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60-481: Holes Bay is an intertidal embayment off Poole Harbour in the county of Dorset on the south coast of England. It lies mostly within the Borough of Poole and is close to Poole town centre. It is an important wetland bird haven. Holes Bay lies on the south coast of England within the Borough of Poole , apart from its northwestern shore which is part of Upton . The bay drains the heathlands around Creekmoor and

120-464: A carbohydrate energy source. The carbohydrate source can be combinations of discrete sugars or can be derived from other sources such as banana , pineapple , peach , or even tomato puree or coconut water . After the preparation of the agar medium, it is poured into test tubes or jars which are then autoclaved (or cooked in a pressure cooker) to sterilize the medium. After cooking, the medium begins to gel as it cools. The taxonomy of this family

180-401: A seta , knocking the pollinator off the flower. After pollination, the sepals and petals fade and wilt, but they usually remain attached to the ovary. In 2011, Bulbophyllum nocturnum was discovered to flower nocturnally. Some species, such as in the genera Phalaenopsis , Dendrobium , and Vanda , produce offshoots or plantlets formed from one of the nodes along the stem , through

240-474: A spur of the labellum ( 8 in the illustration above), or on the point of the sepals, or in the septa of the ovary, the most typical position amongst the Asparagales . In orchids that produce pollinia, pollination happens as some variant of the following sequence: when the pollinator enters into the flower, it touches a viscidium, which promptly sticks to its body, generally on the head or abdomen. While leaving

300-551: A velamen , has the function of absorbing humidity. It is made of dead cells and can have a silvery-grey, white or brown appearance. In some orchids, the velamen includes spongy and fibrous bodies near the passage cells, called tilosomes. The cells of the root epidermis grow at a right angle to the axis of the root to allow them to get a firm grasp on their support. Nutrients for epiphytic orchids mainly come from mineral dust, organic detritus, animal droppings and other substances collecting among on their supporting surfaces. The base of

360-473: A different species of bee, so as to enforce proper cross-pollination. A rare achlorophyllous saprophytic orchid growing entirely underground in Australia, Rhizanthella slateri , is never exposed to light, and depends on ants and other terrestrial insects to pollinate it. Catasetum , a genus discussed briefly by Darwin , actually launches its viscid pollinia with explosive force when an insect touches

420-476: A narrow strip, such as in Pacific islands that have only a narrow tidal range, or can include many meters of shoreline where shallow beach slopes interact with high tidal excursion. The peritidal zone is similar but somewhat wider, extending from above the highest tide level to below the lowest. Organisms in the intertidal zone are well-adapted to their environment, facing high levels of interspecific competition and

480-455: A semiterrestrial or rock-hugging (" lithophyte ") orchid, show a sparkling silver and gold veining on a light green background. The cordate leaves of Psychopsiella limminghei are light brownish-green with maroon-puce markings, created by flower pigments. The attractive mottle of the leaves of lady's slippers from tropical and subtropical Asia ( Paphiopedilum ), is caused by uneven distribution of chlorophyll. Also, Phalaenopsis schilleriana

540-538: A single mass. Each time pollination succeeds, thousands of ovules can be fertilized. Pollinators are often visually attracted by the shape and colours of the labellum. However, some Bulbophyllum species attract male fruit flies ( Bactrocera and Zeugodacus spp.) solely via a floral chemical which simultaneously acts as a floral reward (e.g. methyl eugenol , raspberry ketone , or zingerone ) to perform pollination. The flowers may produce attractive odours. Although absent in most species, nectar may be produced in

600-628: A sticky disc near the top of the column. Just below the pollinia is a second, larger sticky plate called the stigma . The complex mechanisms that orchids have evolved to achieve cross-pollination were investigated by Charles Darwin and described in Fertilisation of Orchids (1862). Orchids have developed highly specialized pollination systems, thus the chances of being pollinated are often scarce, so orchid flowers usually remain receptive for very long periods, rendering unpollinated flowers long-lasting in cultivation. Most orchids deliver pollen in

660-418: Is a pastel pink orchid with leaves spotted dark green and light green. The jewel orchid ( Ludisia discolor ) is grown more for its colorful leaves than its white flowers. Some orchids, such as Dendrophylax lindenii (ghost orchid), Aphyllorchis and Taeniophyllum depend on their green roots for photosynthesis and lack normally developed leaves, as do all of the heterotrophic species. Orchids of

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720-417: Is achieved by removing the pollinia with a small instrument such as a toothpick from the pollen parent and transferring them to the seed parent. Some orchids mainly or totally rely on self-pollination , especially in colder regions where pollinators are particularly rare. The caudicles may dry up if the flower has not been visited by any pollinator, and the pollinia then fall directly on the stigma. Otherwise,

780-548: Is advanced through activities including harvesting fisheries with drag nets and a neglect of the sensitivity of intertidal zones. Orchid Orchids are plants that belong to the family Orchidaceae ( / ˌ ɔːr k ɪ ˈ d eɪ s i . iː , - s i . aɪ / ), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan plants that are found in almost every habitat on Earth except glaciers . The world's richest diversity of orchid genera and species

840-526: Is also a great biodiversity. Organisms in this zone generally are not well adapted to periods of dryness and temperature extremes. Some of the organisms in this area are abalone , sea anemones , brown seaweed , chitons , crabs , green algae , hydroids , isopods , limpets , mussels , nudibranchs , sculpin , sea cucumber , sea lettuce , sea palms , starfish , sea urchins , shrimp , snails , sponges , surf grass , tube worms , and whelks . Creatures in this area can grow to larger sizes because there

900-467: Is bounded in the east by Sterte and Stanley Green , in the north by Creekmoor, in the northwest by Upton, in the west and southwest by Hamworthy and in the southeast by the Old Town area of Poole . Holes Bay is a tidal inland lake which lies to the north of Poole Harbour. It is a designated harbour quiet area. The exit to the bay is a narrow gat between Lower Hamworthy and Poole Old Town, and runs past

960-551: Is found in the tropics . Orchidaceae is one of the two largest families of flowering plants, along with the Asteraceae . It contains about 28,000 currently accepted species distributed across 763 genera . The Orchidaceae family encompasses about 6–11% of all species of seed plants . The largest genera are Bulbophyllum (2,000 species), Epidendrum (1,500 species), Dendrobium (1,400 species) and Pleurothallis (1,000 species). It also includes Vanilla (the genus of

1020-542: Is in constant flux, as new studies continue to clarify the relationships between species and groups of species, allowing more taxa at several ranks to be recognized. The Orchidaceae is currently placed in the order Asparagales by the APG III system of 2009. Five subfamilies are recognised. The cladogram below was made according to the APG system of 1998. It represents the view that most botanists had held up to that time. It

1080-525: Is intended to help reduce traffic jams by ensuring at least one bridge is open to vehicular traffic at any one time. The northern part of the bay is crossed from west to east by the South West Main Line from London to Weymouth , and Poole railway station is less than one hundred metres from the southeastern corner of the bay. The A350 hugs the eastern shore and the A35 swings past its northern shore. In

1140-441: Is more available energy in the localized ecosystem. Also, marine vegetation can grow to much greater sizes than in the other three intertidal subregions due to the better water coverage. The water is shallow enough to allow plenty of sunlight to reach the vegetation to allow substantial photosynthetic activity, and the salinity is at almost normal levels. This area is also protected from large predators such as fish because of

1200-592: Is often called a backbulb. Backbulbs still hold nutrition for the plant, but then a pseudobulb usually takes over, exploiting the last reserves accumulated in the backbulb, which eventually dies off, too. A pseudobulb typically lives for about five years. Orchids without noticeable pseudobulbs are also said to have growths, an individual component of a sympodial plant. Like most monocots , orchids generally have simple leaves with parallel veins , although some Vanilloideae have reticulate venation . Leaves may be ovate, lanceolate, or orbiculate, and very variable in size on

1260-413: Is possible to watch ecological succession over years rather than decades. The burrowing invertebrates that make up large portions of sandy beach ecosystems are known to travel relatively great distances in cross-shore directions as beaches change on the order of days, semilunar cycles, seasons, or years. The distribution of some species has been found to correlate strongly with geomorphic datums such as

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1320-560: Is the New Zealand foreshore and seabed controversy . In legal discussions, the foreshore is often referred to as the wet-sand area . For privately owned beaches in the United States , some states such as Massachusetts use the low-water mark as the dividing line between the property of the State and that of the beach owner; however the public still has fishing, fowling, and navigation rights to

1380-419: Is used as a food reserve for wintry periods, and provides for the development of the other one, from which visible growth develops. In warm and constantly humid climates, many terrestrial orchids do not need pseudobulbs. Epiphytic orchids, those that grow upon a support, have modified aerial roots that can sometimes be a few meters long. In the older parts of the roots, a modified spongy epidermis , called

1440-570: Is used for fishing. Species that occur here include bass , mullet , flounder , corkwing wrasse and gobies . Marine invertebrates such as king ragworm , clams and cockles are also numerous. The mudflats of Holes Bay were rapidly colonized by cordgrass during the 20th century, covering 63% of the intertidal zone between 1901 and 1924, before receding again between 1924 and 1980 due to erosion and die-back. Its vegetation includes woodland wild flowers, saltmarsh plants and grassland species including orchids . In 2015, Poole Borough Council ,

1500-689: The Dorset Wildlife Trust and other interested parties established the Holes Bay Nature Park to bring local people closer to nature and to ensure the habitat is managed for the benefit of the great variety of wildlife found within it. [REDACTED] Media related to Holes Bay at Wikimedia Commons 50°44′N 2°00′W  /  50.733°N 2.000°W  / 50.733; -2.000 Intertidal The intertidal zone also includes steep rocky cliffs , sandy beaches , bogs or wetlands (e.g., vast mudflats ). This area can be

1560-503: The RNLI lifeboat station and part of Poole Quay before entering Poole Harbour itself. Spanning the inlet are two bridges: Poole Bridge and the newer Twin Sails Bridge , opened in 2012. Access to Holes Bay for vessels with a draft greater than 2 metres (6.6 ft) is only possible when the bridges are lifted, which occurs at several fixed times daily and sometimes on request. The new bridge

1620-442: The sea , which is actively moved to the zone by tides. The edges of habitats, in this case the land and sea, are themselves often significant ecosystems , and the littoral zone is a prime example. A typical rocky shore can be divided into a spray zone or splash zone (also known as the supratidal zone ), which is above the spring high-tide line and is covered by water only during storms, and an intertidal zone, which lies between

1680-426: The splash zone (the region above the highest still-tide level, but which receives wave splash). On shores exposed to heavy wave action , the intertidal zone will be influenced by waves, as the spray from breaking waves will extend the intertidal zone. Depending on the substratum and topography of the shore, additional features may be noticed. On rocky shores , tide pools form in depressions that fill with water as

1740-454: The vanilla plant ), the type genus Orchis , and many commonly cultivated plants such as Phalaenopsis and Cattleya . Moreover, since the introduction of tropical species into cultivation in the 19th century, horticulturists have produced many hybrids and cultivars . Orchids are easily distinguished from other plants, as they share some very evident derived characteristics or synapomorphies . Among these are: bilateral symmetry of

1800-461: The L. 2971/01, the foreshore zone is defined as the area of the coast that might be reached by the maximum climbing of the waves on the coast (maximum wave run-up on the coast) in their maximum capacity (maximum referring to the "usually maximum winter waves" and of course not to exceptional cases, such as tsunamis ). The foreshore zone, a part of the exceptions of the law, is public, and permanent constructions are not allowed on it. In Italy, about half

1860-428: The accumulation of growth hormones at that point. These shoots are known as keiki . Epipogium aphyllum exhibits a dual reproductive strategy, engaging in both sexual and asexual seed production. The likelihood of apomixis playing a substantial role in successful reproduction appears minimal. Within certain petite orchid species groups, there is a noteworthy preparation of female gametes for fertilization preceding

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1920-499: The act of pollination. The ovary typically develops into a capsule that is dehiscent by three or six longitudinal slits, while remaining closed at both ends. The seeds are generally almost microscopic and very numerous, in some species over a million per capsule. After ripening, they blow off like dust particles or spores. Most orchid species lack endosperm in their seed and must enter symbiotic relationships with various mycorrhizal basidiomyceteous fungi that provide them

1980-541: The anther may rotate and then enter the stigma cavity of the flower (as in Holcoglossum amesianum ). The slipper orchid Paphiopedilum parishii reproduces by self-fertilization . This occurs when the anther changes from a solid to a liquid state and directly contacts the stigma surface without the aid of any pollinating agent or floral assembly. The labellum of the Cypripedioideae is poke bonnet-shaped , and has

2040-593: The bay include fishing, kayaking and small leisure craft. A large marina known as Cobbs Quay is on the west side of the bay. The bay has a small tidal range of 0.6 m (2 ft) at neap tides and 1.8 m (6 ft) at spring tides , and a double high water . Maximum monthly temperatures range from an average of 8 °C (46 °F) in January to 27 °C (81 °F) in August; minimums range from 3 °C (37 °F) to 16 °C (61 °F). Ground temperatures on

2100-505: The ecosystems, yet forty-four percent of respondents state that there is a fair amount of knowledge used in those regions for fisheries. Intertidal zones are sensitive habitats with an abundance of marine species that can experience ecological hazards associated with tourism and human-induced environmental impacts . A variety of other threats that have been summarized by scientists include nutrient pollution , overharvesting , habitat destruction , and climate change . Habitat destruction

2160-537: The flower ( zygomorphism ), many resupinate flowers, a nearly always highly modified petal (labellum), fused stamens and carpels , and extremely small seeds . All orchids are perennial herbs that lack any permanent woody structure. They can grow according to two patterns: Terrestrial orchids may be rhizomatous or form corms or tubers . The root caps of terrestrial orchids are smooth and white. Some sympodial terrestrial orchids, such as Orchis and Ophrys , have two subterranean tuberous roots . One

2220-536: The flower develops, it undergoes a twisting through 180°, called resupination , so that the labellum lies below the column . The labellum functions to attract insects, and in resupinate flowers, also acts as a landing stage, or sometimes a trap. The reproductive parts of an orchid flower are unique in that the stamens and style are joined to form a single structure, the column . Instead of being released singly, thousands of pollen grains are contained in one or two bundles called pollinia that are attached to

2280-436: The flower, it pulls the pollinium out of the anther, as it is connected to the viscidium by the caudicle or stipe. The caudicle then bends and the pollinium is moved forwards and downwards. When the pollinator enters another flower of the same species, the pollinium has taken such position that it will stick to the stigma of the second flower, just below the rostellum, pollinating it. In horticulture, artificial orchid pollination

2340-409: The flowers to gather volatile chemicals they require to synthesize pheromonal attractants. Males of such species as Euglossa imperialis or Eulaema meriana have been observed to leave their territories periodically to forage for aromatic compounds, such as cineole, to synthesize pheromone for attracting and mating with females. Each type of orchid places the pollinia on a different body part of

2400-510: The function of trapping visiting insects. The only exit leads to the anthers that deposit pollen on the visitor. In some extremely specialized orchids, such as the Eurasian genus Ophrys , the labellum is adapted to have a colour, shape, and odour which attracts male insects via mimicry of a receptive female. Pollination happens as the insect attempts to mate with flowers. Many neotropical orchids are pollinated by male orchid bees , which visit

2460-430: The genus Corallorhiza (coralroot orchids) lack leaves altogether and instead have symbiotic or parasitic associations with fungal mycelium, though which they absorb sugars. Orchid flowers have three sepals , three petals and a three-chambered ovary . The three sepals and two of the petals are often similar to each other but one petal is usually highly modified, forming a "lip" or labellum . In most orchid genera, as

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2520-435: The high and low tidal extremes. Along most shores , the intertidal zone can be clearly separated into the following subzones: high tide zone, middle tide zone, and low tide zone. The intertidal zone is one of a number of marine biomes or habitats , including estuaries , the neritic zone , the photic zone , and deep zones . Marine biologists divide the intertidal region into three zones (low, middle, and high), based on

2580-590: The high tide strand and the water table outcrop. Since the foreshore is alternately covered by the sea and exposed to the air, organisms living in this environment must be adapted to both wet and dry conditions. Intertidal zone biomass reduces the risk of shoreline erosion from high intensity waves. Typical inhabitants of the intertidal rocky shore include sea urchins , sea anemones , barnacles , chitons , crabs , isopods , mussels , starfish , and many marine gastropod molluscs such as limpets and whelks . Sexual and asexual reproduction varies by inhabitants of

2640-455: The individual plant. Their characteristics are often diagnostic. They are normally alternate on the stem, often folded lengthwise along the centre ("plicate"), and have no stipules . Orchid leaves often have siliceous bodies called stegmata in the vascular bundle sheaths (not present in the Orchidoideae ) and are fibrous. The structure of the leaves corresponds to the specific habitat of

2700-420: The intertidal zones. Humans have historically used intertidal zones as foraged food sources during low tide . Migratory birds also rely on intertidal species for feeding areas because of low water habitats consisting of an abundance of mollusks and other marine species. As with the dry sand part of a beach, legal and political disputes can arise over the ownership and use of the foreshore. One recent example

2760-401: The largest orchid in the world, Grammatophyllum speciosum (giant orchid), it can reach three meters. Some Dendrobium species have long, canelike pseudobulbs with short, rounded leaves over the whole length; some other orchids have hidden or extremely small pseudobulbs, completely included inside the leaves. With ageing the pseudobulb sheds its leaves and becomes dormant. At this stage it

2820-407: The littoral zone. With the intertidal zone's high exposure to sunlight , the temperature can range from very hot with full sunshine to near freezing in colder climates. Some microclimates in the littoral zone are moderated by local features and larger plants such as mangroves . Adaptations in the littoral zone allow the utilization of nutrients supplied in high volume on a regular basis from

2880-399: The mudflats can fall below freezing in winter. The prevailing winds are from the west or southwest, but sea breezes can also blow in from the south and southeast. Holes Bay is home to numerous wetland bird species including avocet , black-tailed godwit , curlew , kingfisher , little egret , oystercatcher , red-breasted merganser , redshank , spoonbill , teal and widgeon . The bay

2940-418: The necessary nutrients to germinate, so almost all orchid species are mycoheterotrophic during germination and reliant upon fungi to complete their lifecycles. Only a handful of orchid species have seed that can germinate without mycorrhiza , namely the species within the genus Disa with hydrochorous seeds. As the chance for a seed to meet a suitable fungus is very small, only a minute fraction of all

3000-541: The northwest is Upton Country Park and Upton House, a public facility owned by the Borough of Poole. There is a marina on the western shore, south of the railway line. There is a wooded islet, Pergins Island in the north of the bay which is not open to the public. Holes Bay is the location of the Royal National Lifeboat Institution (RNLI) training school, attached to their Poole headquarters. Uses of

3060-439: The overall average exposure of the zone. The low intertidal zone, which borders on the shallow subtidal zone, is only exposed to air at the lowest of low tides and is primarily marine in character. The mid intertidal zone is regularly exposed and submerged by average tides. The high intertidal zone is only covered by the highest of the high tides, and spends much of its time as terrestrial habitat. The high intertidal zone borders on

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3120-658: The plant. Species that typically bask in sunlight, or grow on sites which can be occasionally very dry, have thick, leathery leaves and the laminae are covered by a waxy cuticle to retain their necessary water supply. Shade-loving species, on the other hand, have long, thin leaves. The leaves of most orchids are perennial, that is, they live for several years, while others, especially those with plicate leaves as in Catasetum , shed them annually and develop new leaves together with new pseudobulbs. The leaves of some orchids are considered ornamental. The leaves of Macodes sanderiana ,

3180-420: The rapidly changing conditions that come with the tides . The intertidal zone is also home to several species from many different phyla ( Porifera , Annelida , Coelenterata , Mollusca , Arthropoda , etc.). The water that comes with the tides can vary from brackish waters , fresh with rain , to highly saline and dry salt , with drying between tidal inundations. Wave splash can dislodge residents from

3240-417: The seeds released grow into adult plants. In cultivation, germination typically takes weeks. Horticultural techniques have been devised for germinating orchid seeds on an artificial nutrient medium, eliminating the requirement of the fungus for germination and greatly aiding the propagation of ornamental orchids. The usual medium for the sowing of orchids in artificial conditions is agar gel combined with

3300-522: The shoreline is owned by the government but leased to private beach clubs called lidos. In the East African and West Indian Ocean, intertidal zone management is often neglected of being a priority due to there being no intent for collective economic productivity. According to workshops performing questionaries, it is stated that eighty-six percent of respondents believe mismanagement of mangrove and coastal ecosystems are due to lack of knowledge to steward

3360-419: The stem of sympodial epiphytes, or in some species essentially the entire stem, may be thickened to form a pseudobulb that contains nutrients and water for drier periods. The pseudobulb typically has a smooth surface with lengthwise grooves, and can have different shapes, often conical or oblong. Its size is very variable; in some small species of Bulbophyllum , it is no longer than two millimeters, while in

3420-420: The tide rises. Under certain conditions, such as those at Morecambe Bay , quicksand may form. This subregion is mostly submerged – it is only exposed at the point of low tide and for a longer period of time during extremely low tides. This area is teeming with life; the most notable difference between this subregion and the other three is that there is much more marine vegetation, especially seaweeds . There

3480-613: The wave action and the relatively shallow water. The intertidal region is an important model system for the study of ecology , especially on wave-swept rocky shores. The region contains a high diversity of species, and the zonation created by the tides causes species ranges to be compressed into very narrow bands. This makes it relatively simple to study species across their entire cross-shore range, something that can be extremely difficult in, for instance, terrestrial habitats that can stretch thousands of kilometres. Communities on wave-swept shores also have high turnover due to disturbance, so it

3540-625: The zone between low and high water. Other states such as California use the high-water mark. In the United Kingdom , the foreshore is generally deemed to be owned by the Crown , with exceptions for what are termed several fisheries , which can be historic deeds to title, dating back to King John 's time or earlier, and the Udal Law , which applies generally in Orkney and Shetland . In Greece , according to

3600-594: Was supported by morphological studies , but never received strong support in molecular phylogenetic studies. Apostasioideae : 2 genera and 16 species, south-eastern Asia Cypripedioideae : 5 genera and 130 species, from the temperate regions of the world, as well as tropical America and tropical Asia Vanilloideae : 15 genera and 180 species, humid tropical and subtropical regions, eastern North America Epidendroideae : more than 500 genera and more or less 20,000 species, cosmopolitan Orchidoideae : 208 genera and 3,630 species, cosmopolitan In 2015,

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