities, particularly at taxonomic levels higher than the species level (genus, family)". The specific goal of
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