155-411: Delphinus phocoena Linnaeus, 1758 The harbour porpoise ( Phocoena phocoena ) is one of eight extant species of porpoise . It is one of the smallest species of cetacean . As its name implies, it stays close to coastal areas or river estuaries, and as such, is the most familiar porpoise to whale watchers . This porpoise often ventures up rivers, and has been seen hundreds of kilometres from
310-453: A moment magnitude of 9.2 (more than a thousand times as powerful as the 1989 San Francisco earthquake ). The time of day (5:36 pm), time of year (spring) and location of the epicenter were all cited as factors in potentially sparing thousands of lives, particularly in Anchorage. Alaska suffered a more severe megathrust earthquake on July 11, 1585, estimated at magnitude 9.25, which remains
465-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
620-409: A territory on May 11, 1912. It was admitted as the 49th state of the U.S. on January 3, 1959. Abundant natural resources have enabled Alaska— with one of the smallest state economies—to have one of the highest per capita incomes , with commercial fishing , and the extraction of natural gas and oil, dominating Alaska's economy . U.S. Armed Forces bases and tourism also contribute to
775-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
930-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
1085-416: A coastal distribution that potentially brings them close to sources of pollution. Porpoises may not experience any toxic effects until they draw on their fat reserves, such as in periods of food shortage, migration or reproduction. An increase in the temperature of the sea water is likely to affect the distribution of porpoises and their prey, but has not been shown to occur. Reduced stocks of sand eel along
1240-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
1395-399: A convoy to Valdez. On the advice of military and civilian leaders, President Lyndon B. Johnson declared all of Alaska a major disaster area the day after the quake. The U.S. Navy and U.S. Coast Guard deployed ships to isolated coastal communities to assist with immediate needs. Bad weather and poor visibility hampered air rescue and observation efforts the day after the quake, but on Sunday
1550-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
1705-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
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#17328520276631860-447: A high energy source. Some studies suggest porpoises are relatively sedentary and usually do not leave a certain area for long. Nevertheless, they have been recorded to move from onshore to offshore waters along coast. Dives of 220 metres (120 fathoms) by harbour porpoises have been recorded. Dives can last five minutes but typically last one minute. The social life of harbour porpoises is not well understood. They are generally seen as
2015-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On
2170-462: A longer coastline than all the other U.S. states combined. At 663,268 square miles (1,717,856 km ) in total area, Alaska is by far the largest state in the United States. Alaska is more than twice the size of the second-largest U.S. state (Texas), and it is larger than the next three largest states (Texas, California, and Montana) combined. Alaska is the seventh largest subnational division in
2325-797: A massive underwater landslide, resulting in the deaths of 32 people between the collapse of the Valdez city harbor and docks, and inside the ship that was docked there at the time. Nearby, a 27-foot (8.2 m) tsunami destroyed the village of Chenega , killing 23 of the 68 people who lived there; survivors out-ran the wave, climbing to high ground. Post-quake tsunamis severely affected Whittier , Seward , Kodiak, and other Alaskan communities, as well as people and property in British Columbia, Washington , Oregon , and California . Tsunamis also caused damage in Hawaii and Japan . Evidence of motion directly related to
2480-430: A maximum weight of around 76 kg (168 lb) compared with the males' 61 kg (134 lb). The body is robust, and the animal is at its maximum girth just in front of its triangular dorsal fin . The beak is poorly demarcated. The flippers, dorsal fin, tail fin and back are a dark grey. The sides are a slightly speckled, lighter grey. The underside is much whiter, though there are usually grey stripes running along
2635-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
2790-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
2945-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that
3100-645: A population that was genetically separate from other native groups present elsewhere in the New World at the end of the Pleistocene . Ben Potter, the University of Alaska Fairbanks archaeologist who unearthed the remains at the Upward Sun River site in 2013, named this new group Ancient Beringian . The Tlingit people developed a society with a matrilineal kinship system of property inheritance and descent in what
3255-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
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#17328520276633410-402: A solitary species. Most of the time, porpoises are either alone or in groups of no more than five animals. Porpoises mate promiscuously. Males produce large amounts of sperm, perhaps for sperm competition . Females become sexually mature by their third or fourth year and can calve each year for several consecutive years, being pregnant and lactating at the same time. The gestation of the porpoise
3565-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
3720-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
3875-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
4030-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
4185-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
4340-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
4495-633: A total of 50 Aleut civilians and eight sailors were interned in Japan. About half of the Aleuts died during the period of internment. Unalaska / Dutch Harbor and Adak became significant bases for the United States Army , United States Army Air Forces and United States Navy. The United States Lend-Lease program involved flying American warplanes through Canada to Fairbanks and then Nome ; Soviet pilots took possession of these aircraft, ferrying them to fight
4650-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
4805-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
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4960-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
5115-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
5270-602: Is a non-contiguous U.S. state on the northwest extremity of North America . Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii . Alaska is also considered to be the northernmost, westernmost, and easternmost (the Aleutian Islands cross the 180th meridian into the eastern hemisphere) state in the United States. It borders the Canadian territory of Yukon and
5425-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
5580-452: Is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use
5735-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
5890-566: Is about 335,000. In the Western Atlantic it is estimated that there are about 33,000 harbour porpoises along the mid-southwestern coast of Greenland (where increasing temperatures have aided them), 75,000 between the Gulf of Maine and Gulf of St. Lawrence , and 27,000 in the Gulf of St. Lawrence. The Pacific population off mainland United States is about 73,000 and off Alaska 89,000. After sharp declines in
6045-787: Is based on the testimony of Chukchi geographer Nikolai Daurkin, who had visited Alaska in 1764–1765 and who had reported on a village on the Kheuveren River, populated by "bearded men" who "pray to the icons ". Some modern researchers associate Kheuveren with Koyuk River . The first European vessel to reach Alaska is generally held to be the St. Gabriel under the authority of the surveyor M. S. Gvozdev and assistant navigator I. Fyodorov on August 21, 1732, during an expedition of Siberian Cossack A. F. Shestakov and Russian explorer Dmitry Pavlutsky (1729–1735). Another European contact with Alaska occurred in 1741, when Vitus Bering led an expedition for
6200-797: Is bordered by Canada's Yukon and British Columbia to the east (making it the only state to border only a Canadian territory ); the Gulf of Alaska and the Pacific Ocean to the south and southwest; the Bering Sea , Bering Strait , and Chukchi Sea to the west; and the Arctic Ocean to the north. Alaska's territorial waters touch Russia's territorial waters in the Bering Strait, as the Russian Big Diomede Island and Alaskan Little Diomede Island are only 3 miles (4.8 km) apart. Alaska has
6355-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
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6510-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,
6665-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
6820-479: Is divided into boroughs . Delegates to the Alaska Constitutional Convention wanted to avoid the pitfalls of the traditional county system and adopted their own unique model. Many of the more densely populated parts of the state are part of Alaska's 16 boroughs, which function somewhat similarly to counties in other states. Unlike county-equivalents in the other states, the boroughs do not cover
6975-594: Is from Medieval Latin porcopiscus , which is a compound of porcus (pig) and piscus (fish). The old word is probably a loan-translation of a Germanic word, compare Danish marsvin and Middle Dutch mereswijn (sea swine). Classical Latin had a similar name, porculus marinus . The species' taxonomic name, Phocoena phocoena , is the Latinized form of the Greek φώκαινα, phōkaina , "big seal", as described by Aristotle ; this from φώκη, phōkē , " seal ". The species
7130-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
7285-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
7440-554: Is home to Mount Shishaldin , which is an occasionally smoldering volcano that rises to 10,000 feet (3,000 m) above the North Pacific. The chain of volcanoes extends to Mount Spurr , west of Anchorage on the mainland. Geologists have identified Alaska as part of Wrangellia , a large region consisting of multiple states and Canadian provinces in the Pacific Northwest , which is actively undergoing continent building . One of
7595-614: Is known for its massive reserves of crude oil and contains both the National Petroleum Reserve–Alaska and the Prudhoe Bay Oil Field . The city of Utqiaġvik , formerly known as Barrow, is the northernmost city in the United States and is located here. The Northwest Arctic area , anchored by Kotzebue and also containing the Kobuk River valley, is often considered part of this region. The respective Inupiat of
7750-451: Is maintained by the Office of History and Archaeology. The survey's inventory of cultural resources includes objects, structures, buildings, sites, districts, and travel ways, with a general provision that they are more than fifty years old. As of 31 January 2012 , more than 35,000 sites have been reported. Alaska is not divided into counties , as most of the other U.S. states, but it
7905-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
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#17328520276638060-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
8215-506: Is now Southeast Alaska , became the capital of Russian America . It remained the capital after the colony was transferred to the United States. The Russians never fully colonized Alaska, and the colony was never very profitable. Evidence of Russian settlement in names and churches survives throughout southeastern Alaska. In 1867, William H. Seward , the United States Secretary of State under President Andrew Johnson , negotiated
8370-605: Is owned and managed by the U.S. federal government as public lands, including a multitude of national forests , national parks, and national wildlife refuges . Of these, the Bureau of Land Management manages 87 million acres (35 million hectares), or 23.8% of the state. The Arctic National Wildlife Refuge is managed by the United States Fish and Wildlife Service . It is the world's largest wildlife refuge, comprising 16 million acres (6.5 million hectares). Of
8525-519: Is proportionally the second highest of any U.S. state, at over 15 percent, after only Hawaii. The name "Alaska" ( Russian : Аля́ска , romanized : Aljáska ) was introduced during the Russian colonial period when it was used to refer to the Alaska Peninsula . It was derived from an Aleut-language idiom , alaxsxaq , meaning "the mainland" or, more literally, "the object towards which
8680-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
8835-441: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Alaska Alaska ( / ə ˈ l æ s k ə / ə- LASS -kə )
8990-714: Is sometimes known as the common porpoise in texts originating in the United Kingdom. In parts of Atlantic Canada it is known colloquially as the puffing pig, and in Norway ‘nise’, derived from an Old Norse word for sneeze; both of which refer to the sound made when porpoises surface to breathe. The harbour porpoise is a little smaller than the other porpoises, at about 67–85 cm ( 26 + 1 ⁄ 2 – 33 + 1 ⁄ 2 in) long at birth, weighing 6.4–10 kg. Adults of both sexes grow to 1.4 to 1.9 m (4 ft 7 in to 6 ft 3 in). The females are heavier, with
9145-565: Is technically part of the continental U.S. , but is not usually included in the colloquial use of the term; Alaska is not part of the contiguous U.S. , often called " the Lower 48 ". The capital city, Juneau , is situated on the mainland of the North American continent but is not connected by road to the rest of the North American highway system. The largest lake in Alaska is Lake Illiamna . The state
9300-594: Is the largest glacier in North America, covering 2,008 square miles (5,200 km ) alone. There are no officially defined borders demarcating the various regions of Alaska, but there are five/six regions that the state is most commonly broken up into: The most populous region of Alaska contains Anchorage , the Matanuska-Susitna Valley and the Kenai Peninsula . Rural, mostly unpopulated areas south of
9455-576: Is the northernmost and westernmost state in the United States, but also has the most easterly longitude in the United States because the Aleutian Islands extend into the Eastern Hemisphere . Alaska is the only non- contiguous U.S. state on continental North America; about 500 miles (800 km) of Canadian territory consisting of British Columbia (in Canada ) separates Alaska from Washington . It
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#17328520276639610-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
9765-601: Is today Southeast Alaska, along with parts of British Columbia and the Yukon . Also in Southeast were the Haida , now well known for their unique arts. The Tsimshian people came to Alaska from British Columbia in 1887, when President Grover Cleveland , and later the U.S. Congress, granted them permission to settle on Annette Island and found the town of Metlakatla, Alaska . All three of these peoples, as well as other indigenous peoples of
9920-487: Is typically 10–11 months. Most births occur in late spring and summer. Calves are weaned after 8–12 months. Their average life-span in the wild is 8–13 years, although exceptionally individuals have reached up to 20, and in captivity up to 28 years. In a study of 239 dead harbour porpoises in the Gulf of Maine–Bay of Fundy, the vast majority were less than 12 years old and the oldest was 17. Harbour porpoises were traditionally hunted for food, as well as for their blubber, which
10075-622: The Exxon Valdez hit a reef in the Prince William Sound, spilling more than 11 million gallons (42 megalitres) of crude oil over 1,100 miles (1,800 km) of coastline. Today, the battle between philosophies of development and conservation is seen in the contentious debate over oil drilling in the Arctic National Wildlife Refuge and the proposed Pebble Mine . Located at the northwest corner of North America , Alaska
10230-573: The Alaska Peninsula are considered part of the Southwest, with the Aleutian Islands often (but not always) being grouped in as well. While primarily part of Southwest Alaska when grouped economically, the Aleutian islands are sometimes recognized as an alternate group from the rest of the region due to the geographic separation from the continent. More than 300 small volcanic islands make up this chain, which stretches more than 1,200 miles (1,900 km) into
10385-727: The Alaska Purchase (referred to pejoratively as Seward's Folly) with the Russians for $ 7.2 million. Russia's contemporary ruler Tsar Alexander II , the Emperor of the Russian Empire , King of Poland and Grand Duke of Finland , also planned the sale; the purchase was made on March 30, 1867. Six months later the commissioners arrived in Sitka and the formal transfer was arranged; the formal flag-raising took place at Fort Sitka on October 18, 1867. In
10540-612: The Alaska Purchase . The region is dominated by the Alexander Archipelago as well as the Tongass National Forest , the largest national forest in the United States. It contains the state capital Juneau , the former capital Sitka , and Ketchikan , at one time Alaska's largest city. The Alaska Marine Highway provides a vital surface transportation link throughout the area and country, as only three communities ( Haines , Hyder and Skagway ) enjoy direct connections to
10695-641: The Alaska Range and west of the Wrangell Mountains also fall within the definition of South Central, as do the Prince William Sound area and the communities of Cordova and Valdez . Also referred to as the Panhandle or Inside Passage , this is the region of Alaska closest to the contiguous states. As such, this was where most of the initial non-indigenous settlement occurred in the years following
10850-636: The Alby People of the east coast of Öland , Sweden . The main threat to porpoises is static fishing techniques such as gill and tangle nets. Bycatch in bottom-set gill nets is considered the main anthropogenic mortality factor for harbour porpoises worldwide. Several thousand die each year in incidental bycatch, which has been reported from the Black Sea, the Baltic Sea, the North Sea, off California, and along
11005-577: The Bering land bridge . The Russian Empire was the first to actively colonize the area beginning in the 18th century, eventually establishing Russian America , which spanned most of the current state and promoted and maintained a native Alaskan Creole population. The expense and logistical difficulty of maintaining this distant possession prompted its sale to the U.S. in 1867 for US$ 7.2 million (equivalent to $ 157 million in 2023). The area went through several administrative changes before becoming organized as
11160-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
11315-639: The National Wildlife Refuge system , parts of 25 rivers to the National Wild and Scenic Rivers system , 3.3 million acres (13,000 km ) to National Forest lands , and 43.6 million acres (176,000 km ) to National Park land . Because of the Act, Alaska now contains two-thirds of all American national parklands. Today, more than half of Alaskan land is owned by the Federal Government . In 1989,
11470-535: The North Sea caught them by chance. A study published by the online journal of the Natural History Museum Rotterdam points out that conjoined twins in whales and dolphins are extremely rare. The vocalizations of the harbour porpoise is made up of short clicks from 0.5 to 5 milliseconds in bursts up to two seconds long. Each click has a frequency between 1000 and 2200 hertz. Aside from communication,
11625-508: The University of Aberdeen in Scotland have also discovered that the local bottlenose dolphins attack and kill harbour porpoises without eating them due to competition for a decreasing food supply. An alternative explanation is that the adult dolphins exhibit infanticidal behaviour and mistake the porpoises for juvenile dolphins which they are believed to kill. Grey seals are also known to attack harbour porpoises by biting off chunks of fat as
11780-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
11935-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
12090-602: The province of British Columbia to the east. It shares a western maritime border, in the Bering Strait , with Russia's Chukotka Autonomous Okrug . The Chukchi and Beaufort Seas of the Arctic Ocean lie to the north, and the Pacific Ocean lies to the south. Technically, it is a semi-exclave of the U.S., and is the largest exclave in the world. Alaska is the largest U.S. state by area, comprising more total area than
12245-418: The public record in Alaska. The state is divided into 34 recording districts which are centrally administered under a state recorder . All recording districts use the same acceptance criteria, fee schedule, etc., for accepting documents into the public record. Whereas many U.S. states use a three-tiered system of decentralization—state/county/township—most of Alaska uses only two tiers—state/borough. Owing to
12400-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
12555-677: The 20th century, populations have rebounded in the inland waters of Washington state . In contrast, some subpopulations are seriously threatened. For example, there are less than 12,000 in the Black Sea, and only about 500 remaining in the Baltic Sea proper, representing a sharp decrease since the mid-1900s. Harbour porpoises prefer temperate and subarctic waters. They inhabit fjords, bays, estuaries and harbours, hence their name. They feed mostly on small pelagic schooling fish, particularly herring , pollack , hake , sardine , cod , capelin , and sprat . They will, however, eat squid and crustaceans in certain places. This species tends to feed close to
12710-465: The 29th the situation improved and rescue helicopters and observation aircraft were deployed. A military airlift immediately began shipping relief supplies to Alaska, eventually delivering 2,570,000 pounds (1,170,000 kg) of food and other supplies. Broadcast journalist, Genie Chance , assisted in recovery and relief efforts, staying on the KENI air waves over Anchorage for more than 24 continuous hours as
12865-656: The Alaska Division of Emergency Services to respond to any future disasters. The 1968 discovery of oil at Prudhoe Bay and the 1977 completion of the Trans-Alaska Pipeline System led to an oil boom. Royalty revenues from oil have funded large state budgets from 1980 onward. Oil production was not the only economic value of Alaska's land. In the second half of the 20th century, Alaska discovered tourism as an important source of revenue. Tourism became popular after World War II when military personnel stationed in
13020-806: The Black Sea endangered . In addition, the harbour porpoise is covered by the Agreement on the Conservation of Small Cetaceans of the Baltic, North East Atlantic, Irish and North Seas ( ASCOBANS ), the Agreement on the Conservation of Cetaceans in the Black Sea, Mediterranean Sea and Contiguous Atlantic Area (ACCOBAMS) and the Memorandum of Understanding Concerning the Conservation of the Manatee and Small Cetaceans of Western Africa and Macaronesia (Western African Aquatic Mammals MoU). Species A species ( pl. : species)
13175-509: The Convention on the Conservation of Migratory Species of Wild Animals ( CMS ). In 2013, the two Baltic Sea subpopulations were listed as vulnerable and critically endangered respectively by HELCOM . Although the species overall is considered to be of Least Concern by the IUCN , they consider the Baltic Sea and Western African populations critically endangered, and the subspecies P. p. relicta of
13330-661: The German invasion of the Soviet Union . The construction of military bases contributed to the population growth of some Alaskan cities. Statehood for Alaska was an important cause of James Wickersham early in his tenure as a congressional delegate. Decades later, the statehood movement gained its first real momentum following a territorial referendum in 1946. The Alaska Statehood Committee and Alaska's Constitutional Convention would soon follow. Statehood supporters also found themselves fighting major battles against political foes, mostly in
13485-615: The North Atlantic and West Africa, P. p. relicta in the Black Sea and Sea of Azov , an unnamed population in the northwest Pacific and P. p. vomerina in the northeast Pacific. Concerning the North Atlantic, an international workshop co-organised by the North Atlantic Marine Mammal Commission and the Norwegian Institute of Marine Research reviewed the status of the species in 2018. It concluded that
13640-573: The North Slope and of the Northwest Arctic seldom consider themselves to be one people. Southwest Alaska is a sparsely inhabited region stretching some 500 miles (800 km) inland from the Bering Sea. Most of the population lives along the coast. Kodiak Island is also located in the Southwest. The massive Yukon–Kuskokwim Delta , one of the largest river deltas in the world, is here. Portions of
13795-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
13950-573: The Pacific Northwest Coast , experienced smallpox outbreaks from the late 18th through the mid- 19th century , with the most devastating epidemics occurring in the 1830s and 1860s, resulting in high fatalities and social disruption. Some researchers believe the first Russian settlement in Alaska was established in the 17th century. According to this hypothesis, in 1648 several koches of Semyon Dezhnyov 's expedition came ashore in Alaska by storm and founded this settlement. This hypothesis
14105-595: The Pacific Northwest. In 1789, a Spanish settlement and fort were built in Nootka Sound . These expeditions gave names to places such as Valdez , Bucareli Sound , and Cordova . Later, the Russian-American Company carried out an expanded colonization program during the early-to-mid-19th century. Sitka , renamed New Archangel from 1804 to 1867, on Baranof Island in the Alexander Archipelago in what
14260-627: The Pacific Ocean. Some of these islands fall in the Eastern Hemisphere, but the International Date Line was drawn west of 180° to keep the whole state, and thus the entire North American continent, within the same legal day. Two of the islands, Attu and Kiska , were occupied by Japanese forces during World War II. According to an October 1998 report by the United States Bureau of Land Management , approximately 65% of Alaska
14415-456: The Russian Navy aboard the St. Peter . After his crew returned to Russia with sea otter pelts judged to be the finest fur in the world, small associations of fur traders began to sail from the shores of Siberia toward the Aleutian Islands. The first permanent European settlement was founded in 1784. Between 1774 and 1800, Spain sent several expeditions to Alaska to assert its claim over
14570-436: The U.S. Congress but also within Alaska. Statehood was approved by the U.S. Congress on July 7, 1958; Alaska was officially proclaimed a state on January 3, 1959. On March 27, 1964, the massive Good Friday earthquake killed 133 people and destroyed several villages and portions of large coastal communities, mainly by the resultant tsunamis and landslides. It was the fourth-most-powerful earthquake in recorded history, with
14725-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
14880-559: The action of the sea is directed". Numerous indigenous peoples occupied Alaska for thousands of years before the arrival of European peoples to the area. Linguistic and DNA studies done here have provided evidence for the settlement of North America by way of the Bering land bridge . At the Upward Sun River site in the Tanana Valley in Alaska, remains of a six-week-old infant were found. The baby's DNA showed that she belonged to
15035-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
15190-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
15345-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
15500-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be
15655-489: The ceremony, 250 uniformed U.S. soldiers marched to the governor's house at "Castle Hill", where the Russian troops lowered the Russian flag and the U.S. flag was raised. This event is celebrated as Alaska Day , a legal holiday on October 18. Alaska was loosely governed by the military initially and was administered as a district starting in 1884, with a governor appointed by the United States president. A federal district court
15810-539: The city proper and the communities of Eagle River, Chugiak, Peters Creek, Girdwood, Bird, and Indian. Fairbanks has a separate borough (the Fairbanks North Star Borough ) and municipality (the City of Fairbanks). The state's most populous city is Anchorage , home to 291,247 people in 2020. The richest location in Alaska by per capita income is Denali ($ 42,245). Yakutat City , Sitka, Juneau, and Anchorage are
15965-524: The clicks are used for echolocation. The harbour porpoise species is widespread in cooler coastal waters of the North Atlantic, North Pacific and the Black Sea . In the Atlantic, harbour porpoises may be present in a curved band of water running from the coast of West Africa to the coasts of Portugal, Spain, France, the United Kingdom, Ireland, Scandinavia, Iceland , Greenland, Nova Scotia and Newfoundland and
16120-492: The collapse of herring in the North Sea caused porpoises to hunt for other prey species. Reduction of prey may result from climate change, overfishing, or both. Noise from ship traffic and oil platforms is thought to affect the distribution of toothed whales, like the harbour porpoise, that use echolocation for communication and prey detection. Noise from shipping traffic, particularly busy sea lanes, appears to instigate evasive behavior, with predominantly lateral movements during
16275-429: The community, disseminating information about shelters and prepared food rations, passing messages of well-being between loved ones, and helping to reunite families. In the longer term, the U.S. Army Corps of Engineers led the effort to rebuild roads, clear debris, and establish new townsites for communities that had been completely destroyed, at a cost of $ 110 million. The West Coast and Alaska Tsunami Warning Center
16430-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
16585-416: The concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part
16740-410: The contiguous North American road system. The Interior is the largest region of Alaska; much of it is uninhabited wilderness. Fairbanks is the only large city in the region. Denali National Park and Preserve is located here. Denali , formerly Mount McKinley, is the highest mountain in North America and is also located here. The North Slope is mostly tundra peppered with small villages. The area
16895-456: The damage done by the earthquake to the city. The team, called the Engineering and Geological Evaluation Group, was headed by Ruth A. M. Schmidt , a geology professor at the University of Alaska Anchorage . The team of scientists came into conflict with local developers and downtown business owners who wanted to immediately rebuild; the scientists wanted to identify future dangers to ensure that
17050-626: The day and deeper dives during the night. The construction of thousands of offshore wind turbines, planned in different areas of North Sea, is known to cause displacement of porpoises from the construction site, particularly if steel monopile foundations are installed by percussive piling, where reactions can occur at distances of more than 20 km (11 nmi). Noise levels from operating wind turbines are low and unlikely to affect porpoises, even at close range. Marine top predators like porpoises and seals accumulate pollutants such as heavy metals, PCBs and pesticides in their fat tissue. Porpoises have
17205-512: The earthquake was also reported from Florida and Texas . Alaska had never experienced a major disaster in a highly populated area before and had very limited resources for dealing with the effects of such an event. In Anchorage, at the urging of geologist Lidia Selkregg , the City of Anchorage and the Alaska State Housing Authority appointed a team of 40 scientists, including geologists, soil scientists, and engineers, to assess
17360-427: The east coast of Scotland, a pattern linked to climate change, appears to be the main reason for the increase in malnutrition in porpoises in the area. Overall, the harbour porpoise is not considered threatened and the total population is in the hundreds of thousands. The harbour porpoise populations of the North Sea, Baltic Sea, western North Atlantic, Black Sea and North West Africa are protected under Appendix II of
17515-553: The east coast of the United States and Canada. Bottom-set gill nets are anchored to the sea floor and are up to 23 kilometres ( 12 + 1 ⁄ 2 nautical miles) in length. It is unknown why porpoises become entangled in gill nets, since several studies indicate they are able to detect these nets using their echolocation . Porpoise-scaring devices, so-called pingers, have been developed to keep porpoises out of nets and numerous studies have demonstrated they are very effective at reducing entanglement. However, concern has been raised over
17670-551: The eastern seaboard of the United States. The population in the Baltic Sea is limited in winter due to sea freezing, and is most common in the southwest parts of the sea. There is another band in the Pacific Ocean running from the Sea of Japan , Vladivostok , the Bering Strait , Alaska , British Columbia , and California . The populations in these regions are not continuous and are classified as separate subspecies with P. p. phocoena in
17825-458: The economy; more than half of the state is federally-owned land containing national forests , national parks , and wildlife refuges . It is among the most irreligious states , one of the first to legalize recreational marijuana , and is known for its libertarian-leaning political culture, generally supporting the Republican Party in national elections. The Indigenous population of Alaska
17980-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
18135-413: The fish they targeted. Almost all the fish they ate were very small, between 3 and 10 cm (1–4 in) long. A study (2024) shown that prey availability is an important driver of seasonal and diel dynamics of harbour porpoise acoustic activity in the Black Sea. In the southeastern region, porpoise activity was primarily nocturnal, with a peak from January to May, aligned with anchovy migration. On
18290-467: The following three largest states of Texas , California , and Montana combined, and is the seventh-largest subnational division in the world . It is the third-least populous and most sparsely populated U.S. state, but is, with a population of 736,081 as of 2020 , the continent's most populous territory located mostly north of the 60th parallel , with more than quadruple the combined populations of Northern Canada and Greenland . The state contains
18445-481: The four largest cities in the U.S. by area . As reflected in the 2020 United States census , Alaska has a total of 355 incorporated cities and census-designated places (CDPs). The tally of cities includes four unified municipalities, essentially the equivalent of a consolidated city–county . The majority of these communities are located in the rural expanse of Alaska known as " The Bush " and are unconnected to that contiguous North American road network. The table at
18600-416: The four largest cities in the United States by area , including the state capital of Juneau . The state's most populous city is Anchorage , and approximately half of Alaska's residents live within its metropolitan area . Indigenous people have lived in Alaska for thousands of years, and it is widely believed that the region served as the entry point for the initial settlement of North America by way of
18755-520: The harbour porpoise population structure is more complex than previously thought, with at least three genetically distinct subspecies in the North Atlantic. Given the structure of the harbour porpoise population, the workshop delineated 18 assessment areas for the North Atlantic. The harbour porpoise has a global population of at least 700,000. In 2016, a comprehensive survey of the Atlantic region in Europe, from Gibraltar to Vestfjorden in Norway, found that
18910-432: The largest private landowner in Alaska in advertisements and other communications. Provisions of ANCSA allowing the corporations' land holdings to be sold on the open market starting in 1991 were repealed before they could take effect. Effectively, the corporations hold title (including subsurface title in many cases, a privilege denied to individual Alaskans) but cannot sell the land. Individual Native allotments are sold on
19065-610: The low population density, most of the land is located in the Unorganized Borough . As the name implies, it has no intermediate borough government but is administered directly by the state government. In 2000, 57.71% of Alaska's area has this status, with 13.05% of the population. Anchorage merged the city government with the Greater Anchorage Area Borough in 1975 to form the Municipality of Anchorage, containing
19220-801: The most powerful earthquake recorded in North American history, and the second most powerful earthquake recorded in world history. The Good Friday earthquake lasted 4 minutes and 38 seconds. Six hundred miles (970 km) of fault ruptured at once and moved up to 60 ft (18 m), releasing about 500 years of stress buildup. Soil liquefaction , fissures, landslides, and other ground failures caused major structural damage in several communities and much damage to property. Anchorage sustained great destruction or damage to many inadequately earthquake-engineered houses, buildings, and infrastructure (paved streets, sidewalks, water and sewer mains, electrical systems, and other human-made equipment), particularly in
19375-406: The noise pollution created by the pingers and whether their efficiency will diminish over time due to porpoises habituating to the sounds. Mortality resulting from trawling bycatch seems to be less of an issue, probably because porpoises are not inclined to feed inside trawls, as dolphins are known to do. Overfishing may reduce preferred prey availability for porpoises. Overfishing resulting in
19530-502: The northwestern shelf, porpoises were more active during daylight from April to October, reflecting the migration patterns of sprat . Harbour porpoises tend to be solitary foragers, but they do sometimes hunt in packs and herd fish together. Young porpoises need to consume about 7% to 8% of their body weight each day to survive, which is approximately 15 pounds or 7 kilograms of fish. Significant predators of harbour porpoises include white sharks and killer whales (orcas). Researchers at
19685-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
19840-508: The often remote and roadless locations. The University of Alaska , as a land grant university , also owns substantial acreage which it manages independently. Another 44 million acres (18 million hectares) are owned by 12 regional, and scores of local, Native corporations created under the Alaska Native Claims Settlement Act (ANCSA) of 1971. Regional Native corporation Doyon, Limited often promotes itself as
19995-412: The open market. Various private interests own the remaining land, totaling about one percent of the state. Alaska is, by a large margin, the state with the smallest percentage of private land ownership when Native corporation holdings are excluded. The Alaska Heritage Resources Survey (AHRS) is a restricted inventory of all reported historic and prehistoric sites within the U.S. state of Alaska; it
20150-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
20305-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
20460-438: The population was about 467,000 harbour porpoises, making it the most abundant cetacean in the region, together with the common dolphin . Based on surveys in 1994, 2005 and 2016, the harbour porpoise population in this region is stable. The highest densities are in the southwestern North Sea and oceans of mainland Denmark ; the latter region alone is home to about 107,000-300,000 harbour porpoises. The entire North Sea population
20615-428: The rebuilt infrastructure would be safe. The team produced a report on May 8, 1964, just a little more than a month after the earthquake. The United States military, which has a large active presence in Alaska, also stepped in to assist within moments of the end of the quake. The U.S. Army rapidly re-established communications with the lower 48 states, deployed troops to assist the citizens of Anchorage, and dispatched
20770-590: The region returned home praising its natural splendor. The Alcan Highway , built during the war, and the Alaska Marine Highway System , completed in 1963, made the state more accessible than before. Tourism has become increasingly important in Alaska, and today over 1.4 million people visit the state each year. With tourism more vital to the economy, environmentalism also rose in importance. The Alaska National Interest Lands Conservation Act (ANILCA) of 1980 added 53.7 million acres (217,000 km ) to
20925-505: The remaining land area, the state of Alaska owns 101 million acres (41 million hectares), its entitlement under the Alaska Statehood Act . A portion of that acreage is occasionally ceded to the organized boroughs presented above, under the statutory provisions pertaining to newly formed boroughs. Smaller portions are set aside for rural subdivisions and other homesteading-related opportunities. These are not very popular due to
21080-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
21235-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
21390-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
21545-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
21700-403: The sea bottom, at least for waters less than 200 metres (110 fathoms) deep. However, when hunting sprat, porpoise may stay closer to the surface. When in deeper waters, porpoises may forage for mid-water fish, such as pearlsides . A study published in 2016 showed that porpoises off the coast of Denmark were hunting 200 fish per hour during the day and up to 550 per hour at night, catching 90% of
21855-487: The sea. The harbour porpoise may be polytypic , with geographically distinct populations representing distinct races: P. p. phocoena in the North Atlantic and West Africa, P. p. relicta in the Black Sea and Sea of Azov, an unnamed population in the northwestern Pacific and P. p. vomerina in the northeastern Pacific. The English word porpoise comes from the French pourpois ( Old French porpais , 12th century), which
22010-545: The several landslide zones along Knik Arm . Two hundred miles (320 km) southwest, some areas near Kodiak were permanently raised by 30 feet (9 m). Southeast of Anchorage, areas around the head of Turnagain Arm near Girdwood and Portage dropped as much as 8 feet (2.4 m), requiring reconstruction and fill to raise the Seward Highway above the new high tide mark. In Prince William Sound , Port Valdez suffered
22165-456: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
22320-461: The state's entire land area. The area not part of any borough is referred to as the Unorganized Borough . The Unorganized Borough has no government of its own, but the U.S. Census Bureau in cooperation with the state divided the Unorganized Borough into 11 census areas solely for the purposes of statistical analysis and presentation. A recording district is a mechanism for management of
22475-530: The throat from the underside of the body. Many anomalously white coloured individuals have been confirmed, mostly in the North Atlantic, but also notably around Turkish and British coasts, and in the Wadden Sea , Bay of Fundy and around the coast of Cornwall . Although conjoined twins are rarely seen in wild mammals, the first known case of a two-headed harbour porpoise was documented in May 2017 when Dutch fishermen in
22630-509: The voice of calm from her temporary post within the Anchorage Public Safety Building. She was effectively designated as the public safety officer by the city's police chief. Chance provided breaking news of the catastrophic events that continued to develop following the magnitude 9.2 earthquake, and she served as the voice of the public safety office, coordinating response efforts, connecting available resources to needs around
22785-537: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
22940-471: The world . If it was an independent nation, it would be the 18th largest country in the world; almost the same size as Iran . With its myriad of islands, Alaska has nearly 34,000 miles (55,000 km) of tidal shoreline. The Aleutian Islands chain extends west from the southern tip of the Alaska Peninsula . Many active volcanoes are found in the Aleutians and in coastal regions. Unimak Island , for example,
23095-513: The world's largest tides occurs in Turnagain Arm , just south of Anchorage, where tidal differences can be more than 35 feet (10.7 m). Alaska has more than 409,000 natural lakes at least one hectare or bigger. Marshlands and wetland permafrost cover 188,320 square miles (487,700 km ) (mostly in northern, western and southwest flatlands). Glacier ice covers about 28,957 square miles (75,000 km ) of Alaska. The Bering Glacier
23250-635: Was banned in 1899), and again in smaller scale during the shortages that occurred in World War I and World War II. A similar, short-lived re-emergence of hunting during the world wars happened in Poland and the Baltic countries. Currently, the species is only hunted as part of the traditional Inuit hunt in the Arctic, notably in Greenland. In prehistoric times, harbour porpoises were also hunted in many areas, for example by
23405-512: Was extremely limited or unavailable until statehood took effect in 1959. Starting in the 1890s and stretching in some places to the early 1910s, gold rushes in Alaska and the nearby Yukon Territory brought thousands of miners and settlers to Alaska. From 1879 to 1920, Alaska produced a cumulative total of over $ 460,000,000 ($ 6,691,927,500 inflation-adjusted) of mineral production. Alaska was officially incorporated as an organized territory in 1912. Alaska's capital, which had been in Sitka until 1906,
23560-483: Was formed as a direct response to the disaster. Federal disaster relief funds paid for reconstruction as well as financially supporting the devastated infrastructure of Alaska's government, spending hundreds of millions of dollars that helped keep Alaska financially solvent until the discovery of massive oil deposits at Prudhoe Bay . At the order of the U.S. Defense Department , the Alaska National Guard founded
23715-408: Was headquartered in Sitka. For most of Alaska's first decade under the United States flag, Sitka was the only community inhabited by American settlers. They organized a "provisional city government", which was Alaska's first municipal government, but not in a legal sense. Legislation allowing Alaskan communities to legally incorporate as cities did not come about until 1900, and home rule for cities
23870-752: Was moved north to Juneau . Construction of the Alaska Governor's Mansion began that same year. European immigrants from Norway and Sweden also settled in southeast Alaska, where they entered the fishing and logging industries. During World War II , the Aleutian Islands Campaign focused on Attu , Agattu and Kiska , all of which were occupied by the Empire of Japan . During the Japanese occupation, an American civilian and two United States Navy personnel were killed at Attu and Kiska respectively, and nearly
24025-614: Was used for lighting fuel. Among others, hunting occurred in the Black Sea , off Normandy , in the Bay of Biscay , off Flanders , in the Little Belt strait, off Iceland , western Norway , in Puget Sound , Bay of Fundy and Gulf of Saint Lawrence. The drive hunt in the Little Belt strait is the best documented example. Thousands of porpoises were caught there until the end of the 19th century (it
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