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128-548: The Tournaisian was named after the Belgian city of Tournai . It was introduced in scientific literature by Belgian geologist André Hubert Dumont in 1832. Like many Devonian and lower Carboniferous stages, the Tournaisian is a unit from West European regional stratigraphy that is now used in the official international time scale. The Tournaisian correlates with the regional North American Kinderhookian and lower Osagean stages and

256-463: A balanced budget, some tax reforms , a labor-market reform, scheduled nuclear phase-out and instigated legislation allowing more stringent war crime and more lenient soft drug usage prosecution. Restrictions on euthanasia were reduced, and in 2003, Belgium became one of the first countries in the world to legalise same-sex marriage . The government promoted active diplomacy in Africa and opposed

384-476: A combination of historical factors such as: speciation , extinction , continental drift , and glaciation . Through observing the geographic distribution of species, we can see associated variations in sea level , river routes, habitat, and river capture . Additionally, this science considers the geographic constraints of landmass areas and isolation, as well as the available ecosystem energy supplies. Over periods of ecological changes, biogeography includes

512-828: A decade after the other regions) is included in both the Flemish and French Communities, and the territory of the German-speaking Community lies wholly within the Walloon Region. Conflicts about jurisdiction between the bodies are resolved by the Constitutional Court of Belgium . The structure is intended as a compromise to allow different cultures to live together peacefully. The Federal State's authority includes justice, defense, federal police, social security, nuclear energy, monetary policy and public debt, and other aspects of public finances. State-owned companies include

640-421: A global scale. GIS can show certain processes on the earth's surface like whale locations, sea surface temperatures , and bathymetry. Current scientists also use coral reefs to delve into the history of biogeography through the fossilized reefs. Two global information systems are either dedicated to, or have strong focus on, biogeography (in the form of the spatial location of observations of organisms), namely

768-419: A great impact on Charles Darwin , who was inspired to consider species adaptations and evolution after learning about botanical geography. De Candolle was the first to describe the differences between the small-scale and large-scale distribution patterns of organisms around the globe. Several additional scientists contributed new theories to further develop the concept of biogeography. Charles Lyell developed

896-749: A habitat and species of organisms describe the ecological application of biogeography. Historical biogeography describes the long-term, evolutionary periods of time for broader classifications of organisms. Early scientists, beginning with Carl Linnaeus , contributed to the development of biogeography as a science. The scientific theory of biogeography grows out of the work of Alexander von Humboldt (1769–1859), Francisco Jose de Caldas (1768–1816), Hewett Cottrell Watson (1804–1881), Alphonse de Candolle (1806–1893), Alfred Russel Wallace (1823–1913), Philip Lutley Sclater (1829–1913) and other biologists and explorers. The patterns of species distribution across geographical areas can usually be explained through

1024-668: A large semi-independent enclave. The Eighty Years' War (1568–1648) was triggered by the Spanish government's policy towards Protestantism , which was becoming popular in the Low Countries. The rebellious northern United Provinces ( Belgica Foederata in Latin , the "Federated Netherlands") eventually separated from the Southern Netherlands ( Belgica Regia , the "Royal Netherlands"). The southern part continued to be ruled successively by

1152-539: A new government . By 30 March 2011, this set a new world record for the elapsed time without an official government, previously held by war-torn Iraq . Finally, in December 2011 the Di Rupo Government led by Walloon socialist Prime Minister Elio Di Rupo was sworn in. The 2014 federal election (coinciding with the regional elections ) resulted in a further electoral gain for the Flemish nationalist N-VA, although

1280-413: A number of methods have been developed to produce arguably more complete "predictive" or "modelled" distributions for species based on their associated environmental or other preferences (such as availability of food or other habitat requirements); this approach is known as either Environmental niche modelling (ENM) or Species distribution modelling (SDM). Depending on the reliability of the source data and

1408-945: A policy of mediation. The Belgians have been strong advocates of European integration . The headquarters of NATO and of several of the institutions of the European Union are located in Belgium. The Belgian Armed Forces had 23,200 active personnel in 2023, including 8,500 in the Land Component , 1,400 in the Naval Component , 4,900 in the Air Component , 1,450 in the Medical Component , and 6,950 in joint service, in addition to 5,900 reserve personnel. In 2019, Belgium's defense budget totaled €4.303 billion ($ 4.921 billion) representing .93% of its GDP. The operational commands of

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1536-987: A result of tectonic uplift (or subsidence ), natural damming created by a landslide , or headward or lateral erosion of the watershed between adjacent basins. Biogeography is a synthetic science, related to geography , biology , soil science , geology , climatology , ecology and evolution . Some fundamental concepts in biogeography include: The study of comparative biogeography can follow two main lines of investigation: There are many types of biogeographic units used in biogeographic regionalisation schemes, as there are many criteria ( species composition , physiognomy , ecological aspects) and hierarchization schemes: biogeographic realms (ecozones), bioregions ( sensu stricto ), ecoregions , zoogeographical regions , floristic regions , vegetation types, biomes , etc. The terms biogeographic unit, biogeographic area can be used for these categories, regardless of rank. In 2008, an International Code of Area Nomenclature

1664-488: A whole suite of predictor variables for biogeographic analysis, including satellite imaging and processing of the Earth. Two main types of satellite imaging that are important within modern biogeography are Global Production Efficiency Model (GLO-PEM) and Geographic Information Systems (GIS). GLO-PEM uses satellite-imaging gives "repetitive, spatially contiguous, and time specific observations of vegetation". These observations are on

1792-581: Is a sovereign state and a federal constitutional monarchy with a parliamentary system . Its institutional organization is complex and is structured on both regional and linguistic grounds. It is divided into three highly autonomous regions : the Flemish Region (Flanders) in the north, the Walloon Region (Wallonia) in the south, and the Brussels-Capital Region in the middle. Brussels

1920-424: Is an alternate view than that of Linnaeus. Buffon's law eventually became a principle of biogeography by explaining how similar environments were habitats for comparable types of organisms. Buffon also studied fossils which led him to believe that the Earth was over tens of thousands of years old, and that humans had not lived there long in comparison to the age of the Earth. Following the period of exploration came

2048-502: Is an integrative field of inquiry that unites concepts and information from ecology , evolutionary biology , taxonomy , geology , physical geography , palaeontology , and climatology . Modern biogeographic research combines information and ideas from many fields, from the physiological and ecological constraints on organismal dispersal to geological and climatological phenomena operating at global spatial scales and evolutionary time frames. The short-term interactions within

2176-440: Is around 23% of the total land area, equivalent to 689,300 hectares (ha) of forest in 2020, up from 677,400 hectares (ha) in 1990. In 2020, naturally regenerating forest covered 251,200 hectares (ha) and planted forest covered 438,200 hectares (ha). For the year 2015, 47% of the forest area was reported to be under public ownership , 53% private ownership and 0% with ownership listed as other or unknown. The territory of Belgium

2304-477: Is based on civil law and originates from the Napoleonic code . The Court of Cassation is the court of last resort, with the courts of appeal one level below. Belgium's political institutions are complex; most political power rests on representation of the main cultural communities . Since about 1970, the significant national Belgian political parties have split into distinct components that mainly represent

2432-495: Is divided into three Regions, two of which, the Flemish Region and Walloon Region , are in turn subdivided into provinces ; the third Region, the Brussels Capital Region , is neither a province nor a part of a province. Belgium is a constitutional , popular monarchy and a federal parliamentary democracy . The bicameral federal parliament is composed of a Senate and a Chamber of Representatives . The former

2560-404: Is in the geological similarities between varying locations around the globe, the geographic distribution of some fossils (including the mesosaurs ) on various continents, and the jigsaw puzzle shape of the landmasses on Earth. Though Wegener did not know the mechanism of this concept of Continental Drift, this contribution to the study of biogeography was significant in the way that it shed light on

2688-619: Is just above this and was used as a base for the Carboniferous in the past. The GSSP for the Tournaisian is near the summit of La Serre hill, in the Lydiennes Formation of the commune of Cabrières , in the Montagne Noire (southern France). The GSSP is in a section on the southern side of the hill, in an 80 cm deep trench, about 125 m south of the summit, 2.5 km southwest of the village of Cabrières and 2.5 km north of

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2816-658: Is lowest in January at 3 °C (37.4 °F) and highest in July at 18 °C (64.4 °F). The average precipitation per month varies between 54 mm (2.1 in) for February and April, to 78 mm (3.1 in) for July. Averages for the years 2000 to 2006 show daily temperature minimums of 7 °C (44.6 °F) and maximums of 14 °C (57.2 °F) and monthly rainfall of 74 mm (2.9 in); these are about 1 °C and nearly 10 millimeters above last century's normal values, respectively. Phytogeographically , Belgium

2944-493: Is made up of 50 senators appointed by the parliaments of the communities and regions and 10 co-opted senators . Prior to 2014, most of the Senate's members were directly elected. The Chamber 's 150 representatives are elected under a proportional voting system from 11 electoral districts . Belgium has compulsory voting and thus maintains one of the highest rates of voter turnout in the world. The King (currently Philippe )

3072-639: Is more than double that of Flanders, which boomed after the Second World War. Belgium is a developed country , with an advanced high-income economy . The country is one of the six founding members of the European Union , and its capital, Brussels, is the de facto capital of the European Union itself, hosting the official seats of the European Commission , the Council of the European Union , and

3200-717: Is now Europe, North America, and China, represent a low-latitude paleobiogeographical realm known as the Amerosinian realm. Divaricating (widely branching) trunks of Lepidodendropsis lycophytes are by far the most abundant and widespread plant fossils of the Tournaisian, yet there was some minor variation in other flora through time and space. In eastern North America, lyginopterids and probable progymnosperms were also common, as indicated by leaf form genera such as Adiantites , Rhodeopteridium , and Genselia . The progymnosperm leaf Triphyllopteris may be more common in Europe while

3328-608: Is now the Benelux, the so-called Burgundian Netherlands . "Burgundy" and "Flanders" were the first two common names used for the Burgundian Netherlands which was the predecessor of the Austrian Netherlands, the predecessor of modern Belgium. The union, technically stretching between two kingdoms, gave the area economic and political stability which led to an even greater prosperity and artistic creation. Born in Belgium,

3456-473: Is on how the environment and humans affect the distribution of species as well as other manifestations of Life such as species or genetic diversity. Biogeography is being applied to biodiversity conservation and planning, projecting global environmental changes on species and biomes, projecting the spread of infectious diseases, invasive species, and for supporting planning for the establishment of crops. Technological evolving and advances have allowed for generating

3584-698: Is shared between the Atlantic European and Central European provinces of the Circumboreal Region within the Boreal Kingdom . According to the World Wide Fund for Nature , the territory of Belgium belongs to the terrestrial ecoregions of Atlantic mixed forests and Western European broadleaf forests . Belgium had a 2018 Forest Landscape Integrity Index mean score of 1.36/10, ranking it 163rd globally out of 172 countries. In Belgium forest cover

3712-614: Is the head of state , though with limited prerogatives . He appoints ministers, including a Prime Minister, that have the confidence of the Chamber of Representatives to form the federal government . The Council of Ministers is composed of no more than fifteen members. With the possible exception of the Prime Minister, the Council of Ministers is composed of an equal number of Dutch-speaking members and French-speaking members. The judicial system

3840-448: Is the branch of biogeography that studies the distribution of plants. Zoogeography is the branch that studies distribution of animals. Mycogeography is the branch that studies distribution of fungi, such as mushrooms . Knowledge of spatial variation in the numbers and types of organisms is as vital to us today as it was to our early human ancestors , as we adapt to heterogeneous but geographically predictable environments . Biogeography

3968-554: Is the smallest and most densely populated region, as well as the richest region in terms of GDP per capita . Belgium is also home to two main linguistic communities: the Flemish Community ( Dutch -speaking), which constitutes about 60 percent of the population, and the French Community ( French -speaking), which constitutes about 40 percent of the population. A small German-speaking Community , making up around one percent of

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4096-605: The 22nd most densely populated country in the world and the 6th most densely populated country in Europe, with a density of 383/km (990/sq mi). Belgium is part of an area known as the Low Countries , historically a somewhat larger region than the Benelux group of states, as it also included parts of northern France. The capital and largest metropolitan region is Brussels ; other major cities are Antwerp , Ghent , Charleroi , Liège , Bruges , Namur , and Leuven . Belgium

4224-558: The Age of Enlightenment in Europe, which attempted to explain the patterns of biodiversity observed by Buffon and Linnaeus. At the birth of the 19th century, Alexander von Humboldt, known as the "founder of plant geography", developed the concept of physique generale to demonstrate the unity of science and how species fit together. As one of the first to contribute empirical data to the science of biogeography through his travel as an explorer, he observed differences in climate and vegetation. The Earth

4352-492: The Belgian Post Group and Belgian Railways . The Federal Government is responsible for the obligations of Belgium and its federalized institutions towards the European Union and NATO. It controls substantial parts of public health, home affairs and foreign affairs. The budget—without the debt—controlled by the federal government amounts to about 50% of the national fiscal income. The federal government employs around 12% of

4480-682: The European Council , as well as one of two seats of the European Parliament (the other being Strasbourg ). Belgium is also a founding member of the Eurozone , NATO , OECD , and WTO , and a part of the trilateral Benelux Union and the Schengen Area . Brussels also hosts the headquarters of many major international organizations, such as NATO . According to Julius Caesar , the Belgae were

4608-639: The European Union  (light green) Belgium , officially the Kingdom of Belgium , is a country in Northwestern Europe . The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to the south, and the North Sea to the west. It covers an area of 30,689 km (11,849 sq mi) and has a population of more than 11.7 million, making it

4736-667: The Global Biodiversity Information Facility (GBIF: 2.57 billion species occurrence records reported as at August 2023) and, for marine species only, the Ocean Biodiversity Information System (OBIS, originally the Ocean Biogeographic Information System : 116 million species occurrence records reported as at August 2023), while at a national scale, similar compilations of species occurrence records also exist such as

4864-715: The Habsburg Emperor Charles ;V was heir of the Burgundians, but also of the royal families of Austria , Castile and Aragon . With the Pragmatic Sanction of 1549 he gave the Seventeen Provinces more legitimacy as a stable entity, rather than just a temporary personal union . He also increased the influence of these Netherlands over the Prince-Bishopric of Liège , which continued to exist as

4992-524: The Holy Roman Empire . The lordships and bishoprics along the "March" (frontier) between the two great kingdoms maintained important connections between each other. For example, the county of Flanders expanded over the Scheldt into the empire, and during several periods was ruled by the same lords as the county of Hainaut . In the 13th and 14th centuries, the cloth industry and commerce boomed especially in

5120-701: The Industrial Revolution , and during the course of the 20th century, possessed a number of colonies , notably the Belgian Congo and Ruanda-Urundi . These colonies gained independence between 1960 and 1962. The second half of the 20th century was marked by rising tensions between the Dutch-speakers and French-speakers, fueled by differences in political culture and the unequal economic development of Flanders and Wallonia. This continuing antagonism has led to several far-reaching state reforms , resulting in

5248-785: The Michel Government . It was the first time N-VA was part of the federal cabinet, while the French-speaking side was represented only by the MR, which achieved a minority of the public votes in Wallonia. In May 2019 federal elections in the Flemish-speaking northern region of Flanders far-right Vlaams Belang party made major gains. In the French-speaking southern area of Wallonia the Socialists were strong. The moderate Flemish nationalist party

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5376-412: The Netherlands ( 450 km ). Its total surface, including water area, is 30,689 km (11,849 sq mi). Before 2018, its total area was believed to be 30,528 km (11,787 sq mi). However, when the country's statistics were measured in 2018, a new calculation method was used. Unlike previous calculations, this one included the area from the coast to the low-water line, revealing

5504-722: The Spanish ( Spanish Netherlands ) and the Austrian House of Habsburgs ( Austrian Netherlands ) and comprised most of modern Belgium. This was the theatre of several more protracted conflicts during much of the 17th and 18th centuries involving France, including the Franco-Dutch War (1672–1678), the Nine Years' War (1688–1697), the War of the Spanish Succession (1701–1714), and part of

5632-721: The United Kingdom of the Netherlands occurred at the dissolution of the First French Empire in 1814, after the abdication of Napoleon. In 1830, the Belgian Revolution led to the re-separation of the Southern Provinces from the Netherlands and to the establishment of a Catholic and bourgeois, officially French-speaking and neutral, independent Belgium under a provisional government and a national congress . Since

5760-712: The War of the Austrian Succession (1740–1748). Following the campaigns of 1794 in the French Revolutionary Wars , the Low Countries ;– including territories that were never nominally under Habsburg rule, such as the Prince-Bishopric of Liège – were annexed by the French First Republic , ending Austrian rule in the region. A reunification of the Low Countries as

5888-650: The Western Roman Empire , the Roman provinces of Belgica and Germania were inhabited by a mix of a Romanized population and Germanic -speaking Franks who came to dominate the military and political class. During the 5th century, the area came under the rule of the Frankish Merovingian kings, who initially established a kingdom ruling over the Romanized population in what is now northern France, and then conquered

6016-413: The miospore index fossil Waltzispora lanzonii . The floral diversity of Tournaisian southern tundra consists almost entirely of relict Devonian genera; this suggests that Late Devonian land plant extinctions in lower latitudes were mostly driven by competition from new tropical species, rather than global environmental pressures. Trilobites experienced their final substantial diversification event in

6144-421: The war against the Islamic State , and in eastern Europe for the NATO presence there. Biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time . Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude , elevation , isolation and habitat area . Phytogeography

6272-422: The 19th century were the Catholic Party and the Liberal Party , with the Belgian Labour Party emerging towards the end of the 19th century. French was originally the official language used by the nobility and the bourgeoisie , especially after the rejection of the Dutch monarchy. French progressively lost its dominance as Dutch began to recover its status. This recognition became official in 1898, and in 1967,

6400-430: The 36 volume Histoire Naturelle, générale et particulière , in which he argued that varying geographical regions would have different forms of life. This was inspired by his observations comparing the Old and New World, as he determined distinct variations of species from the two regions. Buffon believed there was a single species creation event, and that different regions of the world were homes for varying species, which

6528-456: The Amazon basin, Orinoco basin, and Guianas ) with an exceptionally low (flat) topographic relief, the many waterways have had a highly reticulated history over geological time . In such a context, stream capture is an important factor affecting the evolution and distribution of freshwater organisms. Stream capture occurs when an upstream portion of one river drainage is diverted to the downstream portion of an adjacent basin. This can happen as

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6656-471: The Angaran realm. The most common plant fossils in this region were shrub-sized lycophytes such as Ursodendron and Tomiodendron , shorter than their arborescent tropical relatives. Gondwanan plant fossils are uncommon: southernmost Gondwana was covered by dwarf lycophytes, even smaller than those of the Angaran realm. Subtropical and temperate lycophytes such as Lepidodendropsis , Archaeosigillaria , and Frenguellia could be found in some parts of

6784-474: The Ardennes are more rugged and rocky with caves and small gorges . Extending westward into France, this area is eastwardly connected to the Eifel in Germany by the High Fens plateau, on which the Signal de Botrange forms the country's highest point at 694 m (2,277 ft). The climate is maritime temperate with significant precipitation in all seasons ( Köppen climate classification : Cfb ), like most of northwest Europe. The average temperature

6912-428: The Belgian state to assume responsibility for the government of the colony, henceforth called the Belgian Congo . A Belgian commission in 1919 estimated that Congo's population was half what it was in 1879. Germany invaded Belgium in August 1914 as part of the Schlieffen Plan to attack France , and much of the Western Front fighting of World War I occurred in western parts of the country. The opening months of

7040-517: The Chinese Tangbagouan regional stage. In the British Isles, where the Hastarian and Ivorian are difficult to distinguish, the entire Tournaisian was equivalent to the Courceyan regional stage. The base of the Tournaisian (which is also the base of the Carboniferous system ) is at the first appearance of the conodont Siphonodella sulcata within the evolutionary lineage from Siphonodella praesulcata to Siphonodella sulcata . The first appearance of ammonite species Gattendorfia subinvoluta

7168-428: The Communities and Regions were created in 1980, Flemish politicians decided to merge both. Thus the Flemings just have one single institutional body of parliament and government is empowered for all except federal and specific municipal matters. The overlapping boundaries of the Regions and Communities have created two notable peculiarities: the territory of the Brussels-Capital Region (which came into existence nearly

7296-490: The County of Flanders and it became one of the richest areas in Europe. This prosperity played a role in conflicts between Flanders and the king of France . Famously, Flemish militias scored a surprise victory at the Battle of the Golden Spurs against a strong force of mounted knights in 1302, but France soon regained control of the rebellious province. In the 15th century, the Duke of Burgundy in France took control of Flanders, and from there proceeded to unite much of what

7424-548: The European Union, for which Belgium hosts major administrations and institutions, including the European Commission , the Council of the European Union and the extraordinary and committee sessions of the European Parliament . In the early 1990s, Belgium saw several large corruption scandals notably surrounding Marc Dutroux , Andre Cools , the Dioxin Affair , Agusta Scandal and the murder of Karel van Noppen . Belgium shares borders with France ( 620 km ), Germany ( 162/167 km ), Luxembourg ( 148 km ), and

7552-491: The Flemings refused to do so and succeeded progressively in making Dutch an equal language in the education system. Following World War II, Belgian politics became increasingly dominated by the autonomy of its two main linguistic communities. Intercommunal tensions rose and the constitution was amended to minimize the potential for conflict. Based on the four language areas defined in 1962–63 (the Dutch, bilingual, French and German language areas), consecutive revisions of

7680-526: The Gómez Farias Region, Tamaulipas, Mexico , which has been described as "ground-breaking" and "a classic treatise in historical biogeography". Martin applied several disciplines including ecology , botany , climatology , geology , and Pleistocene dispersal routes to examine the herpetofauna of a relatively small and largely undisturbed area, but ecologically complex, situated on the threshold of temperate – tropical (nearctic and neotropical) regions, including semiarid lowlands at 70 meters elevation and

7808-448: The Indian Ocean was much narrower than it is today, and that South America was closer to the Antarctic, one would be hard pressed to explain the presence of many "ancient" lineages of perching birds in Africa, as well as the mainly South American distribution of the suboscines . Paleobiogeography also helps constrain hypotheses on the timing of biogeographic events such as vicariance and geodispersal , and provides unique information on

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7936-400: The Latin name of a large Roman province covering most of Northern Gaul, including the Belgae and Treveri. However, areas closer to the lower Rhine frontier, including the eastern part of modern Belgium, subsequently became part of the frontier province of Germania Inferior , which continued to interact with their neighbours outside the empire. At the time when central government collapsed in

8064-458: The N-VA remained the largest party in parliament. In July 2019 prime minister Charles Michel was selected to hold the post of President of the European Council . His successor Sophie Wilmès was Belgium's first female prime minister. She led the caretaker government since October 2019. The Flemish Liberal party politician Alexander De Croo became new prime minister in October 2020. The parties had agreed on federal government 16 months after

8192-399: The Origin of Species were devoted to geographical distribution. The first discoveries that contributed to the development of biogeography as a science began in the mid-18th century, as Europeans explored the world and described the biodiversity of life. During the 18th century most views on the world were shaped around religion and for many natural theologists, the bible. Carl Linnaeus , in

8320-426: The Regions' and Communities' Governments is the broadest of all the Federating units of all the Federations all over the world. Because of its location at the crossroads of Western Europe , Belgium has historically been the route of invading armies from its larger neighbors. With virtually defenseless borders, Belgium has traditionally sought to avoid domination by the more powerful nations which surround it through

8448-403: The Roman province of Gallia Belgica in 22 BC. In the Middle Ages , the region was part of the Carolingian Empire , and much of it was later part of the Holy Roman Empire , and subsequently the Burgundian Netherlands . Belgium's central location has meant that the area has been relatively prosperous, connected commercially and politically to its bigger neighbours. The country as it exists today

8576-429: The Theory of Continental Drift in 1912, though it was not widely accepted until the 1960s. This theory was revolutionary because it changed the way that everyone thought about species and their distribution around the globe. The theory explained how continents were formerly joined in one large landmass, Pangea , and slowly drifted apart due to the movement of the plates below Earth's surface. The evidence for this theory

8704-603: The Theory of Uniformitarianism after studying fossils. This theory explained how the world was not created by one sole catastrophic event, but instead from numerous creation events and locations. Uniformitarianism also introduced the idea that the Earth was actually significantly older than was previously accepted. Using this knowledge, Lyell concluded that it was possible for species to go extinct. Since he noted that Earth's climate changes, he realized that species distribution must also change accordingly. Lyell argued that climate changes complemented vegetation changes, thus connecting

8832-537: The U.K. National Biodiversity Network , the Atlas of Living Australia , and many others. In the case of the oceans, in 2017 Costello et al. analyzed the distribution of 65,000 species of marine animals and plants as then documented in OBIS, and used the results to distinguish 30 distinct marine realms, split between continental-shelf and offshore deep-sea areas. Since it is self evident that compilations of species occurrence records cannot cover with any completeness, areas that have received either limited or no sampling,

8960-403: The Viséan, and river systems of the Tournaisian were more similar to those of the Late Devonian . The Tournaisian saw a new diversification of arborescent (tree-sized) lycophytes and giant sphenophytes (horsetails). They coexisted alongside ferns and lignophytes ( wood -bearing plants), including early seed plants such as lyginopteridalean pteridosperms ("seed ferns"). The Tournaisian

9088-414: The anticipated effects of climate change can also be used to show potential changes in species distributions that may occur in the future based on such scenarios. Paleobiogeography goes one step further to include paleogeographic data and considerations of plate tectonics . Using molecular analyses and corroborated by fossils , it has been possible to demonstrate that perching birds evolved first in

9216-421: The biotic and abiotic features of the Earth in his book, Cosmos . Augustin de Candolle contributed to the field of biogeography as he observed species competition and the several differences that influenced the discovery of the diversity of life. He was a Swiss botanist and created the first Laws of Botanical Nomenclature in his work, Prodromus. He discussed plant distribution and his theories eventually had

9344-399: The cabinet to the king, as no progress in constitutional reforms had been made. In December 2008, Leterme once more offered his resignation after a crisis surrounding the sale of Fortis to BNP Paribas . At this juncture, his resignation was accepted and Christian Democratic and Flemish Herman Van Rompuy was sworn in as Prime Minister on 30 December 2008. After Herman Van Rompuy

9472-731: The civil servants. Communities exercise their authority only within linguistically determined geographical boundaries, originally oriented towards the individuals of a Community's language: culture (including audiovisual media), education and the use of the relevant language. Extensions to personal matters less directly connected with language comprise health policy (curative and preventive medicine) and assistance to individuals (protection of youth, social welfare, aid to families, immigrant assistance services, and so on.). Regions have authority in fields that can be broadly associated with their territory. These include economy, employment, agriculture, water policy, housing, public works, energy, transport,

9600-562: The country in May 1940 , and 40,690 Belgians, over half of them Jews, were killed during the subsequent occupation and the Holocaust . From September 1944 to February 1945 the Allies liberated Belgium. After World War II , a general strike forced King Leopold III to abdicate in 1951 in favour of his son, Prince Baudouin , since many Belgians thought he had collaborated with Germany during

9728-521: The country to be 160 km (62 sq mi) larger in surface area than previously thought. Its land area alone is 30,494 square kilometers. It lies between latitudes 49°30' and 51°30' N, and longitudes 2°33' and 6°24' E. Belgium has three main geographical regions; the coastal plain in the northwest and the central plateau both belong to the Anglo-Belgian Basin, and the Ardennes uplands in

9856-407: The country's constitution in 1970, 1980, 1988 and 1993 established a unique form of a federal state with segregated political power into three levels: The constitutional language areas determine the official languages in their municipalities, as well as the geographical limits of the empowered institutions for specific matters. Although this would allow for seven parliaments and governments when

9984-420: The development of the fields of conservation biology and landscape ecology . Classic biogeography has been expanded by the development of molecular systematics , creating a new discipline known as phylogeography . This development allowed scientists to test theories about the origin and dispersal of populations, such as island endemics . For example, while classic biogeographers were able to speculate about

10112-633: The elections. Following a usage which can be traced back to the Burgundian and Habsburg courts, in the 19th century it was necessary to speak French to belong to the governing upper class, and those who could only speak Dutch were effectively second-class citizens. Late that century, and continuing into the 20th century, Flemish movements evolved to counter this situation. While the people in Southern Belgium spoke French or dialects of French, and most Brusselers adopted French as their first language,

10240-675: The environment, town and country planning, nature conservation, credit and foreign trade. They supervise the provinces, municipalities and intercommunal utility companies. In several fields, the different levels each have their own say on specifics. With education, for instance, the autonomy of the Communities neither includes decisions about the compulsory aspect nor allows for setting minimum requirements for awarding qualifications, which remain federal matters. Each level of government can be involved in scientific research and international relations associated with its powers. The treaty-making power of

10368-496: The environmental surroundings to varying species. This largely influenced Charles Darwin in his development of the theory of evolution. Charles Darwin was a natural theologist who studied around the world, and most importantly in the Galapagos Islands . Darwin introduced the idea of natural selection, as he theorized against previously accepted ideas that species were static or unchanging. His contributions to biogeography and

10496-597: The factors affecting organism distribution, and to predict future trends in organism distribution. Often mathematical models and GIS are employed to solve ecological problems that have a spatial aspect to them. Biogeography is most keenly observed on the world's islands . These habitats are often much more manageable areas of study because they are more condensed than larger ecosystems on the mainland. Islands are also ideal locations because they allow scientists to look at habitats that new invasive species have only recently colonized and can observe how they disperse throughout

10624-453: The formation of regional biotas. For example, data from species-level phylogenetic and biogeographic studies tell us that the Amazonian teleost fauna accumulated in increments over a period of tens of millions of years, principally by means of allopatric speciation, and in an arena extending over most of the area of tropical South America (Albert & Reis 2011). In other words, unlike some of

10752-477: The former Lifemapper project at the University of Kansas (now continued as a part of BiotaPhy ) and AquaMaps , which as at 2023 contain modelled distributions for around 200,000 terrestrial, and 33,000 species of teleosts , marine mammals and invertebrates, respectively. One advantage of ENM/SDM is that in addition to showing current (or even past) modelled distributions, insertion of changed parameters such as

10880-767: The four components are subordinate to the Staff Department for Operations and Training of the Ministry of Defense , which is headed by the Assistant Chief of Staff Operations and Training , and to the Chief of Defense . The Belgian military consists of volunteers (conscription was abolished in 1995), and citizens of other EU states, Iceland, Norway, Switzerland, or Lichtenstein are also able to join. Belgium has troops deployed in several African countries as part of UN or EU missions, in Iraq for

11008-641: The geographical distribution of organisms around the globe. Alfred Russel Wallace studied the distribution of flora and fauna in the Amazon Basin and the Malay Archipelago in the mid-19th century. His research was essential to the further development of biogeography, and he was later nicknamed the "father of Biogeography". Wallace conducted fieldwork researching the habits, breeding and migration tendencies, and feeding behavior of thousands of species. He studied butterfly and bird distributions in comparison to

11136-535: The government, and on 26 April 2010 King Albert officially accepted the resignation. The Parliamentary elections in Belgium on 13 June 2010 saw the Flemish nationalist N-VA become the largest party in Flanders, and the Socialist Party PS the largest party in Wallonia. Until December 2011, Belgium was governed by Leterme's caretaker government awaiting the end of the deadlocked negotiations for formation of

11264-500: The hamlet of Fontès . The top of the Tournaisian (the base of the Viséan) is at the first appearance of the fusulinid species Eoparastaffella simplex ( morphotype 1/morphotype 2). The Tournaisian contains eight conodont biozones : The Tournaisian coincides with Romer's gap , a period of remarkably few terrestrial fossils, thus constituting a discontinuity between the Devonian and

11392-411: The importance of environmental and geographic similarities or differences as a result of climate and other pressures on the planet. Importantly, late in his career Wegener recognised that testing his theory required measurement of continental movement rather than inference from fossils species distributions. In 1958 paleontologist Paul S. Martin published A Biogeography of Reptiles and Amphibians in

11520-554: The incumbent coalition (composed of Flemish and French-speaking Social Democrats, Liberals, and Christian Democrats) maintains a solid majority in Parliament and in all electoral constituencies. On 22 July 2014, King Philippe nominated Charles Michel (MR) and Kris Peeters (CD&V) to lead the formation of a new federal cabinet composed of the Flemish parties N-VA, CD&V, Open Vld and the French-speaking MR, which resulted in

11648-518: The inhabitants of the northernmost part of Gaul . They lived in a region stretching from Paris to the Rhine , which is much bigger than modern Belgium. However, he also specifically used the Latin word " Belgium " to refer to a politically dominant part of that region, which is now in northernmost France. In contrast, modern Belgium, together with neighbouring parts of the Netherlands and Germany, corresponds to

11776-463: The installation of Leopold I as king on 21 July 1831, now celebrated as Belgium's National Day , Belgium has been a constitutional monarchy and parliamentary democracy , with a laicist constitution based on the Napoleonic code . Although the franchise was initially restricted, universal suffrage for men was introduced after the general strike of 1893 (with plural voting until 1919) and for women in 1949. The main political parties of

11904-467: The invasion of Iraq . It is the only country that does not have age restrictions on euthanasia. Verhofstadt's coalition fared badly in the June 2007 elections . For more than a year, the country experienced a political crisis . This crisis was such that many observers speculated on a possible partition of Belgium . From 21 December 2007 until 20 March 2008 the temporary Verhofstadt III Government

12032-556: The island and change it. They can then apply their understanding to similar but more complex mainland habitats. Islands are very diverse in their biomes , ranging from the tropical to arctic climates. This diversity in habitat allows for a wide range of species study in different parts of the world. One scientist who recognized the importance of these geographic locations was Charles Darwin , who remarked in his journal "The Zoology of Archipelagoes will be well worth examination". Two chapters in On

12160-607: The lands of the most northerly Belgae – the Morini , Menapii , Nervii , Germani Cisrhenani , and Aduatuci . Caesar found these peoples particularly warlike and economically undeveloped, and described them as kinsmen of the Germanic tribes east of the Rhine. The area around Arlon in southern Belgium was a part of the country of the powerful Treveri , to whom some of them paid tribute. After Caesar's conquests, Gallia Belgica first came to be

12288-547: The lycophyte Sublepidodendron characterizes Tournaisian China. Late Devonian seed plants like Rhacopteris also persisted into the Tournaisian tropics. Lepidodendron , a massive arborescent lycophyte which would dominate coal forests through the rest of the Carboniferous, first appeared near the Tournaisian-Viséan boundary. Northern Asia ( Kazakhstan and Siberia ) was positioned within subtropical or temperate northern latitudes, and developed its own endemic floras,

12416-479: The mid-18th century, improved our classifications of organisms through the exploration of undiscovered territories by his students and disciples. When he noticed that species were not as perpetual as he believed, he developed the Mountain Explanation to explain the distribution of biodiversity; when Noah's ark landed on Mount Ararat and the waters receded, the animals dispersed throughout different elevations on

12544-799: The mid-late Tournaisian, briefly regaining a level of diversity not seen since the Middle Devonian . Almost all new species belonged to the recently-evolved family Phillipsiidae , while the few surviving Devonian-type trilobites declined. Most early Tournaisian trilobites were widespread deep-water species. By the late Tournaisian, they had recolonized shallower environments and divided into three different biogeographic zones corresponding to North America, Europe, and East Asia. 43°33′20″N 3°21′26″E  /  43.5556°N 3.3572°E  / 43.5556; 3.3572 Belgium – in Europe  (light green & dark grey) – in

12672-507: The more modern terrestrial ecosystems of the Carboniferous. The middle of the Tournaisian is marked by a southern glaciation event, of a slightly lesser extent than the glaciations which swept over Gondwana in the later Carboniferous and the very end of the Devonian. During the Tournaisian, South America was located at south polar latitudes and formed the westernmost part of the supercontinent Gondwana . The southwestern coastline of Gondwana

12800-515: The most important and consequential developments in biogeography has been to show how multiple organisms, including mammals like monkeys and reptiles like squamates , overcame barriers such as large oceans that many biogeographers formerly believed were impossible to cross. See also Oceanic dispersal . Biogeography now incorporates many different fields including but not limited to physical geography, geology, botany and plant biology, zoology, general biology, and modelling. A biogeographer's main focus

12928-499: The mountain. This showed different species in different climates proving species were not constant. Linnaeus' findings set a basis for ecological biogeography. Through his strong beliefs in Christianity, he was inspired to classify the living world, which then gave way to additional accounts of secular views on geographical distribution. He argued that the structure of an animal was very closely related to its physical surroundings. This

13056-683: The nature of the models employed (including the scales for which data are available), maps generated from such models may then provide better representations of the "real" biogeographic distributions of either individual species, groups of species, or biodiversity as a whole, however it should also be borne in mind that historic or recent human activities (such as hunting of great whales , or other human-induced exterminations) may have altered present-day species distributions from their potential "full" ecological footprint. Examples of predictive maps produced by niche modelling methods based on either GBIF (terrestrial) or OBIS (marine, plus some freshwater) data are

13184-476: The northernmost cloud forest in the western hemisphere at over 2200 meters. The publication of The Theory of Island Biogeography by Robert MacArthur and E.O. Wilson in 1967 showed that the species richness of an area could be predicted in terms of such factors as habitat area, immigration rate and extinction rate. This added to the long-standing interest in island biogeography . The application of island biogeography theory to habitat fragments spurred

13312-571: The origins of species in the Hawaiian Islands , phylogeography allows them to test theories of relatedness between these populations and putative source populations on various continents, notably in Asia and North America . Biogeography continues as a point of study for many life sciences and geography students worldwide, however it may be under different broader titles within institutions such as ecology or evolutionary biology. In recent years, one of

13440-635: The other Frankish kingdoms. During the 8th century, the empire of the Franks came to be ruled by the Carolingian dynasty , whose centre of power included the area which is now eastern Belgium. Over the centuries, it was divided up in many ways, but the Treaty of Verdun in 843 divided the Carolingian Empire into three kingdoms whose borders had a lasting impact on medieval political boundaries. Most of modern Belgium

13568-823: The parliament accepted a Dutch version of the Constitution . The Berlin Conference of 1885 ceded control of the Congo Free State to King Leopold II as his private possession. From around 1900 there was growing international concern for the extreme and savage treatment of the Congolese population under Leopold II , for whom the Congo was primarily a source of revenue from ivory and rubber production. Many Congolese were killed by Leopold's agents for failing to meet production quotas for ivory and rubber. In 1908, this outcry led

13696-504: The political and linguistic interests of these communities. The major parties in each community, though close to the political center, belong to three main groups: Christian Democrats , Liberals , and Social Democrats . Further notable parties came into being well after the middle of last century, mainly to represent linguistic , nationalist, or environmental interests, and recently smaller ones of some specific liberal nature. A string of Christian Democrat coalition governments from 1958

13824-572: The population, exists in the East Cantons . The Brussels-Capital Region is officially bilingual in French and Dutch, although French is the majority language and lingua franca . Belgium's linguistic diversity and related political conflicts are reflected in its complex system of governance, made up of six different governments . In antiquity the region around Belgium was inhabited by the Belgae , and became

13952-448: The presence or absence of geographical barriers. His observations led him to conclude that the number of organisms present in a community was dependent on the amount of food resources in the particular habitat. Wallace believed species were dynamic by responding to biotic and abiotic factors. He and Philip Sclater saw biogeography as a source of support for the theory of evolution as they used Darwin's conclusion to explain how biogeography

14080-420: The region of Australia or the adjacent Antarctic (which at that time lay somewhat further north and had a temperate climate). From there, they spread to the other Gondwanan continents and Southeast Asia – the part of Laurasia then closest to their origin of dispersal – in the late Paleogene , before achieving a global distribution in the early Neogene . Not knowing that at the time of dispersal,

14208-523: The southeast to the Hercynian orogenic belt . The Paris Basin reaches a small fourth area at Belgium's southernmost tip, Belgian Lorraine . The coastal plain consists mainly of sand dunes and polders . Further inland lies a smooth, slowly rising landscape irrigated by numerous waterways, with fertile valleys and the northeastern sandy plain of the Campine ( Kempen ). The thickly forested hills and plateaus of

14336-428: The study of plant and animal species in: their past and/or present living refugium habitat ; their interim living sites; and/or their survival locales. As writer David Quammen put it, "...biogeography does more than ask Which species? and Where . It also asks Why? and, what is sometimes more crucial, Why not? ." Modern biogeography often employs the use of Geographic Information Systems (GIS), to understand

14464-602: The supercontinent, such as Argentina and Australia . In the middle Tournaisian glaciation, species-poor frigid tundra developed in western Argentina. These south polar tundras hosted the oldest known seed plants in Gondwanan territories, which likely spread south across a land bridge once the Rheic Ocean closed between Laurussia and Gondwana. Tournaisian terrestrial sediments in South America are additionally characterized by

14592-416: The theory of evolution were different from those of other explorers of his time, because he developed a mechanism to describe the ways that species changed. His influential ideas include the development of theories regarding the struggle for existence and natural selection. Darwin's theories started a biological segment to biogeography and empirical studies, which enabled future scientists to develop ideas about

14720-448: The transition from a unitary to a federal arrangement between 1970 and 1993. Despite the reforms, tensions have persisted: there is particularly significant separatist sentiment among the Flemish ; language laws such as the municipalities with language facilities have been the source of much controversy; and the government formation period following the 2010 federal election set a world record at 589 days. Unemployment in Wallonia

14848-704: The war were known as the Rape of Belgium due to German excesses. Belgium assumed control of the German colonies of Ruanda-Urundi (modern-day Rwanda and Burundi ) during the war, and in 1924 the League of Nations mandated them to Belgium. In the aftermath of the First World War, Belgium annexed the Prussian districts of Eupen and Malmedy in 1925, thereby causing the presence of a German-speaking minority. German forces again invaded

14976-676: The war. The Belgian Congo gained independence in 1960 during the Congo Crisis ; Ruanda-Urundi followed with its independence two years later. Belgium joined NATO as a founding member and formed the Benelux group of nations with the Netherlands and Luxembourg. Belgium became one of the six founding members of the European Coal and Steel Community in 1951 and of the European Atomic Energy Community and European Economic Community , established in 1957. The latter has now become

15104-447: The well-known insular faunas ( Galapagos finches , Hawaiian drosophilid flies, African rift lake cichlids ), the species-rich Amazonian ichthyofauna is not the result of recent adaptive radiations . For freshwater organisms, landscapes are divided naturally into discrete drainage basins by watersheds , episodically isolated and reunited by erosional processes. In regions like the Amazon Basin (or more generally Greater Amazonia,

15232-478: Was a transitional stage for lignophyte evolution: Devonian progymnosperm taxa such as Archaeopteris had gone extinct, but new types of woody trees such as Pitus and Protopitys set the stage for even greater morphological diversity. There is still much debate over the proportion of spore-bearing (progymnosperm) to seed-bearing (spermatophyte) woody plants, but both were evidently major parts of Tournaisian ecosystems. Tropical and subtropical swamps, in what

15360-554: Was broken in 1999 after the first dioxin crisis , a major food contamination scandal. A "rainbow coalition" emerged from six parties: the Flemish and the French-speaking Liberals, Social Democrats and Greens. Later, a " purple coalition " of Liberals and Social Democrats formed after the Greens lost most of their seats in the 2003 election . The government led by Prime Minister Guy Verhofstadt from 1999 to 2007 achieved

15488-402: Was bustling with distinctive cold-water brachiopod and bivalve faunas. Coal is less common in the Tournaisian than in the rest of the Carboniferous, and forests and swamps were at low-density despite some trees reaching heights of up to 40 meters (131 feet). Anabranching channels and anastomosing rivers (with permanent channels splitting around large vegetated islands) would not develop until

15616-534: Was designated the first permanent President of the European Council on 19 November 2009, he offered the resignation of his government to King Albert II on 25 November 2009. A few hours later, the new government under Prime Minister Yves Leterme was sworn in. On 22 April 2010, Leterme again offered the resignation of his cabinet to the king after one of the coalition partners, the OpenVLD , withdrew from

15744-400: Was divided into regions which he defined as tropical, temperate, and arctic and within these regions there were similar forms of vegetation. This ultimately enabled him to create the isotherm, which allowed scientists to see patterns of life within different climates. He contributed his observations to findings of botanical geography by previous scientists, and sketched this description of both

15872-515: Was established following the 1830 Belgian Revolution , when it seceded from the United Kingdom of the Netherlands , which had incorporated the Southern Netherlands (which comprised most of modern-day Belgium) after the Congress of Vienna in 1815. Belgium has been called "the Battlefield of Europe", a reputation reinforced in the 20th century by both world wars . Belgium was an early participant in

16000-529: Was important to a George Louis Buffon's rival theory of distribution. Closely after Linnaeus, Georges-Louis Leclerc, Comte de Buffon observed shifts in climate and how species spread across the globe as a result. He was the first to see different groups of organisms in different regions of the world. Buffon saw similarities between some regions which led him to believe that at one point continents were connected and then water separated them and caused differences in species. His hypotheses were described in his work,

16128-524: Was in office. This was a coalition of the Flemish and Francophone Christian Democrats , the Flemish and Francophone Liberals together with the Francophone Social Democrats . On that day a new government , led by Flemish Christian Democrat Yves Leterme , the actual winner of the federal elections of June 2007 , was sworn in by the king. On 15 July 2008 Leterme offered the resignation of

16256-531: Was in the Middle Kingdom , later known as Lotharingia , but the coastal county of Flanders , west of the Scheldt , became the northernmost part of West Francia , the predecessor of France . In 870 in the Treaty of Meerssen , modern Belgium lands all became part of the western kingdom for a period, but in 880 in the Treaty of Ribemont , Lotharingia came under the lasting control of the eastern kingdom, which became

16384-496: Was similar to a record of species inheritance. Key findings, such as the sharp difference in fauna either side of the Wallace Line , and the sharp difference that existed between North and South America prior to their relatively recent faunal interchange , can only be understood in this light. Otherwise, the field of biogeography would be seen as a purely descriptive one. Moving on to the 20th century, Alfred Wegener introduced

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