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127-414: The hartebeest ( / ˈ h ɑːr t ə ˌ b iː s t / ; Alcelaphus buselaphus ), also known as kongoni or kaama , is an African antelope . It is the only member of the genus Alcelaphus . Eight subspecies have been described, including two sometimes considered to be independent species. A large antelope, the hartebeest stands just over 1 m (3 ft 3 in) at the shoulder, and has

254-456: A 1999 genetic study by P. Arctander of the University of Copenhagen and colleagues, which sampled the control region of the mitochondrial DNA , found that these two formed a clade within A. buselaphus , and that recognising these as species would render A. buselaphus paraphyletic (an unnatural grouping). The same study found A. b. major to be the most divergent, having branched off before

381-426: A black tuft. The other distinctive features of the hartebeest are its long legs (often with black markings), short neck, and pointed ears. A study correlated the size of hartebeest species to habitat productivity and rainfall. The western hartebeest is the largest subspecies, and has a characteristic white line between the eyes. The red hartebeest is also large, with a black forehead and a contrasting light band between

508-449: A colourless fluid in the Lelwel hartebeest. Both sexes of all subspecies have horns, with those of females being more slender. Horns can reach lengths of 45–70 cm (18–28 in); the maximum horn length is 74.9 cm ( 29 + 1 ⁄ 2  in), recorded from a Namibian red hartebeest. The horns of the western hartebeest are thick and appear U-shaped from the front and Z-shaped from

635-414: A different shape. The horns of female bovids are believed to have evolved for defence against predators or to express territoriality, as nonterritorial females, which are able to use crypsis for predator defence, often do not have horns. Females possess horns only in half of the bovid genera, and females in these genera are heavier than those in the rest. Females use horns mainly for stabbing. In bovids,

762-578: A drastic decline due to habitat destruction , hunting, human settlement, and competition with livestock for food. Each of the eight subspecies of the hartebeest has a different conservation status. The Bubal hartebeest was declared extinct by the International Union for Conservation of Nature (IUCN) in 1994. While the populations of the red hartebeest are on the rise, those of the Tora hartebeest , already Critically Endangered , are falling. The hartebeest

889-417: A female in oestrus holds out her tail slightly to signal her receptivity, and the male tries to block the female's way. She may eventually stand still and allow the male to mount her. Copulation is brief and is often repeated, sometimes twice or more in a minute. Any intruder at this time is chased away. In large herds, females often mate with several males. Gestation is eight to nine months long, after which

1016-667: A few such as the buffalo, bushbuck, reedbuck, and grysbok are exceptions. Social activity and feeding usually peak during dawn and dusk. The bovids usually rest before dawn, during midday, and after dark. Grooming is usually by licking with the tongue. Rarely do antelopes roll in mud or dust. Wildebeest and buffalo usually wallow in mud, whereas the hartebeest and topi rub their heads and horns in mud and then smear it over their bodies. Bovids use different forms of vocal, olfactory, and tangible communication. These involve varied postures of neck, head, horns, hair, legs, and ears to convey sexual excitement, emotional state, or alarm. One such expression

1143-520: A fight is marked with a series of head movements and stances, as well as depositing droppings on dung piles. The opponents drop onto their knees and, after giving a hammer-like blow, begin wrestling, their horns interlocking. One attempts to fling the head of the other to one side to stab the neck and shoulders with his horns. Fights are rarely serious, but can be fatal if they are. Like the sassabies, hartebeest produce quiet quacking and grunting sounds. Juveniles tend to be more vocal than adults, and produce

1270-474: A final backward turn. The horns of Swayne's hartebeest are thin and shaped like parentheses, curving upward and then backward. The horns of the Tora hartebeest are particularly thin and spread out sideways, diverging more than in any other subspecies. Apart from its long face, the large chest and the sharply sloping back differentiate the hartebeest from other antelopes. The hartebeest shares several physical traits with

1397-1346: A fraction of West Africa, but harbours 322 of West Africa's fish species, with 247 being restricted to this area and 129 being restricted to even smaller ranges. The central river's fauna comprises 194 fish species, with 119 endemics and only 33 restricted to small areas. The marine diversity is greatest near the Indian Ocean shore with about 2000 species. Characteristic to African fauna are Perciformes ( Lates , tilapias , Dichistiidae , Anabantidae , Mudskippers , Parachanna , Acentrogobius , Croilia , Glossogobius , Hemichromis , Nanochromis , Oligolepis , Oreochromis , Redigobius , Sarotherodon , Stenogobius and others), Gonorhynchiformes ( Kneriidae , Phractolaemidae ), some lungfishes ( Protopterus ), many Characiformes ( Distichodontidae , Hepsetidae , Citharinidae , Alestiidae ), Osteoglossiformes ( African knifefish , Gymnarchidae , Mormyridae , Pantodontidae ), Siluriformes ( Amphiliidae , Anchariidae , Ariidae , Austroglanididae , Clariidae , Claroteidae , Malapteruridae , Mochokidae , Schilbeidae ), Osmeriformes ( Galaxiidae ), Cyprinodontiformes ( Aplocheilidae , Poeciliidae ) and Cypriniformes ( Labeobarbus , Pseudobarbus , Tanakia and others). Endemic to Africa are

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1524-447: A half years, longer than in other Alcelaphini. Often the mortality rate of male juveniles is high, as they have to face the aggression of territorial adult males and are also deprived of good forage by them. The lifespan is 12 to 15 years. Hartebeest inhabit dry savannas, open plains and wooded grasslands, often moving into more arid places after rainfall. They are more tolerant of wooded areas than other Alcelaphini, and are often found on

1651-509: A level muzzle. Fighting techniques differ amongst the bovid families and also depend on their build. While the hartebeest fight on knees, others usually fight on all fours. Gazelles of various sizes use different methods of combat. Gazelles usually box, and in serious fights may clash and fence, consisting of hard blows from short range. Ibex, goat and sheep males stand upright and clash into each other downwards. Wildebeest use powerful head butting in aggressive clashes. If horns become entangled,

1778-615: A notable affinity between the A. b. lelwel and A. b. tora divisions. The eight subspecies, including the two controversial ones, are: In 2000, a study scrutinised two major populations of the Swayne's hartebeest, from the Senkele Wildlife Sanctuary and the Nechisar National Park , for mitochondrial (D-loop) and nuclear ( microsatellite ) variability in an attempt to estimate the levels of genetic variation between

1905-465: A pale white (as in the Arabian oryx ) to black (as in the black wildebeest ). However, only the intermediate shades, such as brown and reddish brown (as in the reedbuck ), are commonly observed. In several species, females and juveniles exhibit a light-coloured coat, while those of males darken with age. As in the wildebeest , the coat may be marked with prominent or faint stripes. In some species such as

2032-487: A permanent sheath of keratin . Most bovids bear 30 to 32 teeth. Most bovids are diurnal . Social activity and feeding usually peak during dawn and dusk. Bovids typically rest before dawn, during midday, and after dark. They have various methods of social organisation and social behaviour , which are classified into solitary and gregarious behaviour. Bovids use different forms of vocal, olfactory, and tangible communication. Most species alternately feed and ruminate throughout

2159-825: A permanent sheath of keratin. Although horns occur in a single pair on almost all bovid species, there are exceptions such as the four-horned antelope and the Jacob sheep . The unique horn structure is the only unambiguous morphological feature of bovids that distinguishes them from other pecorans . A high correlation exists between horn morphology and fighting behaviour of the individual. For instance, long horns are intended for wrestling and fencing, whereas curved horns are used in ramming. Males with horns directed inwards are monogamous and solitary, while those with horns directed outwards tend to be polygynous . These results were independent of body size. Male horn development has been linked to sexual selection , Horns are small spikes in

2286-548: A possible origin of Alcelaphus in eastern Africa. Alcelaphus quickly radiated across the African savannas, replacing several previous forms (such as a relative of the hirola ). Flagstad and colleagues showed an early split in the hartebeest populations into two distinct lineages around 0.5 million years ago – one to the north and the other to the south of the equator. The northern lineage further diverged into eastern and western lineages, nearly 0.4 million years ago, most probably as

2413-417: A quacking call when alarmed or pursued. The hartebeest uses defecation as an olfactory and visual display. Herds are generally sedentary, and tend to migrate only under adverse conditions such as natural calamities. The hartebeest is the least migratory in the tribe Alcelaphini (which also includes wildebeest and sassabies), and also consumes the least amount of water and has the lowest metabolic rate among

2540-603: A result of the expanding central African rainforest belt and subsequent contraction of savanna habitats during a period of global warming. The eastern lineage gave rise to the Coke's, Swayne's, Tora and Lelwel hartebeest; and from the western lineage evolved the Bubal and western hartebeest. The southern lineage gave rise to Lichtenstein's and red hartebeest. These two taxa are phylogenetically close, having diverged only 0.2 million years ago. The study concluded that these major events throughout

2667-409: A single calf weighing about 9 kg (20 lb) is born. Births usually peak in the dry season, and take place in thickets – unlike the wildebeest, which give birth in groups on the plains. Though calves can move about on their own shortly after birth, they usually lie in the open in close proximity of their mothers. The calf is weaned at four months, but young males stay with their mothers for two and

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2794-414: A study of grass selectivity among the wildebeest, zebra , and the Coke's hartebeest, the hartebeest showed the highest selectivity. All animals preferred Themeda triandra over Pennisetum mezianum and Digitaria macroblephara . More grass species were eaten in the dry season than in the wet season. Mating in hartebeest takes place throughout the year, with one or two peaks that can be influenced by

2921-416: A study on the effect of place and sex on carcass characteristics, the average carcass weight of the male red hartebeest was 79.3 kg ( 174 + 3 ⁄ 4  lb) and that of females was 56 kg (123 lb). The meat of the animals from Qua-Qua region had the highest lipid content—1.3 g (20 gr) per 100 g ( 3 + 1 ⁄ 2  oz) of meat. Negligible differences were found in

3048-593: A subspecies of A. buselaphus . Vrba dissolved the new genus in 1997 after reconsideration. An MtDNA analysis could find no evidence to support a separate genus for Lichtenstein's hartebeest. It also showed the tribe Alcelaphini to be monophyletic , and discovered close affinity between the Alcelaphus and the sassabies (genus Damaliscus )—both genetically and morphologically. Eight subspecies are identified, of which two – A. b. caama and A. b. lichtensteinii – have been considered to be independent species. However,

3175-513: A tribe of Caprinae, in Africa date back to the late Miocene. The earliest Hippotragine fossils date back to the late Miocene, and were excavated from sites such as Lothagam and Awash Valley . The first African fossils of Reduncinae date back to 6-7 Mya. Reduncinae and Peleinae probably diverged in the mid-Miocene. All bovids have the similar basic form - a snout with a blunt end, one or more pairs of horns (generally present on males) immediately after

3302-401: A typical head-and-body length of 200 to 250 cm (79 to 98 in). The weight ranges from 100 to 200 kg (220 to 440 lb). It has a particularly elongated forehead and oddly-shaped horns , a short neck, and pointed ears. Its legs, which often have black markings, are unusually long. The coat is generally short and shiny. Coat colour varies by the subspecies, from the sandy brown of

3429-489: A vast range. South Africa has one of the highest proportions of Lycaenid butterflies (48%) for any region in the world with many species restricted in range. North Africa is in the Palaearctic region and has a different species assemblage. Genera which are species rich in Africa include Charaxes , Acraea , Colotis and Papilio , most notably Papilio antimachus and Papilio zalmoxis . The tribe Liptenini

3556-662: Is endemic to the Afrotropics and includes species rich genera such as Ornipholidotos , Liptenara , Pentila , Baliochila , Hypophytala , Teriomima , Deloneura and Mimacraea . The Miletinae are mostly African, notably Lachnocnema . Other endemic lycaenids include the genus Alaena . Endemic Nymphalidae include Euphaedra , Bebearia , Heteropsis , Precis , Pseudacraea , Bicyclus and Euxanthe . Endemic Pieridae include Pseudopontia paradoxa and Mylothris . Endemic skippers include Sarangesa and Kedestes . The highest species diversity

3683-414: Is a reddish tan. Coke's hartebeest is reddish to tawny in the upper parts, but has relatively lighter legs and rump. Lichtenstein's hartebeest is reddish brown, though the flanks are a lighter tan and the rump whitish. The Tora hartebeest is a dark reddish brown in the upper part of the body, the face, the forelegs and the rump, but the hindlegs and the underbelly are a yellowish white. The Swayne's hartebeest

3810-457: Is a rich chocolate brown with fine spots of white that are actually the white tips of its hairs. Its face is black save for the chocolate band below the eyes. The shoulders and upper part of the legs are black. Fine textured, the body hair of the hartebeest is about 25 mm (1 in) long. The hartebeest has preorbital glands (glands near the eyes) with a central duct, that secrete a dark sticky fluid in Coke's and Lichtenstein's hartebeest, and

3937-577: Is believed to have come into being in the Pliocene, but became extinct in the middle Pleistocene . Several genera of Hippotraginae are known since the Pliocene and Pleistocene. This subfamily appears to have diverged from the Alcelaphinae in the latter part of early Miocene. The Bovinae are believed to have diverged from the rest of the Bovidae in the early Miocene . The Boselaphini became extinct in Africa in

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4064-708: Is disputed, with suggestions of as many as ten and as few as two subfamilies. However, molecular, morphological and fossil evidence indicates the existence of eight distinct subfamilies: Aepycerotinae (consisting of just the impala), Alcelaphinae (bontebok, hartebeest, wildebeest and relatives), Antilopinae (several antelopes, gazelles, and relatives), Bovinae (cattle, buffaloes, bison and other antelopes), Caprinae (goats, sheep, ibex, serows and relatives), Cephalophinae (duikers), Hippotraginae (addax, oryx and relatives) and Reduncinae (reedbuck and kob antelopes). In addition, three extinct subfamilies are known: Hypsodontinae (mid- Miocene ), Oiocerinae ( Turolian ) and

4191-418: Is eight to nine months long, after which a single calf is born. Births usually peak in the dry season. The lifespan is 12 to 15 years. Inhabiting dry savannas and wooded grasslands, hartebeest often move to more arid places after rainfall. They have been reported from altitudes on Mount Kenya up to 4,000 m (13,000 ft). The hartebeest was formerly widespread in Africa, but populations have undergone

4318-462: Is especially rich in Afromontane regions, and there are some endemic families in Africa (e.g. Achatinidae , Chlamydephoridae ) but other tropical families are common too ( Charopidae , Streptaxidae , Cyclophoridae , Subulinidae , Rhytididae ). 156 tardigrade species have been found, and about 8000 species of arachnids . The African millipede Archispirostreptus gigas is one of the largest in

4445-469: Is extinct in Algeria, Egypt, Lesotho, Libya, Morocco, Somalia, and Tunisia; but has been introduced into Eswatini and Zimbabwe . It is a popular game animal due to its highly regarded meat. The vernacular name "hartebeest" may have originated from the obsolete Dutch word hertebeest , literally deer beast , based on the resemblance (to early Dutch settlers ) of the antelope to deer . The first use of

4572-549: Is home to numerous songbirds ( pipits , orioles , antpeckers , brubrus , cisticolas , nigritas , olivebacks , pytilias , green-backed twinspot , crimson-wings , seedcrackers , bluebills , firefinches , waxbills , amandavas , quailfinches , munias , weavers , tit-hylia , Amadina , Anthoscopus , Mirafra , Hypargos , Eremomela , Euschistospiza , Erythrocercus , Malimbus , Pitta , Uraeginthus , pied crow , white-necked raven , thick-billed raven , Cape crow and others). The red-billed quelea

4699-554: Is in the Democratic Republic of the Congo , home to 2,040 species 181 of which are endemic. Africa is the richest continent of freshwater fish, with about 3000 species. The East African Great Lakes ( Victoria , Malawi , and Tanganyika ) are the center of biodiversity of many fish, especially cichlids (they harbor more than two-thirds of the estimated 2000 species in the family). The West African coastal rivers region covers only

4826-487: Is in the savannas of Eastern Africa . Other bovid species also occur in Europe, Asia, and North America. Bovidae includes a number of domesticated species, including three whose use has spread worldwide, namely cattle , sheep , and goats . Dairy products , such as milk , butter , and cheese , are manufactured largely from domestic cattle. Bovids are also raised for their leather , meat , and wool . The name "Bovidae"

4953-494: Is listed under a different conservation status by the International Union for Conservation of Nature . The species as a whole is classified as Least Concern by the IUCN. The hartebeest is extinct in Algeria, Egypt, Lesotho, Libya, Morocco, Somalia, and Tunisia. Hartebeest are popular game and trophy animals as they are prominently visible and hence easy to hunt. Pictorial as well as epigraphic evidence from Egypt suggests that in

5080-711: Is restricted to Madagascar . The African list of ungulates is longer than in any other continent. The largest number of modern bovids is found in Africa ( African buffalo , duikers , impala , rhebok , Reduncinae , oryx , dik-dik , klipspringer , oribi , gerenuk , true gazelles , hartebeest , wildebeest , dibatag , eland , Tragelaphus , Hippotragus , Neotragus , Raphicerus , Damaliscus ). Other even-toed ungulates include giraffes , okapis , hippopotamuses , warthogs , giant forest hogs , red river hogs and bushpigs . Odd-toed ungulates are represented by three species of zebras , African wild ass , black and white rhinoceros . The biggest African mammal

5207-461: Is the African bush elephant , the second largest being its smaller counterpart, the African forest elephant . Four species of pangolins can be found in Africa. African fauna contains 216 species of primates . Four species of great apes ( Hominidae ) are endemic to Africa: both species of gorilla ( western gorilla , Gorilla gorilla , and eastern gorilla , Gorilla beringei ) and both species of

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5334-402: Is the flehmen response . Bovids usually stand motionless, with the head high and an intent stare, when they sense danger. Some like the impala, kudu, and eland can even leap to heights of a few feet. Bovids may roar or grunt to caution others and warn off predators. Bovids such as gazelles stot or pronk in response to predators, making high leaps on stiff legs, indicating honestly both that

5461-580: Is the most abundant bird species in the world. Of the 589 species of birds (excluding seabirds) that breed in the Palaearctic (temperate Europe and Asia), 40% spend the winter elsewhere. Of those species that leave for the winter, 98% travel south to Africa. More than 1100 mammal species live in Africa. Africa has three endemic orders of mammals, the Tubulidentata ( aardvarks ), Afrosoricida ( tenrecs and golden moles ), and Macroscelidea ( elephant shrews ). The current mammalian phylogeny recognizes

5588-718: Is the only bovid that is both solitary and not territorial. This antelope hardly displays aggression, and tends to isolate itself or form loose herds, though in a favourable habitat, several bushbuck may be found quite close to one another. Excluding the cephalophines (duikers), tragelaphines (spiral-horned antelopes) and the neotragines, most African bovids are gregarious and territorial. Males are forced to disperse on attaining sexual maturity, and must form their own territories, while females are not required to do so. Males that do not hold territories form bachelor herds. Competition takes place among males to acquire dominance, and fights tend to be more rigorous in limited rutting seasons . With

5715-1113: The Antilopinae , was later placed in its own subfamily, Pantholopinae . However, molecular and morphological analysis supports the inclusion of Pantholops in Caprinae . Below is a cladogram based on Yang et al . , 2013 and Calamari, 2021: Bovini (bison, buffalo, cattle, etc.) [REDACTED] Boselaphini (nilgai and four-horned antelope) [REDACTED] Tragelaphini (kudus, nyalas etc.) [REDACTED] Aepycerotinae (impala) [REDACTED] Nesotraginae (suni and bates's antelope) Antilopinae (gazelles, springbok, dik-dik, royal antelope, saiga, etc.) [REDACTED] Cephalophinae (duikers etc.) [REDACTED] Oreotraginae (klipspringer) Reduncinae (kobs, reedbucks, waterbucks etc.) [REDACTED] Caprinae (chamois, sheep, ibexes, goats, muskox, etc.) [REDACTED] Alcelaphinae (hartebeest, topi, wildebeest etc.) [REDACTED] Hippotraginae (sable antelopes, oryxes etc.) [REDACTED] Alternatively, all members of

5842-509: The Messinian salinity crisis (for example introduction of Murinae , immigrants from southern Asia ) During the early Tertiary , Africa was covered by a vast evergreen forest inhabited by an endemic forest fauna with many types common to southern Asia. In the Pliocene the climate became dry and most of the forest was destroyed, the forest animals taking refuge in the remaining forest islands. At

5969-583: The Nazinga Game Ranch in Burkina Faso found that the hartebeest's skull structure eased the acquisition and chewing of highly fibrous foods. The hartebeest has much lower food intake than the other members of Alcelaphini. The long thin muzzle of the hartebeest assists in feeding on leaf blades of short grasses and nibbling off leaf sheaths from grass stems. In addition to this, it can derive nutritious food even from tall senile grasses. These adaptations of

6096-527: The Upper Palaeolithic age , Egyptians hunted hartebeest and domesticated them. The hartebeest was a prominent source of meat, but its economic significance was lower than that of gazelles and other desert species. However, from the beginning of the Neolithic age, hunting became less common and consequently the remains of the hartebeest from this period in ancient Egypt, where it is now extinct, are rare. In

6223-601: The West African crocodile , two species of slender-snouted crocodile , and 1-3 species of dwarf crocodile ) are also present. There live (temporarily or permanently) more than 2600 bird species in Africa (about 1500 of them passerines ). Some 114 of them are threatened species. The Afrotropic has various endemic bird families, including ostriches (Struthionidae), mesites , sunbirds , secretary bird (Sagittariidae), guineafowl (Numididae), and mousebirds (Coliidae). Also, several families of passerines are limited to

6350-491: The addax , the coat colour can vary by the season. Scent glands and sebaceous glands are often present. Some species, such as the gemsbok , sable antelope , and Grant's gazelle , are camouflaged with strongly disruptive facial markings that conceal the highly recognisable eye. Many species, such as gazelles, may be made to look flat, and hence to blend into the background, by countershading . The outlines of many bovids are broken up with bold disruptive colouration,

6477-410: The biological family of cloven-hoofed , ruminant mammals that includes cattle , bison , buffalo , antelopes (including goat-antelopes ), sheep and goats . A member of this family is called a bovid . With 143 extant species and 300 known extinct species , the family Bovidae consists of 11 (or two) major subfamilies and thirteen major tribes. The family evolved 20 million years ago, in

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6604-514: The buzzards , harriers , Old World vultures , bateleur , Circaetus , Melierax and others. Trogons are represented by one genus ( Apaloderma ). African penguin is the only penguin species. Madagascar was once home to the now extinct elephant birds . Mauritius was once home to the now extinct endemic bird species, the being notably the Dodo bird and also the Mauritius blue pigeon . Africa

6731-399: The dental pad , that provides a surface to grip grasses and foliage. They are hypsodont and selenodont , since the molars and premolars are low- crowned and crescent-shaped cusps . The lower incisors and canines project forward. The incisors are followed by a long toothless gap, known as the diastema . The general dental formula for bovids is 0.0.2-3.3 3.1.3.3 . Most members of

6858-997: The pythonids ( Python ), typhlopids ( Typhlops ) and leptotyphlopids ( Leptotyphlops , Rhinoleptus ). Of the lizards , many species of geckos ( day geckos , Afroedura , Afrogecko , Colopus , Pachydactylus , Hemidactylus , Narudasia , Paroedura , Pristurus , Quedenfeldtia , Rhoptropus , Tropiocolotes , Uroplatus ), Cordylidae , as well as Lacertidae ( Nucras , Lacerta , Mesalina , Acanthodactylus , Pedioplanis ), Agamas , skinks , plated lizards and some monitor lizards are common. There are 12 genera and 58 species of African amphisbaenians (e.g. Chirindia , Zygaspis , Monopeltis , Dalophia ). Several genera of tortoises ( Kinixys , Pelusios , Psammobates , Geochelone , Homopus , Chersina ), turtles ( Pelomedusidae , Cyclanorbis , Cycloderma , Erymnochelys ), and 5-7 species of crocodiles (the Nile crocodile ,

6985-696: The red hartebeest have been found in Elandsfontein, Cornelia (Free State) and Florisbad in South Africa , as well as in Kabwe in Zambia . In Israel , hartebeest remains have been found in northern Negev , Shephelah , Sharon Plain and Tel Lachish . This population of the hartebeest was originally limited to the open country of the southernmost regions of the southern Levant . It was probably hunted in Egypt, which affected

7112-469: The western hartebeest to the chocolate brown of the Swayne's hartebeest . Both sexes of all subspecies have horns, with those of females being more slender. Horns can reach lengths of 45–70 cm (18–28 in). Apart from its long face, the large chest and the sharply sloping back differentiate the hartebeest from other antelopes. A conspicuous hump over the shoulders is due to the long dorsal processes of

7239-524: The Aegodontia, can be classified within the subfamily Antilopinae, with the individual subfamilies being tribes in this treatment. In the early Miocene, bovids began diverging from the cervids (deer) and giraffids . The earliest bovids, whose presence in Africa and Eurasia in the latter part of early Miocene (20 Mya ) has been ascertained, were small animals, somewhat similar to modern gazelles , and probably lived in woodland environments. Eotragus ,

7366-981: The Afrotropical amphibians, there is least concern about their conservation status; approximately 130 species are endangered, about one-fourth of which are at a critical stage. Almost all of the amphibians of Madagascar (238 species ) are endemic to that region. The West African goliath frog is the largest frog species in the world. The center of chameleon diversity is Madagascar. Snakes found in Africa include atractaspidids , elapids ( cobras , Aspidelaps , Boulengerina , Dendroaspis , Elapsoidea , Hemachatus , Homoroselaps and Paranaja ), viperines , ( Atheris , Bitis , Cerastes , Causus , Echis , Macrovipera , Montatheris , Proatheris , Vipera ), colubrids ( Dendrolycus , Dispholidus , Gonionotophis , Grayia , Hormonotus , Lamprophis , Psammophis , Leioheterodon , Madagascarophis , Poecilopholis , Dasypeltis etc.),

7493-933: The Afrotropics. These include rock-jumpers (Chaetopidae), bushshrikes (Malaconotidae), wattle-eyes , (Platysteiridae) and rockfowl (Picathartidae). Other common birds include parrots ( lovebirds , Poicephalus , Psittacus ), various cranes ( crowned cranes , blue crane , wattled crane ), storks ( marabous , Abdim's stork , saddle-billed stork ), herons ( slaty egret , black heron , goliath heron ), shoebill , bustards ( kori bustard , Neotis , Eupodotis , Lissotis ), sandgrouse ( Pterocles ), Coraciiformes ( bee-eaters , hornbills , Ceratogymna ), phasians , Congo peafowl , blue quail , harlequin quail , stone partridge , Madagascar partridge ). The woodpeckers and allies include honeyguides , African barbets , African piculet , ground woodpecker , Dendropicos and Campethera . The birds of prey include

7620-440: The Bovidae, where for example Bos , Ovis , bontebok and gemsbok have white stockings. Again, communication is the likely function. Excepting some domesticated forms, all male bovids have horns, and in many species, females, too, possess horns. The size and shape of the horns vary greatly, but the basic structure is a pair of simple bony protrusions without branches, often having a spiral, twisted, or fluted form, each covered in

7747-552: The Bovidae-Moschidae clade 27 to 28 million years ago. The following cladogram is based on the 2003 study. Tragulidae [REDACTED] Antilocapridae [REDACTED] Giraffidae [REDACTED] Cervidae [REDACTED] Moschidae [REDACTED] Bovidae [REDACTED] Molecular studies have supported monophyly in the family Bovidae (a group of organisms comprises an ancestral species and all their descendants). The number of subfamilies in Bovidae

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7874-519: The Nechisar National Park. Additionally, the Swayne hartebeest populations were compared with a large red hartebeest population, and both subspecies were found to have a high degree of genetic variation. The study advocated in situ conservation of the Swayne's hartebeest and a renewed attempt at its translocation in order to conserve genetic diversity and increase its population in both the protected areas. The diploid number of chromosomes in

8001-410: The Pliocene. By the late Miocene, around 10 Mya, the bovids rapidly diversified , leading to the creation of 70 new genera. This late Miocene radiation was partly because many bovids became adapted to more open, grassland habitats. The Aepycerotinae first appeared in the late Miocene, and no significant difference in the sizes of the primitive and modern impala has been noted. Fossils of ovibovines,

8128-440: The afternoon. Temperature is regulated through sweating in cattle, whereas goats use panting for the same. The right lung , consisting of four to five lobes , is around 1.5 times larger than the left, which has three lobes. Most bovids bear 30 to 32 teeth. While the upper incisors are absent, the upper canines are either reduced or absent. Instead of the upper incisors, bovids have a thick and tough layer of tissue, called

8255-403: The availability of food. Both males and females reach sexual maturity at one to two years of age. Reproduction varies by the subspecies and local factors. Mating takes place in the territories defended by a single male, mostly in open areas. The males may fight fiercely for dominance, following which the dominant male smells the female's genitalia, and follows her if she is in oestrus . Sometimes

8382-417: The beginning of the 21st century it was understood that the family Moschidae (musk deer) was sister to Cervidae . However, a 2003 phylogenetic study by Alexandre Hassanin (of National Museum of Natural History, France ) and colleagues, based on mitochondrial and nuclear analyses, revealed that Moschidae and Bovidae form a clade sister to Cervidae . According to the study, Cervidae diverged from

8509-679: The best known group of insects. The caterpillars of mopani moth are part of the South African cuisine. Among the numerous species of African beetles are the famous sacred scarab , the centaurus beetle , manticora tiger beetles and enormous Goliath beetles . Hotspots for butterflies include the Congolian forests and the Guinean forest-savanna mosaic . Some butterflies ( Hamanumida daedalus , Precis , Eurema ) are grassland or savannah specialists. Many of these have very large populations and

8636-515: The biggest number of termite genera of all continents, and over 1,000 termite species. Of Diptera , the number of described African species is about 17,000. Natalimyzidae, a new family of acalyptrate flies has been recently described from South Africa . Anopheles gambiae , Aedes aegypti and tsetse fly are important vectors of diseases. 1600 species of bees and 2000 species of ants among other Hymenopterans are known from Africa. There live also 3,607 species of butterflies , being

8763-568: The break-up of Gondwanaland early in the Cretaceous , but was probably connected to the mainland again in the Eocene . The first Neogene faunal interchange took place in the Middle Miocene (the introduction of Myocricetodontinae , Democricetodontinae , and Dendromurinae ). A major terrestrial faunal exchange between North Africa and Europe began at about 6.1 Ma, some 0.4 Myr before the beginning of

8890-1257: The clade Afrotheria (often viewed as a superorder ), which includes the exclusively African orders, as well as others believed to be of African origin. The East African plains are well known for their diversity of large mammals. African Eulipotyphla include the subfamilies Myosoricinae and Crocidurinae . Hedgehogs include desert hedgehogs , Atelerix and others. The rodents are represented by African bush squirrels , African ground squirrels , African striped squirrels , gerbils , cane rats , acacia rats , Nesomyidae , springhare , mole rats , dassie rats , striped grass mice , sun squirrels , thicket rats , Old World porcupines , target rats , maned rats , Deomyinae , Aethomys , Arvicanthis , Colomys , Dasymys , Dephomys , Epixerus , Grammomys , Graphiurus , Hybomys , Hylomyscus , Malacomys , Mastomys , Mus , Mylomys , Myomyscus , Oenomys , Otomys , Parotomys , Pelomys , Praomys , Rhabdomys , Stenocephalemys and many others. African rabbits and hares include riverine rabbit , Bunyoro rabbit , Cape hare , scrub hare , Ethiopian highland hare , African savanna hare , Abyssinian hare and several species of Pronolagus . Among

9017-399: The coat varies in colour according to subspecies. The western hartebeest is a pale sandy-brown, but the front of the legs are darker. The red hartebeest is a reddish-brown, with a dark face. Black markings can be observed on the chin, the back of the neck, shoulders, hips and legs; these are in sharp contrast with the broad white patches that mark its flanks and lower rump. The Lelwel hartebeest

9144-409: The complex cellulose into simpler fatty acids , which are then absorbed through the rumen wall. Bovids have a long small intestine ; the length of the small intestine in cattle is 29–49 m (95–161 ft). Body temperature fluctuates through the day; for instance, in goats the temperature can change slightly from nearly 37 °C (99 °F) in the early morning to 40 °C (104 °F) in

9271-430: The concentrations of individual fatty acids , amino acids , and minerals . The study considered hartebeest meat to be healthy, as the ratio of polyunsaturated to saturated fatty acids was 0.78, slightly more than the recommended 0.7. Fauna of Africa Whereas the earliest traces of life in fossil record of Africa date back to the earliest times, the formation of African fauna as we know it today, began with

9398-592: The day. While small bovids forage in dense and closed habitat, larger species feed on high-fiber vegetation in open grasslands. Most bovids are polygynous . Mature bovids mate at least once a year and smaller species may even mate twice. In some species, neonate bovids remain hidden for a week to two months, regularly nursed by their mothers; in other species, neonates are followers, accompanying their dams, rather than tending to remain hidden. The greatest diversities of bovids occur in Africa . The maximum concentration of species

9525-506: The duiker browse for a few hours during day or night. Feeding habits are related to body size; while small bovids forage in dense and closed habitat, larger species feed upon high-fiber vegetation in open grasslands. Subfamilies exhibit different feeding strategies. While Bovinae species graze extensively on fresh grass and diffused forage, Cephalophinae species (with the exception of Sylvicapra ) primarily consume fruits. Reduncinae and Hippotraginae species depend on unstable food sources, but

9652-492: The earliest known bovid, weighed 18 kg (40 lb) and was nearly the same in size as the Thomson's gazelle . Early in their evolutionary history, the bovids split into two main clades: Boodontia (of Eurasian origin) and Aegodontia (of African origin). This early split between Boodontia and Aegodontia has been attributed to the continental divide between these land masses. When these continents were later rejoined, this barrier

9779-419: The early Miocene . The bovids show great variation in size and pelage colouration. Except some domesticated forms , all male bovids have two or more horns , and in many species, females possess horns, too. The size and shape of the horns vary greatly, but the basic structure is always one or more pairs of simple bony protrusions without branches, often having a spiral, twisted or fluted form, each covered in

9906-510: The early Pliocene; their latest fossils were excavated in Langebaanweg (South Africa) and Lothagam (Kenya). The middle Miocene marked the spread of the bovids into China and the Indian subcontinent. According to Vrba, the radiation of the subfamily Alcelaphinae began in the latter part of middle Miocene. The Caprinae tribes probably diverged in the early middle Miocene. The Caprini emerged in

10033-717: The edge of woodlands. They have been reported from altitudes on Mount Kenya up to 4,000 m (13,000 ft). The red hartebeest is known to move across large areas, and females roam home ranges of over 1,000 km (390 sq mi), with male territories 200 km (77 sq mi) in size. Females in the Nairobi National Park (Kenya) have individual home ranges stretching over 3.7–5.5 km ( 1 + 3 ⁄ 8 – 2 + 1 ⁄ 8  sq mi), which are not particularly associated with any one female group. Average female home ranges are large enough to include 20 to 30 male territories. Each hartebeest subspecies

10160-572: The endemic fauna and partly from steppe forms that adapted themselves to forest life, while the present savanna fauna is similarly explained. The isolation in past times has resulted in the presence of closely related subspecies in widely separated regions Africa, where humans originated, shows much less evidence of loss in the Pleistocene megafaunal extinction, perhaps because co-evolution of large animals alongside early humans provided enough time for them to develop effective defenses. Its situation in

10287-723: The exception of migratory males, males generally hold the same territory throughout their lives. In the waterbuck, some male individuals, known as "satellite males", may be allowed into the territories of other males and have to wait till the owner grows old so they may acquire his territory. Lek mating , where males gather together and competitively display to potential mates, is known to exist among topis , kobs , and lechwes . The tragelaphines, cattle, sheep, and goats are gregarious and not territorial. In these species, males must gain absolute dominance over all other males, and fights are not confined to territories. Males, therefore, spend years in body growth. Most bovids are diurnal, although

10414-456: The eyes. The large Lelwel hartebeest has dark stripes on the front of its legs. Coke's hartebeest is moderately large, with a shorter forehead and longer tail in comparison to the other subspecies. Lichtenstein's hartebeest is smaller, with dark stripes on the front of the legs, as in the Lelwel hartebeest. The Swayne's hartebeest is smaller than the Tora hartebeest, but both have a shorter forehead and similar appearance. Generally short and shiny,

10541-869: The families Arthroleptidae , Astylosternidae , Heleophrynidae , Hemisotidae , Hyperoliidae , Petropedetidae , Mantellidae . Also widespread are Bufonidae ( Bufo , Churamiti , Capensibufo , Mertensophryne , Nectophryne , Nectophrynoides , Schismaderma , Stephopaedes , Werneria , Wolterstorffina ), Microhylidae ( Breviceps , Callulina , Probreviceps , Cophylinae , Dyscophus , Melanobatrachinae , Scaphiophryninae ), Rhacophoridae ( Chiromantis ), Ranidae ( Afrana , Amietia , Amnirana , Aubria , Conraua , Hildebrandtia , Lanzarana , Ptychadena , Strongylopus , Tomopterna ) and Pipidae ( Hymenochirus , Pseudhymenochirus , Xenopus ). The 2002–2004 'Global Amphibian Assessment' by IUCN , Conservation International and NatureServe revealed that for only about 50% of

10668-647: The family are herbivorous , but most duikers are omnivorous . Like other ruminants, bovids have four-chambered stomachs, which allow them to digest plant material, such as grass , that cannot be used by many other animals. Ruminants (and some others like kangaroos , rabbits , and termites ) are able to use micro-organisms living in their guts to break down cellulose by fermentation . The bovids have various methods of social organisation and social behaviour, which are classified into solitary and gregarious behaviour. Further, these types may each be divided into territorial and nonterritorial behaviour. Small bovids such as

10795-448: The fauna as a whole (it has been estimated that the African insects make up about 10-20% of the global insect species richness, and about 15% of new species descriptions come from Afrotropics ). The only endemic African insect order is Mantophasmatodea . About 875 African species of dragonflies have been recorded. The migratory locust and desert locust have been serious threats to African economies and human welfare. Africa has

10922-637: The females with their young. The females form groups of five to 12 animals, with four generations of young in the group. Females fight for dominance over the herd. Sparring between males and females is common. At three or four years of age, the males can attempt to take over a territory and its female members. A resident male defends his territory and will fight if provoked. The male marks the border of his territory through defecation. Hartebeest are remarkably alert and cautious animals with highly developed brains . Generally calm in nature, hartebeest can be ferocious when provoked. While feeding, one individual stays on

11049-512: The few Onychophorans , Peripatopsis and Opisthopatus live in Africa. Greatest diversity of freshwater mollusks is found in East African lakes. Of marine snails, less diversity is present in Atlantic coast, more in tropical Western Indian Ocean region (over 3000 species of gastropods with 81 endemic species). Cowry shells have been used as a money by native Africans. The land snail fauna

11176-454: The formidable horns are used to ward off the predator. The elevated position of the eyes enables the hartebeest to inspect its surroundings continuously even as it is grazing. The muzzle has evolved so as to derive maximum nutrition from even a frugal diet. The horns are also used during fights among males for dominance in the breeding season; the clash of the horns is loud enough that it can be heard from hundreds of metres away. The beginning of

11303-524: The genus Pan ( chimpanzee , Pan troglodytes , and bonobo , Pan paniscus ). Humans and their ancestors originated in Africa. Other primates include colobuses , baboons , geladas , vervet monkeys , guenons , macaques , mandrills , crested mangabeys , white-eyelid mangabeys , kipunji , Allen's swamp monkeys , Patas monkeys and talapoins . Lemurs and aye-aye are characteristic of Madagascar. See also Lists of mammals of Africa . Bovidae Alternate taxonomy: The Bovidae comprise

11430-501: The ground. The bovids show great variation in size: the gaur can weigh more than 1,500 kg (3,300 lb), and stand 2.2 m (87 in) high at the shoulder. The royal antelope , in sharp contrast, is only 25 cm (9.8 in) tall and weighs at most 3 kg (6.6 lb). The klipspringer , another small antelope, stands 45–60 cm (18–24 in) at the shoulder and weighs just 10–20 kg (22–44 lb). Differences occur in pelage colouration, ranging from

11557-520: The hartebeest enable the animal to feed well even in the dry season, which is usually a difficult period for grazers. For instance, in comparison with the roan antelope , the hartebeest is better at procuring and chewing the scarce regrowth of perennial grasses at times when forage is least available. These unique abilities could have allowed the hartebeest to prevail over other animals millions of years ago, leading to its successful radiation across Africa. Grasses generally comprise at least 80 per cent of

11684-412: The hartebeest is Alcelaphus buselaphus . First described by German zoologist Peter Simon Pallas in 1766, it is classified in the genus Alcelaphus and placed in the family Bovidae . In 1979, palaeontologist Elisabeth Vrba supported Sigmoceros as a separate genus for Lichtenstein's hartebeest , a kind of hartebeest, as she assumed it was related to Connochaetes (wildebeest). She had analysed

11811-459: The hartebeest is 40. Hybrids are usually reported from areas where ranges of two subspecies overlap. Hybrids between the Lelwel and Tora hartebeest have been reported in eastern Sudan and western Ethiopia, in a stretch southward from the Blue Nile to about 9° N latitude. A study proved a male hybrid of the red hartebeest and the blesbok ( Damaliscus pygargus ) to be sterile . Sterility of the hybrid

11938-453: The hartebeest's diet, but they account for over 95 per cent of their food in the wet season, October to May. Jasminum kerstingii is part of the hartebeest's diet at the start of the rainy season. Between seasons, they mainly feed on the culms of grasses. A study found that the hartebeest is able to digest a higher proportion of food than the topi and the wildebeest. In areas with scarce water, it can survive on melons, roots, and tubers. In

12065-408: The hartebeest's evolution are strongly related to climatic factors, and that there had been successive bursts of radiation from a more permanent population—a refugium —in eastern Africa; this could be vital to understanding the evolutionary history of not only the hartebeest but also other mammals of the African savanna. The earliest fossil record dates back to nearly 0.7 million years ago. Fossils of

12192-513: The head of one Zambian individual without any pathogenicity . Nematodes , cestodes , paramphistomes ; and the roundworm Setaria labiatopapillosa have also been isolated from the hartebeest. In 1931, a red hartebeest in Gobabis (southwestern Africa) was infected with long, thin worms. These were named Longistrongylus meyeri after their collector, T. Meyer. Hartebeest are primarily grazers, and their diets consist mostly of grasses. A study in

12319-482: The klipspringer, oribi , and steenbok are generally solitary and territorial. They hold small territories into which other members of the species are not allowed to enter. These antelopes form monogamous pairs. Many species such as the dik-dik use pheromone secretions from the preorbital glands and sometimes dung, as well, to mark their territories. The offspring disperse at the time of adolescence, and males must acquire territories prior to mating. The bushbuck

12446-843: The lineage split to give a combined caama/lichtensteinii lineage and another that gave rise to the remaining extant subspecies. Conversely a 2001 phylogenetic study, based on D–loop and cytochrome b analysis by Øystein Flagstad (of the Norwegian Institute for Nature Research, Trondheim ) and colleagues, found that the southern lineage of A. b. caama and A. lichtensteinii diverged earliest. Analysis of skull structure supports partition into three major divisions: A. b. buselaphus division ( nominate , also including A. b. major division), A. b. tora division (also including A. b. cokii and A. b. swaynei ) and A. b. lelwel division. Another analysis of cytochrome b and D-loop sequence data shows

12573-439: The lookout for danger, often standing on a termite mound to see farther. At times of danger, the whole herd flees in a single file after an individual suddenly starts off. Adult hartebeest are preyed upon by lions, leopards , hyenas and wild dogs ; cheetahs and jackals target juveniles . Crocodiles may also prey on hartebeest. The thin long legs of the hartebeest provide for a quick escape in an open habitat; if attacked,

12700-494: The marine mammals there are several species of dolphins , 2 species of sirenians and seals (e.g. Cape fur seals ). Of the carnivorans there are 60 species, including the conspicuous hyenas , lions , leopards , cheetahs , serval , African wild dog as well as the less prominent and understudied Side-striped jackal , striped polecat , African striped weasel , caracal , honey badger , speckle-throated otter , several mongooses , foxes and civets . The family Eupleridae

12827-594: The members of the tribe. Several parasites have been isolated from the hartebeest. These parasites regularly alternate between hartebeest and gazelles or wildebeest. Hartebeest can be infected with theileriosis due to Rhipicephalus evertsi and Theileria species. South of the Sahara , common parasites include Loewioestrus variolosus , Gedoelstia cristata and G. hassleri . The latter two species can cause serious diseases such as encephalitis . However, parasites are not always harmful – 252 larvae were found in

12954-494: The middle Miocene, and seem to have been replaced by other bovids and cervids in Eurasia. The earliest fossils of the antilopines are from the middle Miocene, though studies show the existence of the subfamily from the early Miocene. Speciation occurred in the tribe Antilopini during the middle or upper Miocene, mainly in Eurasia. Tribe Neotragini seems to have appeared in Africa by the end of Miocene, and had become widespread by

13081-431: The monogamous duikers and other small antelopes, whereas in the polygynous, they are large and elaborately formed (for example in a spiral structure, as in the giant eland ). Thus, to some extent, horns depict the degree of competition among males in a species. However, the presence of horns in females is likely due to natural selection . The horns of females are usually smaller than those of males, and are sometimes of

13208-498: The numbers in the Levant, and disconnected it from its main population in Africa. A large antelope with a particularly elongated forehead and oddly shaped horns, the hartebeest stands just over 1 m (3 ft 3 in) at the shoulder, and has a typical head-and-body length of 200 to 250 cm (79 to 98 in). The weight ranges from 100 to 200 kg (220 to 440 lb). The tail, 40 to 60 cm (16 to 24 in) long, ends in

13335-413: The opponent's head rather than its body. The S-shaped horns, such as those on the impala , have various sections that help in ramming, holding, and stabbing. Serious fights leading to injury are rare. Most bovids alternately feed and ruminate throughout the day. While those that feed on concentrate feed and digest in short intervals, the roughage feeders take longer intervals. Only small species such as

13462-429: The opponents move in a circular manner to unlock them. Muskoxen will ram into each other at high speeds. As a rule, only two bovids of equal build and level of defence engage in a fight, which is intended to determine the superior of the two. Individuals that are evidently inferior to others would rather flee than fight; for example, immature males do not fight with the mature bulls. Generally, bovids direct their attacks on

13589-442: The oval or pointed ears, a distinct neck and limbs, and a tail varying in length and bushiness among the species. Most bovids exhibit sexual dimorphism , with males usually larger as well as heavier than females. Sexual dimorphism is more prominent in medium- to large-sized bovids. All bovids have four toes on each foot – they walk on the central two (the hooves ), while the outer two (the dewclaws ) are much smaller and rarely touch

13716-478: The populations and within the subspecies. The results showed a remarkable differentiation between the two populations; that from the Senkele Wildlife Sanctuary showed more genetic diversity than the one from the Nechisar National Park. Another revelation was that the translocation of the individuals from the Senkele Wildlife Sanctuary in 1974 had not made a significant contribution to the gene pool of

13843-498: The predator has been seen, and that the stotting individual is strong and not worth chasing. In the mating season, rutting males bellow to make their presence known to females. Muskoxen roar during male-male fights, and male saigas force air through their noses, producing a roar to deter rival males and attract females. Mothers also use vocal communication to locate their calves if they get separated. During fights over dominance, males tend to display themselves in an erect posture with

13970-477: The recognition of Peleinae and Pantholopinae , comprising the genera Pelea and Pantholops respectively, as subfamilies . In 2000, American biologist George Schaller and palaeontologist Elisabeth Vrba suggested the inclusion of Pelea in Reduncinae , though the grey rhebok, the sole species of Pelea , is highly different from kobs and reduncines in morphology. Pantholops , earlier classified in

14097-455: The red hartebeest. In one study, the highest dimorphism was found in skull weight. Another study concluded that the length of the breeding season is a good predictor of dimorphism in pedicle (the bony structures from which the horns grow) height and skull weight, and the best predictor of the horn circumference. Active mainly during daytime, the hartebeest grazes in the early morning and late afternoon, and rests in shade around noon. Gregarious ,

14224-401: The same time a broad land-bridge connected Africa with Asia and there was a great invasion of animals of the steppe fauna into Africa. At the beginning of the Pleistocene a moist period set in and much of the forest was renewed while the grassland fauna was divided and isolated, as the forest fauna had previously been. The present forest fauna is therefore of double origin, partly descended of

14351-518: The sassabies (genus Damaliscus ), such as an elongated and narrow face, the shape of the horns, the pelage texture and colour, and the terminal tuft of the tail. The wildebeest have more specialised skull and horn features than the hartebeest. The hartebeest exhibits sexual dimorphism , but only slightly, as both sexes bear horns and have similar body masses. The degree of sexual dimorphism varies by subspecies. Males are 8% heavier than females in Swayne's and Lichtenstein's hartebeest, and 23% heavier in

14478-460: The sides, growing backward at first and then forward, ending with a sharp backward turn. The horns of the red and the Lelwel hartebeest are similar to those of the western hartebeest, but appear V-shaped when viewed from the front. The Lichtenstein's hartebeest has thick parallel ringed horns, with a flat base. Its horns are shorter than those of other subspecies, curving upward then sharply forward, followed by an inward turn at an angle of about 45° and

14605-468: The skull characters of living and extinct species of antelope to make a cladogram , and argued that a wide skull linked Lichtenstein's hartebeest with Connochaetes . However, this finding was not replicated by Alan W. Gentry of the Natural History Museum , who classified it as an independent species of Alcelaphus . Zoologists such as Jonathan Kingdon and Theodor Haltenorth considered it to be

14732-455: The species forms herds of up to 300 individuals. Larger numbers gather in places with abundant grass. In 1963, a congregation of 10,000 animals was recorded on the plains near Sekoma Pan in Botswana. However, moving herds are not so cohesive, and tend to disperse frequently. The members of a herd can be divided into four groups: territorial adult males, non-territorial adult males, young males, and

14859-815: The splitting up of the Gondwana supercontinent in the mid- Mesozoic era . After that, four to six faunal assemblages, the so-called African Faunal Strata (AFSs) can be distinguished. The isolation of Africa was broken intermittently by discontinuous "filter routes" that linked it to some other Gondwanan continents ( Madagascar , South America , and perhaps India ), but mainly to Laurasia . Interchanges with Gondwana were rare and mainly "out-of-Africa" dispersals, whereas interchanges with Laurasia were numerous and bidirectional, although mainly from Laurasia to Africa. Despite these connections, isolation resulted in remarkable absences, poor diversity, and emergence of endemic taxa in Africa. Madagascar separated from continental Africa during

14986-463: The strongly contrasting patterns helping to delay recognition by predators. However, all the Hippotraginae (including the gemsbok) have pale bodies and faces with conspicuous markings. The zoologist Tim Caro describes this as difficult to explain, but given that the species are diurnal, he suggests that the markings may function in communication. Strongly contrasting leg colouration is common only in

15113-645: The subfamily Tethytraginae , which contains Tethytragus (mid- Miocene ). In 1992, Alan W . Gentry of the Natural History Museum, London divided the eight major subfamilies of Bovidae into two major clades on the basis of their evolutionary history: the Boodontia, which comprised only the Bovinae, and the Aegodontia, which consisted of the rest of the subfamilies. Boodonts have somewhat primitive teeth, resembling those of oxen , whereas aegodonts have more advanced teeth like those of goats. A controversy exists about

15240-405: The third and fourth metapodials are combined into the cannon bone . The ulna and fibula are reduced, and fused with the radius and tibia, respectively. Long scapulae are present, whereas the clavicles are absent. Being ruminants , the stomach is composed of four chambers: the rumen (80%), the omasum , the reticulum , and the abomasum . The ciliates and bacteria of the rumen ferment

15367-754: The tropics spared it also from Pleistocene glaciations and the climate has not changed much. There are large gaps in human knowledge about African invertebrates. East Africa has a rich coral fauna with about 400 known species. More than 400 species of Echinoderms and 500 species of Bryozoa live there too, as well as one Cubozoan species ( Carybdea alata ). Of nematodes , the Onchocerca volvulus , Necator americanus , Wuchereria bancrofti and Dracunculus medinensis are human parasites. Some of important plant-parasitic nematodes of crops include Meloidogyne , Pratylenchus , Hirschmanniella , Radopholus , Scutellonema and Helicotylenchus . Of

15494-431: The vertebrae in this region. Gregarious animals, hartebeest form herds of 20 to 300 individuals. They are very alert and non-aggressive. They are primarily grazers , with their diets consisting mainly of grasses. Mating in hartebeest takes place throughout the year with one or two peaks, and depends upon the subspecies and local factors. Both males and females reach sexual maturity at one to two years of age. Gestation

15621-490: The word "hartebeest" in South African literature was in Dutch colonial administrator Jan van Riebeeck 's journal Daghregister in 1660. He wrote: " Meester Pieter ein hart-beest geschooten hadde (Master Pieter [van Meerhoff] had shot one hartebeest)". Another name for the hartebeest is kongoni , a Swahili word. Kongoni is often used to refer in particular to one of its subspecies , Coke's hartebeest . The scientific name of

15748-501: The world. 20 genera of freshwater crabs are present. The soil animal communities tropical Africa are poorly known. A few ecological studies have been undertaken on macrofauna, mainly in West Africa. Earthworms are being extensively studied in West and South Africa. Approximately 100,000 species of insects have been described from Sub-Saharan Africa , but there are very few overviews of

15875-504: Was attributed to difficulties in segregation during meiosis , indicated by azoospermia and a low number of germ cells in its seminiferous tubules . There are three well-defined hybrids between the subspecies: The genus Alcelaphus emerged about 4.4 million years ago in a clade whose other members were Damalops , Numidocapra , Rabaticeras , Megalotragus , Oreonagor , and Connochaetes . An analysis using phylogeographic patterns within hartebeest populations suggested

16002-493: Was given by the British zoologist John Edward Gray in 1821. The word "Bovidae" is the combination of the prefix bov- (originating from Latin bos , "ox", through Late Latin bovinus ) and the suffix -idae . The family Bovidae is placed in the order Artiodactyla (which includes the even-toed ungulates ). It includes 143 extant species, accounting for nearly 55% of the ungulates , and 300 known extinct species. Until

16129-555: Was removed, and both groups expanded into the territory of the other. The tribes Bovini and Tragelaphini diverged in the early Miocene. Bovids are known to have reached the Americas in the Pleistocene by crossing the Bering land bridge . The present genera of Alcelaphinae appeared in the Pliocene . The extinct Alcelaphine genus Paramularius , which was the same in size as the hartebeest,

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