36-428: Adelges tsugae , the hemlock woolly adelgid ( / ə ˈ d ɛ l . dʒ ɪ d / ) or HWA , is an insect of the order Hemiptera (true bugs) native to East Asia . It feeds by sucking sap from hemlock and spruce trees ( Tsuga spp.; Picea spp.). In its native range, HWA is not a serious pest because populations are managed by natural predators and parasitoids and by host resistance. In eastern North America it
72-470: A 47 to 88% reduction in adelgid densities within 5 months of introduction. The beetle's lifecycle is in parallel to the lifecycle of the hemlock woolly adelgid. Both lay eggs in the spring and hatching occurs nearly simultaneously. When hatched, S. tsugae larvae are highly mobile and feed on hemlock woolly adelgid eggs and larvae. Each S. tsugae larva can effectively consume about 500 adelgid eggs or nearly 100 developing adelgid nymphs. Laricobius nigrinus
108-417: A considerable dieback of adelgids and subsequent regrowth of infested trees was observed across Connecticut. The same phenomenon was repeated after the prolonged winter of 2013–2014, in time to save numerous nearly succumbed forests. A 2009 study conducted by scientists with the U.S. Forest Service Southern Research Station suggests the hemlock woolly adelgid is killing hemlock trees faster than expected in
144-574: A healthy dark green. In the northern portion of the hemlock's range, death typically occurs 4 to 10 years after infestation. Trees that survive the direct effects of the infection are usually weakened and may die from secondary causes. The presence of HWA can be identified by its egg sacs, which resemble small tufts of cotton clinging to the underside of hemlock branches. In North America, the hemlock woolly adelgid asexually reproduces and can have two generations per year. Both generations are parthenogenetic and exclusively female. In its native Asian habitat,
180-552: A hundred species, are marine. Insects such as snow scorpionflies flourish in cold habitats including the Arctic and at high altitude. Insects such as desert locusts , ants, beetles, and termites are adapted to some of the hottest and driest environments on earth, such as the Sonoran Desert . Insects form a clade , a natural group with a common ancestor, among the arthropods . A phylogenetic analysis by Kjer et al. (2016) places
216-538: A pupa, developing through a series of increasingly adult-like nymphal stages. The higher level relationship of the insects is unclear. Fossilized insects of enormous size have been found from the Paleozoic Era, including giant dragonfly-like insects with wingspans of 55 to 70 cm (22 to 28 in). The most diverse insect groups appear to have coevolved with flowering plants . Adult insects typically move about by walking and flying; some can swim. Insects are
252-427: A relative of L. nigrinus . They have shown promise in field trials. Other natural predators of adelgids include aphid fly larvae (family Chamaemyiidae), certain midge larvae, green lacewings (family Chrysopidae) and brown lacewings (family Hemerobiidae). The environmentally safest chemical control methods for treating individual trees are nontoxic insecticidal soap and horticultural oil . These are sprayed on
288-418: A system of paired openings along their sides, connected to small tubes that take air directly to the tissues. The blood therefore does not carry oxygen; it is only partly contained in vessels, and some circulates in an open hemocoel . Insect vision is mainly through their compound eyes , with additional small ocelli . Many insects can hear, using tympanal organs , which may be on the legs or other parts of
324-470: A third winged generation called sexupera occurs; although this generation's sexual reproduction requires a species of spruce not found in the Eastern United States, and therefore dies, Between 100 and 300 eggs are laid by each individual in the woolly egg sacs beneath the branches. Larvae emerge in spring and can spread on their own or with the assistance of wind, birds, or mammals. In the nymph stage,
360-525: A variety of ways. Male moths can sense the pheromones of female moths over great distances. Other species communicate with sounds: crickets stridulate , or rub their wings together, to attract a mate and repel other males. Lampyrid beetles communicate with light. Humans regard many insects as pests , especially those that damage crops, and attempt to control them using insecticides and other techniques. Others are parasitic , and may act as vectors of diseases . Insect pollinators are essential to
396-498: Is imidacloprid , which can be effective for several years if absorbed through the soil. Tree roots absorb and transport the product into the foliage and kill hemlock woolly adelgid. Soil drenches must be applied when soil moisture is adequate for the tree roots to absorb the product. These products should not be used in close proximity to bodies of water. Trunk injections are used for large trees that are near water or where soils are too rocky for soil injections or drenches. The chemical
SECTION 10
#1733115954148432-507: Is a destructive pest that threatens the eastern hemlock ( Tsuga canadensis ) and the Carolina hemlock ( Tsuga caroliniana ). HWA is also found in western North America, where it has likely been present for thousands of years. In western North America, it primarily attacks western hemlock Tsuga heterophylla and has only caused minor damage due to natural predators and host resistance. Accidentally introduced to North America from Japan , HWA
468-414: Is a native fish species to the eastern United States and is known to prefer the cool, shaded streams of hemlock forests during spawning events. Vulnerable animal populations are expected to diminish as a result of loss of hemlock habitat to the invasive hemlock woolly adelgid. One factor giving hope is that the adelgid does not seem able to survive prolonged or bitter cold. Following the winter of 1999–2000,
504-520: Is affected by the presence of Rhododendron maximum , which often coexists with hemlock, because a combination of influences restricts regeneration to shade and otherwise understory-tolerant plant species. Major changes in ecosystem structure and function, including hydrologic processes, are expected with the loss of hemlock. Loss of the eastern and Carolina hemlock from hemlock woolly adelgid infestation will likely result in many ecological shifts in eastern North America. The understory of hemlock forests
540-478: Is also valued both as an ornamental and as an important source of lumber . Unlike the balsam woolly adelgid that attacked only mature balsam fir, HWA infests hemlocks of all ages. Where hemlocks occur in pure stands in that region, the most commonly observed tree species to succeed it is black (sweet) birch . In the southern extreme of its range, hemlock typically occurs not in pure stands, but in linear riparian areas and other moist sites. Succession in these areas
576-431: Is another predatory beetle used as a biological control in response to hemlock woolly adelgid. Native to the western United States and Canada, L. nigrinus is known to prey exclusively on various woolly adelgids. L. nigrinus adults lay their eggs on top of wintering adelgid larvae in early spring, and upon hatching, the larval beetles feed on hemlock woolly adelgid. Also under study is Laricobius osakensis from Japan,
612-403: Is characterized as dark, damp, and cool and is an ideal habitat for various other organisms. The moist environment is preferred by many native amphibian species, particularly newts and salamanders. Some species of birds have close association with the hemlock, especially during mating and nesting periods. Aquatic systems adjunct to hemlock stands are also affected by the trees' decline. Brook trout
648-550: Is injected directly into the tree and transported to the twigs and needles where the hemlock woolly adelgids are feeding. Adequate soil moisture is also necessary for the tree to take up these products. Hemlock is a vital component of the New England forest system, and is the third-most prevalent tree in Vermont . It provides protection from erosion along stream banks, food for deer and wildlife, and shelter for deer in winter. The tree
684-476: Is oval in shape. The tiny brown-colored insect has four thread-like stylets that are bundled together and function as a mouthpart. Three times the length of its body, the stylet bundle pierces the host plant's parenchymatic ray tissue to derive nutrition from stored reserves. It may also inject a toxin while feeding. The resulting desiccation causes the tree to lose needles and not produce new growth. Hemlocks stricken by HWA frequently become grayish-green rather than
720-830: The Latin word insectum from in , "cut up", as insects appear to be cut into three parts. The Latin word was introduced by Pliny the Elder who calqued the Ancient Greek word ἔντομον éntomon "insect" (as in entomology ) from ἔντομος éntomos "cut in pieces"; this was Aristotle 's term for this class of life in his biology , also in reference to their notched bodies. The English word insect first appears in 1601 in Philemon Holland 's translation of Pliny. In common speech, insects and other terrestrial arthropods are often called bugs . Entomologists to some extent reserve
756-2227: The Paraneoptera , and Kjer et al. 2016 for the Holometabola . The numbers of described extant species (boldface for groups with over 100,000 species) are from Stork 2018. Archaeognatha (hump-backed/jumping bristletails, 513 spp) [REDACTED] Zygentoma (silverfish, firebrats, fishmoths, 560 spp) [REDACTED] Odonata (dragonflies and damselflies, 5,899 spp) [REDACTED] Ephemeroptera (mayflies, 3,240 spp) [REDACTED] Zoraptera (angel insects, 37 spp) [REDACTED] Dermaptera (earwigs, 1,978 spp) [REDACTED] Plecoptera (stoneflies, 3,743 spp) [REDACTED] Orthoptera (grasshoppers, crickets, katydids, 23,855 spp) [REDACTED] Grylloblattodea (ice crawlers, 34 spp) [REDACTED] Mantophasmatodea (gladiators, 15 spp) [REDACTED] Phasmatodea (stick insects, 3,014 spp) [REDACTED] Embioptera (webspinners, 463 spp) [REDACTED] Mantodea (mantises, 2,400 spp) [REDACTED] Blattodea (cockroaches and termites, 7,314 spp) [REDACTED] Psocodea (book lice, barklice and sucking lice, 11,000 spp) [REDACTED] [REDACTED] Hemiptera (true bugs, 103,590 spp) [REDACTED] Thysanoptera (thrips, 5,864 spp) [REDACTED] Hymenoptera (sawflies, wasps, bees, ants, 116,861 spp) [REDACTED] Strepsiptera (twisted-wing flies, 609 spp) [REDACTED] Coleoptera (beetles, 386,500 spp) [REDACTED] Raphidioptera (snakeflies, 254 spp) [REDACTED] Neuroptera (lacewings, 5,868 spp) [REDACTED] Megaloptera (alderflies and dobsonflies, 354 spp) [REDACTED] Lepidoptera (butterflies and moths, 157,338 spp) [REDACTED] Trichoptera (caddisflies, 14,391 spp) [REDACTED] Diptera (true flies, 155,477 spp) [REDACTED] Mecoptera (scorpionflies, 757 spp) [REDACTED] Siphonaptera (fleas, 2,075 spp) [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] Laricobius osakensis Laricobius osakensis
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#1733115954148792-561: The Pennsylvania Department of Conservation and Natural Resources 's Bureau of Forestry has released hundreds of thousands of adult S. tsugae beetles into affected hemlock forests of the eastern United States to determine its effectiveness at controlling the spread of the adelgid. From 1995 to 1997, experiments in Connecticut and Virginia found that releasing adult Sasajiscymnus tsugae beetles into infested hemlock stands resulted in
828-491: The arthropod phylum . Insects have a chitinous exoskeleton , a three-part body ( head , thorax and abdomen ), three pairs of jointed legs , compound eyes , and a pair of antennae . Insects are the most diverse group of animals, with more than a million described species ; they represent more than half of all animal species. The insect nervous system consists of a brain and a ventral nerve cord . Most insects reproduce by laying eggs . Insects breathe air through
864-618: The Hemiptera (true bugs), Lepidoptera (butterflies and moths), Diptera (true flies), Hymenoptera (wasps, ants, and bees), and Coleoptera (beetles), each with more than 100,000 described species. Insects are distributed over every continent and almost every terrestrial habitat. There are many more species in the tropics , especially in rainforests , than in temperate zones. The world's regions have received widely differing amounts of attention from entomologists. The British Isles have been thoroughly surveyed, so that Gullan and Cranston 2014 state that
900-451: The adelgid is immobile and settles on a single tree. The current leading biological control method of hemlock woolly adelgid is Sasajiscymnus tsugae , [originally called Pseudoscymnus tsugae ]. S. tsugae is a black lady beetle that is relatively host-specific, feeding only on three known aldegid species, including HWA. This beetle was discovered in 1992 while feeding on hemlock woolly adelgid in its natural range of Japan. Since 1995,
936-459: The body. Their sense of smell is via receptors, usually on the antennae and the mouthparts. Nearly all insects hatch from eggs . Insect growth is constrained by the inelastic exoskeleton, so development involves a series of molts . The immature stages often differ from the adults in structure, habit and habitat. Groups that undergo four-stage metamorphosis often have a nearly immobile pupa . Insects that undergo three-stage metamorphosis lack
972-505: The foliage and smother the insects as they dry. Most trees need to be treated on a yearly basis. Toxic systemic insecticides may be applied to the foliage and bark of a tree and can persist in killing the adelgid for up to four years after application. Caution must be used, and restraint exercised around bodies of water. Soil drenches/soil injections/bark sprays are used in larger trees that cannot be completely sprayed with insecticidal soaps or foliage insecticides. The most common insecticide
1008-592: The insects among the Hexapoda , six-legged animals with segmented bodies; their closest relatives are the Diplura (bristletails). Collembola (springtails) [REDACTED] Protura (coneheads) [REDACTED] Diplura (two-pronged bristletails) [REDACTED] Insecta (=Ectognatha) [REDACTED] The internal phylogeny is based on the works of Wipfler et al. 2019 for the Polyneoptera , Johnson et al. 2018 for
1044-402: The name "bugs" for a narrow category of " true bugs ", insects of the order Hemiptera , such as cicadas and shield bugs . Other terrestrial arthropods, such as centipedes , millipedes , woodlice , spiders , mites and scorpions , are sometimes confused with insects, since they have a jointed exoskeleton. Adult insects are the only arthropods that ever have wings, with up to two pairs on
1080-554: The only invertebrates that can achieve sustained powered flight; insect flight evolved just once. Many insects are at least partly aquatic , and have larvae with gills; in some species, the adults too are aquatic. Some species, such as water striders , can walk on the surface of water. Insects are mostly solitary, but some, such as bees , ants and termites , are social and live in large, well-organized colonies . Others, such as earwigs , provide maternal care, guarding their eggs and young. Insects can communicate with each other in
1116-504: The reported 9-year decline of some infested hemlock trees in the Northeast." In fact, as of 2007, the rate of HWA expansion was recorded as 15.6 km/year south of Pennsylvania and 8.13 km/year (or less) in the northern section of the HWA's range. Insect Insects (from Latin insectum ) are hexapod invertebrates of the class Insecta . They are the largest group within
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1152-535: The reproduction of many flowering plants and so to their ecosystems. Many insects are ecologically beneficial as predators of pest insects, while a few provide direct economic benefit. Two species in particular are economically important and were domesticated many centuries ago: silkworms for silk and honey bees for honey . Insects are consumed as food in 80% of the world's nations, by people in roughly 3000 ethnic groups. Human activities are having serious effects on insect biodiversity . The word insect comes from
1188-465: The southern Appalachians, and rapidly altering the carbon cycle of these forests. According to ScienceDaily , the pest could kill most of the region's hemlock trees within the next decade. According to the study, researchers found "hemlock woolly adelgid infestation is rapidly impacting the carbon cycle in [hemlock] tree stands", and "adelgid-infested hemlock trees in the South are declining much faster than
1224-527: The thorax. Whether winged or not, adult insects can be distinguished by their three-part body plan, with head, thorax, and abdomen; they have three pairs of legs on the thorax. Estimates of the total number of insect species vary considerably, suggesting that there are perhaps some 5.5 million insect species in existence, of which about one million have been described and named. These constitute around half of all eukaryote species, including animals , plants , and fungi . The most diverse insect orders are
1260-518: The total of around 22,500 species is probably within 5% of the actual number there; they comment that Canada's list of 30,000 described species is surely over half of the actual total. They add that the 3000 species of the American Arctic must be broadly accurate. In contrast, a large majority of the insect species of the tropics and the southern hemisphere are probably undescribed. Some 30–40,000 species inhabit freshwater ; very few insects, perhaps
1296-461: Was first found in the eastern United States near Richmond, Virginia, in 1951. The pest is now found from northern Georgia to coastal Maine and southwestern Nova Scotia as well as areas of western Michigan near the eastern Lake Michigan shoreline. As of 2015, HWA has affected 90% of the geographic range of eastern hemlock in North America. An adult individual body length is typically 0.8 mm, and
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