81-562: Jadera is a genus of true bugs in the soapberry bug subfamily. Members of this genus are only found in the Americas (from southern Canada through Argentina). These 19 species belong to the genus Jadera : Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net This Coreoidea article is a stub . You can help Misplaced Pages by expanding it . Hemiptera Hemiptera ( / h ɛ ˈ m ɪ p t ər ə / ; from Ancient Greek hemipterus 'half-winged')
162-505: A partial calque of French laque en écailles , 'lac in thin pieces', later gomme-laque , 'gum lac'. Most European languages (except Romance ones and Greek) have borrowed the word for the substance from English or from the German equivalent Schellack . Shellac is scraped from the bark of the trees where the female lac bug, Kerria lacca (order Hemiptera, family Kerriidae , also known as Laccifer lacca ), secretes it to form
243-408: A pupal stage and undergo complete metamorphosis . The majority of species are terrestrial, including a number of important agricultural pests, but some are found in freshwater habitats. These include the water boatmen , backswimmers , pond skaters , and giant water bugs . Hemiptera belong to the insect superorder Paraneoptera , which includes lice ( Psocodea ), thrips ( Thysanoptera ), and
324-501: A binding agent in artist's pigments in Spain in the year 1220. The use of overall paint or varnish decoration on large pieces of furniture was first popularised in Venice (then later throughout Italy). There are a number of 13th-century references to painted or varnished cassone , often dowry cassone that were made deliberately impressive as part of dynastic marriages. The definition of varnish
405-411: A complex mix of aliphatic and alicyclic hydroxy acids and their polymers that varies in exact composition depending upon the source of the shellac and the season of collection. The major component of the aliphatic component is aleuritic acid , whereas the main alicyclic component is shellolic acid . Shellac is UV-resistant, and does not darken as it ages (though the wood under it may do so, as in
486-504: A component of heat-cured capping cement used to fasten the caps or bases to the bulbs of electric lamps. It is the central element of the traditional " French polish " method of finishing furniture, fine string instruments , and pianos . Shellac, being edible, is used as a glazing agent on pills (see excipient ) and sweets, in the form of pharmaceutical glaze (or, "confectioner's glaze"). Because of its acidic properties (resisting stomach acids), shellac-coated pills may be used for
567-673: A double layer of grains, coarser on the outside. The Amazon rain forest cicada Hemisciera maculipennis displays bright red deimatic flash coloration on its hindwings when threatened; the sudden contrast helps to startle predators, giving the cicada time to escape. The coloured patch on the hindwing is concealed at rest by an olive green patch of the same size on the forewing, enabling the insect to switch rapidly from cryptic to deimatic behaviour. Some hemipterans such as firebugs have bold aposematic warning coloration, often red and black, which appear to deter passerine birds. Many hemipterans including aphids, scale insects and especially
648-443: A greater or lesser degree. The nymphs moult several times as they grow, and each instar resembles the adult more than the previous one. Wing buds grow in later stage nymphs; the final transformation involves little more than the development of functional wings (if they are present at all) and functioning sexual organs, with no intervening pupal stage as in holometabolous insects. Many aphids are parthenogenetic during part of
729-600: A herbaceous plant. Phloem sap, which has a higher concentration of sugars and nitrogen, is under positive pressure unlike the more dilute xylem sap. Most of the Sternorrhyncha and a number of Auchenorrhynchan groups feed on phloem. Phloem feeding is common in the Fulgoromorpha, most Cicadellidae and in the Heteroptera. The Typhlocybine Cicadellids specialize in feeding on non-vascular mesophyll tissue of leaves, which
810-404: A jam jar in and out. Stridulatory sounds are produced among the aquatic Corixidae and Notonectidae (backswimmers) using tibial combs rubbed across rostral ridges. Hemipterans are hemimetabolous , meaning that they do not undergo metamorphosis , the complete change of form between a larval phase and an adult phase . Instead, their young are called nymphs , and resemble the adults to
891-593: A melting point of 75 °C (167 °F), it can be classed as a thermoplastic used to bind wood flour , the mixture can be moulded with heat and pressure. Shellac scratches more easily than most lacquers and varnishes, and application is more labour-intensive, which is why it has been replaced by plastic in most areas. Shellac is much softer than Urushi lacquer, for instance, which is far superior with regard to both chemical and mechanical resistance. But damaged shellac can easily be touched up with another coat of shellac (unlike polyurethane, which chemically cures to
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#1732852447842972-418: A moulding compound. Shellac (both orange and white varieties) was used both in the field and laboratory to glue and stabilise dinosaur bones until about the mid-1960s. While effective at the time, the long-term negative effects of shellac (being organic in nature) on dinosaur bones and other fossils is debated, and shellac is very rarely used by professional conservators and fossil preparators today. Shellac
1053-460: A protective film. Shellac as wood finish is natural and non-toxic in its pure form. A finish made of shellac is UV-resistant. For water-resistance and durability, it does not keep up with synthetic finishing products. Because it is compatible with most other finishes, shellac is also used as a barrier or primer coat on wood to prevent the bleeding of resin or pigments into the final finish, or to prevent wood stain from blotching. Shellac
1134-488: A single species. This kind of polymorphism tends to be helpful when habitats are temporary with more energy put into reproduction when food is available and into dispersal through flight when food becomes scarce. In aphids, both winged and wingless forms occur with winged forms produced in greater numbers when food resources are depleted. Aphids and whiteflies can sometimes be transported very long distances by atmospheric updrafts and high altitude winds. Wing-length polymorphism
1215-433: A small amount of wax (3%–5% by volume), which comes from the lac bug. In some preparations, this wax is removed (the resulting product being called "dewaxed shellac"). This is done for applications where the shellac will be coated with something else (such as paint or varnish), so the topcoat will adhere. Waxy (non-dewaxed) shellac appears milky in liquid form, but dries clear. Shellac comes in many warm colours, ranging from
1296-411: A solid) because the new coat merges with and bonds to the existing coat(s). Shellac is soluble in alkaline solutions of ammonia , sodium borate , sodium carbonate , and sodium hydroxide , and also in various organic solvents . When dissolved in alcohol (typically denatured ethanol ) for application, shellac yields a coating of good durability and hardness. Upon mild hydrolysis shellac gives
1377-699: A stain if the user wants the wood to be coloured; clear or blonde shellac may be applied over a stain without affecting the colour of the finished piece, as a protective topcoat. "Wax over shellac" (an application of buffed-on paste wax over several coats of shellac) is often regarded as a beautiful, if fragile, finish for hardwood floors. Luthiers still use shellac to French polish fine acoustic stringed instruments, but it has been replaced by synthetic plastic lacquers and varnishes in many workshops, especially high-volume production environments. Shellac dissolved in alcohol, typically more dilute than as used in French polish,
1458-601: A take-off velocity of up to 2.7 metres per second and an acceleration of up to 250 g. The instantaneous power output is much greater than that of normal muscle, implying that energy is stored and released to catapult the insect into the air. Cicadas, which are much larger, extend their hind legs for a jump in under a millisecond, again implying elastic storage of energy for sudden release. Instead of relying on any form of locomotion, most Sternorrhyncha females are sedentary or completely sessile, attached to their host plants by their thin feeding stylets which cannot be taken out of
1539-469: A timed enteric or colonic release. Shellac is used as a 'wax' coating on citrus fruit to prolong its shelf/storage life. It is also used to replace the natural wax of the apple , which is removed during the cleaning process. When used for this purpose, it has the food additive E number E904. Shellac is an odour and stain blocker and so is often used as the base of "all-purpose" primers. Although its durability against abrasives and many common solvents
1620-409: A tunnel-like tube as it traverses the branches of the tree. Though these tunnels are sometimes referred to as " cocoons ", they are not cocoons in the entomological sense. This insect is in the same superfamily as the insect from which cochineal is obtained. The insects suck the sap of the tree and excrete " sticklac " almost constantly. The least-coloured shellac is produced when the insects feed on
1701-409: A variety of modes of locomotion including swimming, skating on a water surface and jumping, as well as walking and flying like other insects. Several families of Heteroptera are water bugs , adapted to an aquatic lifestyle , such as the water boatmen (Corixidae), water scorpions (Nepidae), and backswimmers (Notonectidae). They are mostly predatory, and have legs adapted as paddles to help
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#17328524478421782-404: A very light blonde ("platina") to a very dark brown ("garnet"), with many varieties of brown, yellow, orange and red in between. The colour is influenced by the sap of the tree the lac bug is living on and by the time of harvest. Historically, the most commonly sold shellac is called "orange shellac", and was used extensively as a combination stain and protectant for wood panelling and cabinetry in
1863-531: A water surface, in the same way that a toy soap boat propels itself. Water bugs in the genus Microvelia ( Veliidae ) can travel at up to 17 cm/s, twice as fast as they can walk, by this means. Flight is well developed in the Hemiptera although mostly used for short distance movement and dispersal. Wing development is sometimes related to environmental conditions. In some groups of Hemiptera, there are variations of winged, short-winged, and wingless forms within
1944-451: A wide variety of habitats, generally terrestrial, though some are adapted to life in or on the surface of fresh water (e.g. pondskaters , water boatmen , giant water bugs ). Hemipterans are hemimetabolous , with young nymphs that somewhat resemble adults. Many aphids are capable of parthenogenesis , producing young from unfertilised eggs; this helps them to reproduce extremely rapidly in favourable conditions. Humans have interacted with
2025-405: Is an order of insects , commonly called true bugs , comprising over 80,000 species within groups such as the cicadas , aphids , planthoppers , leafhoppers , assassin bugs , bed bugs , and shield bugs . They range in size from 1 mm (0.04 in) to around 15 cm (6 in), and share a common arrangement of piercing-sucking mouthparts . The name "true bugs" is often limited to
2106-411: Is even lower in amino acids and contains monosaccharides rather than sucrose , as well as organic acids and minerals. No digestion is required (except for the hydrolysis of sucrose) and 90% of the nutrients in the xylem sap can be utilised. Some phloem sap feeders selectively mix phloem and xylem sap to control the osmotic potential of the liquid consumed. A striking adaptation to a very dilute diet
2187-502: Is found in many hemipterans: a filter chamber, a part of the gut looped back on itself as a countercurrent exchanger, which permits nutrients to be separated from excess water. The residue, mostly water with sugars and amino acids, is quickly excreted as sticky "honey dew", notably from aphids but also from other Auchenorrhycha and Sternorrhyncha. Some Sternorrhyncha including Psyllids and some aphids are gall formers. These sap-sucking hemipterans inject fluids containing plant hormones into
2268-637: Is more nutritious than the leaf epidermis. Most Heteroptera also feed on mesophyll tissue where they are more likely to encounter defensive secondary plant metabolites which often leads to the evolution of host specificity. Obligate xylem feeding is a special habit that is found in the Auchenorrhyncha among Cicadoidea, Cercopoidea and in Cicadelline Cicadellids. Some phloem feeders may take to xylem sap facultatively, especially when facing dehydration. Xylem feeders tend to be polyphagous; to overcome
2349-575: Is not always clear, but it seems to have been a spirit varnish based on gum benjamin or mastic , both traded around the Mediterranean. At some time, shellac began to be used as well. An article from the Journal of the American Institute of Conservation describes using infrared spectroscopy to identify shellac coating on a 16th-century cassone. This is also the period in history where "varnisher"
2430-530: Is not very good, shellac provides an excellent barrier against water vapour penetration. Shellac-based primers are an effective sealant to control odours associated with fire damage. Shellac has traditionally been used as a dye for cotton and, especially, silk cloth in Thailand, particularly in the north-eastern region. It yields a range of warm colours from pale yellow through to dark orange-reds and dark ochre. Naturally dyed silk cloth, including that using shellac,
2511-408: Is notably rare in tree-living Hemiptera. Many Auchenorrhyncha including representatives of the cicadas, leafhoppers, treehoppers, planthoppers, and froghoppers are adapted for jumping (saltation). Treehoppers, for example, jump by rapidly depressing their hind legs. Before jumping, the hind legs are raised and the femora are pressed tightly into curved indentations in the coxae. Treehoppers can attain
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2592-402: Is now commonly sold as "sanding sealer" by several companies. It is used to seal wooden surfaces, often as preparation for a final more durable finish; it reduces the amount of final coating required by reducing its absorption into the wood. Shellac is a natural bioadhesive polymer and is chemically similar to synthetic polymers. It can thus be considered a natural form of plastic . With
2673-423: Is placed in canvas tubes (much like long socks) and heated over a fire. This causes the shellac to liquefy, and it seeps out of the canvas, leaving the bark and bugs behind. The thick, sticky shellac is then dried into a flat sheet and broken into flakes, or dried into "buttons" (pucks/cakes), then bagged and sold. The end-user then crushes it into a fine powder and mixes it with ethyl alcohol before use, to dissolve
2754-424: Is their "beak" in which the modified mandibles and maxillae form a "stylet" which is sheathed within a modified labium . The stylet is capable of piercing tissues and sucking liquids, while the labium supports it. The stylet contains a channel for the outward movement of saliva and another for the inward movement of liquid food. A salivary pump drives saliva into the prey; a cibarial pump extracts liquid from
2835-418: Is true of many aquatic species which are predatory, either as nymphs or adults. The predatory shield bug for example stabs caterpillars with its beak and sucks out the body fluids. The saliva of predatory heteropterans contains digestive enzymes such as proteinase and phospholipase , and in some species also amylase . The mouthparts of these insects are adapted for predation. There are toothed stylets on
2916-454: Is used as a brush-on colorant, food glaze and wood finish . Shellac functions as a tough natural primer , sanding sealant , tannin -blocker, odour -blocker, stain , and high-gloss varnish . Shellac was once used in electrical applications as it possesses good insulation qualities and seals out moisture. Phonograph and 78 rpm gramophone records were made of shellac until they were gradually replaced by vinyl . By 1948 shellac
2997-498: Is widely available in the rural northeast, especially in Ban Khwao District , Chaiyaphum province . The Thai name for the insect and the substance is "khrang" (Thai: ครั่ง). Wood finishing is one of the most traditional and still popular uses of shellac mixed with solvents or alcohol. This dissolved shellac liquid, applied to a piece of wood, is an evaporative finish: the alcohol of the shellac mixture evaporates, leaving behind
3078-476: The Asian citrus psyllid which transmits citrus greening disease . Shellac Shellac ( / ʃ ə ˈ l æ k / ) is a resin secreted by the female lac bug on trees in the forests of India and Thailand. Chemically, it is mainly composed of aleuritic acid , jalaric acid, shellolic acid, and other natural waxes. It is processed and sold as dry flakes and dissolved in alcohol to make liquid shellac, which
3159-485: The Early Devonian period. Hemipterans can dramatically cut the mass of affected plants, especially in major outbreaks. They sometimes also change the mix of plants by predation on seeds or feeding on roots of certain species. Some sap-suckers move from one host to another at different times of year. Many aphids spend the winter as eggs on a woody host plant and the summer as parthenogenetically reproducing females on
3240-518: The Malpighian tubules in their proximal segment that produce mucopolysaccharides, which form the froth around spittlebugs, offering a measure of protection. Parental care is found in many species of Hemiptera especially in members of the Membracidae and numerous Heteroptera. In many species of shield bug, females stand guard over their egg clusters to protect them from egg parasitoids and predators. In
3321-573: The Psylloidea and Aleyrodoidea . Aphids and Coccoids appear in the Triassic . The Coleorrhyncha extend back to the Lower Jurassic . The Heteroptera first appeared in the Triassic . The present members of the order Hemiptera (sometimes referred to as Rhynchota) were historically placed into two orders, the so-called Homoptera and Heteroptera/Hemiptera, based on differences in wing structure and
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3402-519: The Sternorrhyncha and Auchenorrhyncha , or partially hardened, as in most Heteroptera . The name "Hemiptera" is from the Greek ἡμι- ( hemi ; "half") and πτερόν ( pteron ; "wing"), referring to the forewings of many heteropterans which are hardened near the base, but membranous at the ends. Wings modified in this manner are termed hemelytra (singular: hemelytron ), by analogy with
3483-482: The honeydew that these hemipterans secrete. The relationship is mutualistic , as both ant and aphid benefit. Ants such as the yellow anthill ant, Lasius flavus , breed aphids of at least four species, Geoica utricularia , Tetraneura ulmi , Forda marginata and Forda formicaria , taking eggs with them when they found a new colony; in return, these aphids are obligately associated with the ant, breeding mainly or wholly asexually inside anthills. Ants may also protect
3564-595: The lovebug is a fly and the Maybug and ladybug are beetles . The term is occasionally extended to colloquial names for freshwater or marine crustaceans (e.g. Balmain bug , Moreton Bay bug , mudbug ) and used by physicians and bacteriologists for disease-causing germs (e.g. superbugs ). Most hemipterans feed on plants, using their sucking and piercing mouthparts to extract plant sap . Some are bloodsucking, or hematophagous , while others are predators that feed on other insects or small invertebrates . They live in
3645-424: The 1930s, it was estimated that half of all shellac was used for gramophone records . Use of shellac for records was common until the 1950s and continued into the 1970s in some non-Western countries, as well as for some children's records. Until recent advances in technology, shellac ( French polish ) was the only glue used in the making of ballet dancers' pointe shoes , to stiffen the box (toe area) to support
3726-428: The 20th century. Shellac was once very common anywhere paints or varnishes were sold (such as hardware stores). However, cheaper and more abrasion- and chemical-resistant finishes, such as polyurethane , have almost completely replaced it in decorative residential wood finishing such as hardwood floors, wooden wainscoting plank panelling, and kitchen cabinets. These alternative products, however, must be applied over
3807-452: The Hemiptera for millennia. Some species, including many aphids, are significant agricultural pests , damaging crops by sucking the sap. Others harm humans more directly as vectors of serious viral diseases. The bed bug is a persistent parasite of humans, and some kissing bugs can transmit Chagas disease . Some species have been used for biological control of insect pests or of invasive plants. A few hemipterans, have been cultivated for
3888-560: The US. In the American South , use of knotty pine plank panelling covered with orange shellac was once as common in new construction as drywall is today. It was also often used on kitchen cabinets and hardwood floors, prior to the advent of polyurethane . Until the advent of vinyl , most gramophone records were pressed from shellac compounds. From 1921 to 1928, 18,000 tons of shellac were used to create 260 million records for Europe. In
3969-448: The amount (in pounds) of shellac flakes dissolved in a gallon of denatured alcohol. For example: a 1-lb. cut of shellac is the strength obtained by dissolving one pound of shellac flakes in a gallon of alcohol (equivalent to 120 grams per litre). Most pre-mixed commercial preparations come at a 3-lb. cut. Multiple thin layers of shellac produce a significantly better end result than a few thick layers. Thick layers of shellac do not adhere to
4050-414: The animal move through the water. The pondskaters or water striders (Gerridae) are also associated with water, but use the surface tension of standing water to keep them above the surface; they include the sea skaters in the genus Halobates , the only truly marine group of insects. Marangoni effect propulsion exploits the change in surface tension when a soap-like surfactant is released on to
4131-413: The aquatic Belostomatidae, females lay their eggs on the back of the male which guards the eggs. Protection provided by ants is common in the Auchenorrhyncha. Although many species of Hemiptera are significant pests of crops and garden plants, including many species of aphid and scale insects , other species are harmless. The damage done is often not so much the deprivation of the plant of its sap, but
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#17328524478424212-505: The case of pine). The earliest written evidence of shellac goes back 3,000 years, but shellac is known to have been used earlier. According to the ancient Indian epic poem, the Mahabharata , an entire palace was built out of dried shellac. Shellac was uncommonly used as a dyestuff for as long as there was a trade with the East Indies . According to Merrifield, shellac was first used as
4293-513: The completely hardened elytra of beetles , and occur only in the suborder Heteroptera . In all suborders, the hindwings – if present at all – are entirely membranous and usually shorter than the forewings. The forewings may be held "roofwise" over the body (typical of Sternorrhyncha and Auchenorrhyncha ), or held flat on the back, with the ends overlapping (typical of Heteroptera ). The antennae in Hemiptera typically consist of four or five segments, although they can still be quite long, and
4374-574: The dancer en pointe. Many manufacturers of pointe shoes still use the traditional techniques, and many dancers use shellac to revive a softening pair of shoes. Shellac was historically used as a protective coating on paintings. Sheets of Braille were coated with shellac to help protect them from wear due to being read by hand. Shellac was used from the mid-nineteenth century to produce small moulded goods such as picture frames , boxes , toilet articles, jewelry , inkwells and even dentures . Advances in plastics have rendered shellac obsolete as
4455-496: The dangerous Chagas disease . The first known hemipteran to feed in this way on vertebrates was the extinct assassin bug Triatoma dominicana found fossilized in amber and dating back about twenty million years. Faecal pellets fossilised beside it show that it transmitted a disease-causing Trypanosoma and the amber included hairs of the likely host, a bat . Some species of ant protect and farm aphids (Sternorrhyncha) and other sap-sucking hemipterans, gathering and eating
4536-818: The disparity in the analyses is suggested to be the long branch attraction effect in phylogenetic analysis, due to rapidly evolving DNA regions. The cladogram shows Hemiptera's placement within Paraneoptera , as well as how Hemiptera's four suborders are related. English names are given in parentheses where possible. Other insects [REDACTED] (barklice, booklice, lice) [REDACTED] Thysanoptera (thrips) [REDACTED] (aphids, scale insects, etc) [REDACTED] (shield bugs, assassin bugs, etc) [REDACTED] (moss bugs) [REDACTED] Fulgoromorpha (planthoppers) [REDACTED] Cicadomorpha (cicadas, leafhoppers, spittlebugs, etc) [REDACTED] The defining feature of hemipterans
4617-609: The extraction of dyestuffs such as cochineal and carmine , and for shellac . Cicadas have been used as food, and have appeared in literature since the Iliad in Ancient Greece . Hemiptera is the largest order of hemimetabolous insects (not undergoing complete metamorphosis; though some examples such as male scale insects do undergo a form of complete metamorphosis ), containing over 95,000 named species. Other insect orders with more species are all holometabolous , meaning they have
4698-429: The fact that they transmit serious viral diseases between plants. They often produce copious amounts of honeydew which encourages the growth of sooty mould . Significant pests include the cottony cushion scale , a pest of citrus fruit trees, the green peach aphid and other aphids which attack crops worldwide and transmit diseases, and jumping plant lice which can be plant-specific and transmit diseases, as with
4779-440: The flakes and make liquid shellac. Liquid shellac has a limited shelf life (about 1 year), so is sold in dry form for dissolution before use. Liquid shellac sold in hardware stores is often marked with the production (mixing) date, so the consumer can know whether the shellac inside is still good. Some manufacturers (e.g., Zinsser) have ceased labeling shellac with the production date, but the production date may be discernible from
4860-453: The heteropteran Oncopeltus fasciatus when it consumes milkweeds , while the coreid stinkbug Amorbus rubiginosus acquires 2- hexenal from its food plant, Eucalyptus . Some long-legged bugs mimic twigs, rocking to and fro to simulate the motion of a plant part in the wind. The nymph of the Masked hunter bug camouflages itself with sand grains, using its hind legs and tarsal fan to form
4941-501: The kusum tree ( Schleichera ). The number of lac bugs required to produce 1 kilogram (2.2 lb) of shellac has variously been estimated between 50,000 and 300,000. The root word lakh is a unit in the Indian numbering system for 100,000 and presumably refers to the huge numbers of insects that swarm on host trees, up to 150 per square inch (23/cm ). The raw shellac, which contains bark shavings and lac bugs removed during scraping,
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#17328524478425022-426: The life cycle, such that females can produce unfertilized eggs, which are clones of their mother. All such young are females ( thelytoky ), so 100% of the population at these times can produce more offspring. Many species of aphid are also viviparous : the young are born live rather than laid as eggs. These adaptations enable aphids to reproduce extremely rapidly when conditions are suitable. Hemipterans make use of
5103-438: The mandibles able to cut into and abrade tissues of their prey. There are further stylets on the maxillae, adapted as tubular canals to inject saliva and to extract the pre-digested and liquified contents of the prey. A few hemipterans are haematophagic ectoparasites ), feeding on the blood of larger animals. These include bedbugs and the triatomine kissing bugs of the assassin bug family Reduviidae , which can transmit
5184-582: The mouthparts of Siphonaptera , some Diptera and Thysanoptera superficially resemble the rostrum of the Hemiptera, but on closer inspection the differences are considerable. Aside from the mouthparts, various other insects can be confused with Hemiptera, but they all have biting mandibles and maxillae instead of the rostrum. Examples include cockroaches and psocids , both of which have longer, many-segmented antennae, and some beetles , but these have fully hardened forewings which do not overlap. The forewings of Hemiptera are either entirely membranous, as in
5265-471: The negative pressure of xylem requires a special cibarial pump. Phloem feeding hemiptera typically have symbiotic micro-organisms in their gut that help to convert amino acids . Phloem feeders produce honeydew from their anus. A variety of organisms that feed on honeydew form symbiotic associations with phloem-feeders. Phloem sap is a sugary liquid low in amino acids, so insects have to process large quantities to meet their nutritional requirements. Xylem sap
5346-942: The plant bugs from their natural enemies, removing the eggs of predatory beetles and preventing access by parasitic wasps. Some leafhoppers (Auchenorrhyncha) are similarly "milked" by ants. In the Corcovado rain forest of Costa Rica, wasps compete with ants to protect and milk leafhoppers; the leafhoppers preferentially give more honeydew, more often, to the wasps, which are larger and may offer better protection. Hemiptera form prey to predators including vertebrates, such as birds, and other invertebrates such as ladybirds . In response, hemipterans have evolved antipredator adaptations . Ranatra may feign death ( thanatosis ). Others such as Carpocoris purpureipennis secrete toxic fluids to ward off arthropod predators; some Pentatomidae such as Dolycoris are able to direct these fluids at an attacker. Toxic cardenolide compounds are accumulated by
5427-603: The plant quickly. Most hemipterans are phytophagous, using their sucking and piercing mouthparts to feed on plant sap. These include cicadas, leafhoppers, treehoppers, planthoppers, froghoppers, aphids, whiteflies, scale insects , and some other groups. Some are monophages , being host specific and only found on one plant taxon, others are oligophages , feeding on a few plant groups, while others again are less discriminating polyphages and feed on many species of plant. The relationship between hemipterans and plants appears to be ancient, with piercing and sucking of plants evident in
5508-454: The plant tissues inducing the production of tissue that covers to protects the insect and also act as sinks for nutrition that they feed on. The hackleberry gall psyllid for example, causes a woody gall on the leaf petioles of the hackleberry tree it infests, and the nymph of another psyllid produces a protective lerp out of hardened honeydew. Most other hemipterans are predatory , feeding on other insects, or even small vertebrates. This
5589-459: The planthoppers secrete wax to protect themselves from threats such as fungi, parasitoidal insects and predators, as well as abiotic factors like desiccation. Hard waxy coverings are especially important in the sedentary Sternorrhyncha such as scale insects, which have no means of escaping from predators; other Sternorrhyncha evade detection and attack by creating and living inside plant galls. Nymphal Cicadoidea and Cercopoidea have glands attached to
5670-440: The position of the rostrum . "Homoptera" was established as paraphyletic group and an obsolete name. The order is now divided into four suborders, Heteroptera , Sternorrhyncha , Auchenorrhyncha , and Coleorrhyncha . The earlier work was based on nuclear DNA , but later phylogenetic analysis using mitochondrial DNA suggests that Homoptera may be monophyletic after all, a sister group of Heteroptera . The cause of
5751-455: The prey. Both pumps are powered by substantial dilator muscles in the head. The beak is usually folded under the body when not in use. The diet is typically plant sap , but some hemipterans such as assassin bugs are predators . Both herbivorous and predatory hemipterans inject enzymes to begin digestion extra-orally (before the food is taken into the body). These enzymes include amylase to hydrolyse starch , polygalacturonase to weaken
5832-405: The production lot code. Alternatively, old shellac may be tested to see if it is still usable: a few drops on glass should dry to a hard surface in roughly 15 minutes. Shellac that remains tacky for a long time is no longer usable. Storage life depends on peak temperature, so refrigeration extends shelf life. The thickness (concentration) of shellac is measured by the unit "pound cut", referring to
5913-568: The suborder Heteroptera . Entomologists reserve the term bug for Hemiptera or Heteroptera, which does not include other arthropods or insects of other orders such as ants , bees , beetles , or butterflies . In some varieties of English, all terrestrial arthropods (including non-insect arachnids , and myriapods ) also fall under the colloquial understanding of bug . Many insects with "bug" in their common name , especially in American English , belong to other orders; for example,
5994-424: The substrate or to each other well, and thus can peel off with relative ease; in addition, thick shellac will obscure fine details in carved designs in wood and other substrates. Shellac naturally dries to a high-gloss sheen. For applications where a flatter (less shiny) sheen is desired, products containing amorphous silica, such as "Shellac Flat", may be added to the dissolved shellac. Shellac naturally contains
6075-401: The tarsi of the legs have two or three segments. Many hemipterans can produce sound for communication. The "song" of male cicadas, the loudest of any insect, is produced by tymbal organs on the underside of the abdomen, and is used to attract mates. The tymbals are drumlike disks of cuticle, which are clicked in and out repeatedly, making a sound in the same way as popping the metal lid of
6156-406: The tough cell walls of plants, and proteinases to break down proteins. Although the Hemiptera vary widely in their overall form, their mouthparts form a distinctive "rostrum". Other insect orders with mouthparts modified into anything like the rostrum and stylets of the Hemiptera include some Phthiraptera , but for other reasons they generally are easy to recognize as non-hemipteran. Similarly,
6237-759: The true bugs of Hemiptera. Within Paraneoptera, Hemiptera is most closely related to the sister clade Thysanoptera. The fossil record of hemipterans goes back to the Carboniferous ( Moscovian ). The oldest fossils are of the Archescytinidae from the Lower Permian and are thought to be basal to the Auchenorrhyncha . Fulgoromorpha and Cicadomorpha appear in the Upper Permian , as do Sternorrhyncha of
6318-497: The windings to the rotor or armature . In more recent times, shellac has been replaced in these applications by synthetic resins such as polyester resin . Some applications use shellac mixed with other natural or synthetic resins, such as pine resin or phenol- formaldehyde resin, of which Bakelite is the best known, for electrical use. Mixed with other resins, barium sulfate , calcium carbonate , zinc sulfide , aluminium oxide and/or cuprous carbonate ( malachite ), shellac forms
6399-513: Was identified as a distinct trade, separate from both carpenter and artist. Another use for shellac is sealing wax . The widespread use of shellac seals in Europe dates back to the 17th century , thanks to the increasing trade with India. In the early- and mid-twentieth century, orange shellac was used as a one-product finish (combination stain and varnish-like topcoat) on decorative wood panelling used on walls and ceilings in homes, particularly in
6480-481: Was no longer used to make records. From the time shellac replaced oil and wax finishes in the 19th century, it was one of the dominant wood finishes in the western world until it was largely replaced by nitrocellulose lacquer in the 1920s and 1930s. Besides wood finishing, shellac is used as an ingredient in food, medication and candy such as confectioner's glaze , as well as a means of preserving harvested citrus fruit . Shellac comes from shell and lac ,
6561-543: Was used for fixing inductor , motor , generator and transformer windings. It was applied directly to single-layer windings in an alcohol solution. For multi-layer windings, the whole coil was submerged in shellac solution, then drained and placed in a warm location to allow the alcohol to evaporate. The shellac locked the wire turns in place, provided extra insulation, prevented movement and vibration and reduced buzz and hum. In motors and generators it also helps transfer force generated by magnetic attraction and repulsion from
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