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A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water , either permanently, for years or decades, or only seasonally. Flooding results in oxygen -poor ( anoxic ) processes taking place, especially in the soils . Wetlands form a transitional zone between waterbodies and dry lands , and are different from other terrestrial or aquatic ecosystems due to their vegetation 's roots having adapted to oxygen-poor waterlogged soils . They are considered among the most biologically diverse of all ecosystems, serving as habitats to a wide range of aquatic and semi-aquatic plants and animals , with often improved water quality due to plant removal of excess nutrients such as nitrates and phosphorus .

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73-524: Southern James Bay is a coastal wetland complex in northeastern Ontario , Canada bordering James Bay and Quebec . It was designated as a wetland of international importance via the Ramsar Convention on May 27, 1987. The shallow waters of the James Bay region represent an important late autumn staging area for migratory, Arctic-breeding waterbirds . Two migratory bird sanctuaries are located in

146-711: A sink or a source of carbon, depending on the specific wetland. If they function as a carbon sink, they can help with climate change mitigation . However, wetlands can also be a significant source of methane emissions due to anaerobic decomposition of soaked detritus , and some are also emitters of nitrous oxide . Humans are disturbing and damaging wetlands in many ways, including oil and gas extraction , building infrastructure, overgrazing of livestock , overfishing , alteration of wetlands including dredging and draining, nutrient pollution , and water pollution . Wetlands are more threatened by environmental degradation than any other ecosystem on Earth, according to

219-531: A crucial role in the global carbon cycle, as it serves as the primary mechanism for removing CO 2 (carbon dioxide) from the atmosphere and incorporating it into living biomass. The primary production of organic compounds allows carbon to enter the biosphere . Carbon is considered essential for life as a base element for building organic compounds. The element of carbon forms the bases biogeochemical cycles (or nutrient cycles ) and drives communities of living organisms. Understanding biological carbon fixation

292-402: A frequency and duration to support, and that under normal circumstances do support, a prevalence of vegetation typically adapted for life in saturated soil conditions. Wetlands generally included swamps, marshes, bogs, and similar areas.' For each of these definitions and others, regardless of the purpose, hydrology is emphasized (shallow waters, water-logged soils). The soil characteristics and

365-1248: A polar climate, wetland temperatures can be as low as −50 °C (−58 °F). Peatlands in arctic and subarctic regions insulate the permafrost , thus delaying or preventing its thawing during summer, as well as inducing its formation. The amount of precipitation a wetland receives varies widely according to its area. Wetlands in Wales , Scotland , and western Ireland typically receive about 1,500 mm (59 in) per year. In some places in Southeast Asia , where heavy rains occur, they can receive up to 10,000 mm (390 in). In some drier regions, wetlands exist where as little as 180 mm (7.1 in) precipitation occurs each year. Temporal variation: Surface flow may occur in some segments, with subsurface flow in other segments. Wetlands vary widely due to local and regional differences in topography , hydrology , vegetation , and other factors, including human involvement. Other important factors include fertility, natural disturbance, competition, herbivory , burial and salinity. When peat accumulates, bogs and fens arise. The most important factor producing wetlands

438-540: A portion of the overall water cycle, which also includes atmospheric water (precipitation) and groundwater . Many wetlands are directly linked to groundwater and they can be a crucial regulator of both the quantity and quality of water found below the ground. Wetlands that have permeable substrates like limestone or occur in areas with highly variable and fluctuating water tables have especially important roles in groundwater replenishment or water recharge. Substrates that are porous allow water to filter down through

511-432: A process carried out by a diverse community of microorganisms. During decomposition, complex organic compounds are broken down into simpler molecules by the action of enzymes produced by bacteria, fungi, and other soil organisms. As organic matter is decomposed, carbon is released in various forms, including carbon dioxide (CO2) and dissolved organic carbon (DOC). However, not all of the carbon released during decomposition

584-429: A protective barrier to coastal shoreline. Mangroves stabilize the coastal zone from the interior and will migrate with the shoreline to remain adjacent to the boundary of the water. The main conservation benefit these systems have against storms and storm surges is the ability to reduce the speed and height of waves and floodwaters. Carbon fixation Biological carbon fixation , or сarbon assimilation ,

657-422: A strong influence on wetland water chemistry, particularly in coastal wetlands and in arid and semiarid regions with large precipitation deficits. Natural salinity is regulated by interactions between ground and surface water, which may be influenced by human activity. Carbon is the major nutrient cycled within wetlands. Most nutrients, such as sulfur , phosphorus , carbon , and nitrogen are found within

730-410: A unique kind of wetland where lush plant growth and slow decay of dead plants (under anoxic conditions) results in organic peat accumulating; bogs, fens, and mires are different names for peatlands. Variations of names for wetland systems: Some wetlands have localized names unique to a region such as the prairie potholes of North America's northern plain, pocosins , Carolina bays and baygalls of

803-508: A wetland is "an ecosystem that arises when inundation by water produces soils dominated by anaerobic and aerobic processes, which, in turn, forces the biota, particularly rooted plants, to adapt to flooding". Sometimes a precise legal definition of a wetland is required. The definition used for regulation by the United States government is: 'The term "wetlands" means those areas that are inundated or saturated by surface or ground water at

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876-422: A wetland system includes its plants ( flora ) and animals ( fauna ) and microbes (bacteria, fungi). The most important factor is the wetland's duration of flooding. Other important factors include fertility and salinity of the water or soils. The chemistry of water flowing into wetlands depends on the source of water, the geological material that it flows through and the nutrients discharged from organic matter in

949-403: A wetland) affects hydro-periods (temporal fluctuations in water levels) by controlling the water balance and water storage within a wetland. Landscape characteristics control wetland hydrology and water chemistry. The O 2 and CO 2 concentrations of water depend upon temperature , atmospheric pressure and mixing with the air (from winds or water flows). Water chemistry within wetlands

1022-558: A whole. To replace these wetland ecosystem services , enormous amounts of money would need to be spent on water purification plants, dams, levees, and other hard infrastructure, and many of the services are impossible to replace. Floodplains and closed-depression wetlands can provide the functions of storage reservoirs and flood protection. The wetland system of floodplains is formed from major rivers downstream from their headwaters . "The floodplains of major rivers act as natural storage reservoirs, enabling excess water to spread out over

1095-577: A wide area, which reduces its depth and speed. Wetlands close to the headwaters of streams and rivers can slow down rainwater runoff and spring snowmelt so that it does not run straight off the land into water courses. This can help prevent sudden, damaging floods downstream." Notable river systems that produce wide floodplains include the Nile River , the Niger river inland delta, the Zambezi River flood plain,

1168-412: A willow-alder shrub area into a drier forest interspersed with fens and bogs ". Gradually rising inland from sea level, it attains elevations of no more than 50 metres (160 ft). Wetland Wetlands exist on every continent , except Antarctica . The water in wetlands is either freshwater , brackish or saltwater . The main types of wetland are defined based on the dominant plants and

1241-443: Is glyceraldehyde 3-phosphate (GAP) together with dihydroxyacetone phosphate (DHAP): An alternative perspective accounts for NADPH (source of e ) and ATP: The formula for inorganic phosphate (P i ) is HOPO 3 + 2H . Formulas for triose and TP are C 2 H 3 O 2 -CH 2 OH and C 2 H 3 O 2 -CH 2 OPO 3 + 2H The reverse Krebs cycle , also known as the reverse TCA cycle (rTCA) or reductive citric acid cycle ,

1314-440: Is a very expensive pathway: 7 ATP molecules are used for the synthesis of the new pyruvate and 3 ATP for the phosphate triose. An important characteristic of this cycle is that it allows the co-assimilation of numerous compounds making it suitable for the mixotrophic organisms. A variant of the 3-hydroxypropionate cycle was found to operate in the aerobic extreme thermoacidophile archaeon Metallosphaera sedula . This pathway

1387-400: Is an alternative to the standard Calvin-Benson cycle for carbon fixation. It has been found in strict anaerobic or microaerobic bacteria (as Aquificales ) and anaerobic archea . It was discovered by Evans, Buchanan and Arnon in 1966 working with the photosynthetic green sulfur bacterium Chlorobium limicola . In particular, it is one of the most used pathways in hydrothermal vents by

1460-462: Is called the 3-hydroxypropionate/4-hydroxybutyrate (3-HP/4-HB) cycle. Yet another variant of the 3-hydroxypropionate cycle is the dicarboxylate/4-hydroxybutyrate (DC/4-HB) cycle. It was discovered in anaerobic archaea. It was proposed in 2008 for the hyperthermophile archeon Ignicoccus hospitalis . CO 2 fixation is catalyzed by enoyl-CoA carboxylases/reductases. Although no heterotrophs use carbon dioxide in biosynthesis, some carbon dioxide

1533-465: Is carbon dioxide (CO 2 ). It is estimated that approximately 250 billion tons of carbon dioxide are converted by photosynthesis annually. The majority of the fixation occurs in terrestrial environments, especially the tropics. The gross amount of carbon dioxide fixed is much larger since approximately 40% is consumed by respiration following photosynthesis. Historically, it is estimated that approximately 2×10 billion tons of carbon has been fixed since

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1606-475: Is composed of two cycles and the name of this way comes from the 3-Hydroxyporopionate which corresponds to an intermediate characteristic of it. The first cycle is a way of synthesis of glyoxylate . During this cycle, two equivalents of bicarbonate are fixed by the action of two enzymes: the Acetyl-CoA carboxylase catalyzes the carboxylation of the Acetyl-CoA to Malonyl-CoA and Propionyl-CoA carboxylase catalyses

1679-546: Is cyclic due to the regeneration of the oxaloacetate. The bacteria Gammaproteobacteria and Riftia pachyptila switch from the Calvin-Benson cycle to the rTCA cycle in response to concentrations of H 2 S . The reductive acetyl CoA pathway (CoA) pathway, also known as the Wood-Ljungdahl pathway uses CO 2 as electron acceptor and carbon source, and H 2 as an electron donor to form acetic acid. This metabolism

1752-440: Is determined by the pH , salinity , nutrients, conductivity , soil composition, hardness , and the sources of water. Water chemistry varies across landscapes and climatic regions. Wetlands are generally minerotrophic (waters contain dissolved materials from soils) with the exception of ombrotrophic bogs that are fed only by water from precipitation. Because bogs receive most of their water from precipitation and humidity from

1825-440: Is essential for comprehending ecosystem dynamics , climate regulation, and the sustainability of life on Earth. Organisms that grow by fixing carbon, such as most plants and algae , are called autotrophs . These include photoautotrophs (which use sunlight) and lithoautotrophs (which use inorganic oxidation ). Heterotrophs , such as animals and fungi , are not capable of carbon fixation but are able to grow by consuming

1898-412: Is hydrology, or flooding . The duration of flooding or prolonged soil saturation by groundwater determines whether the resulting wetland has aquatic, marsh or swamp vegetation . Other important factors include soil fertility, natural disturbance, competition, herbivory , burial, and salinity. When peat from dead plants accumulates, bogs and fens develop. Wetland hydrology is associated with

1971-459: Is immediately lost to the atmosphere; a significant portion is retained in the soil through processes collectively known as soil carbon sequestration. Soil microbes, particularly bacteria and fungi, play a pivotal role in this process by incorporating decomposed organic carbon into their biomass or by facilitating the formation of stable organic compounds, such as humus and soil organic matter. One key mechanism by which soil microbes sequester carbon

2044-455: Is incorporated in their metabolism. Notably pyruvate carboxylase consumes carbon dioxide (as bicarbonate ions) as part of gluconeogenesis , and carbon dioxide is consumed in various anaplerotic reactions . 6-phosphogluconate dehydrogenase catalyzes the reductive carboxylation of ribulose 5-phosphate to 6-phosphogluconate in E. coli under elevated CO 2 concentrations. Some carboxylases , particularly RuBisCO , preferentially bind

2117-452: Is only found in certain archaea and accounts for 80% of global methanogenesis, is also based on the reductive acetyl CoA pathway. The Carbon Monoxide Dehydrogenase / Acetyl-CoA Synthase is the oxygen-sensitive enzyme that permits the reduction of CO 2 to CO and the synthesis of acetyl-CoA in several reactions. One branch of this pathway, the methyl branch, is similar but non-homologous between bacteria and archaea. In this branch happens

2190-493: Is the process by which living organisms convert inorganic carbon (particularly carbon dioxide ) to organic compounds . These organic compounds are then used to store energy and as structures for other biomolecules . Carbon is primarily fixed through photosynthesis , but some organisms use chemosynthesis in the absence of sunlight . Chemosynthesis is carbon fixation driven by chemical energy rather than from sunlight.   The process of biological carbon fixation plays

2263-451: Is the only place in the world where both crocodiles and alligators coexist. The saltwater crocodile inhabits estuaries and mangroves. Snapping turtles also inhabit wetlands. Birds , particularly waterfowl and waders use wetlands extensively. Mammals of wetlands include numerous small and medium-sized species such as voles , bats , muskrats and platypus in addition to large herbivorous and apex predator species such as

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2336-407: Is through the process of microbial biomass production. Bacteria and fungi assimilate carbon from decomposed organic matter into their cellular structures as they grow and reproduce. This microbial biomass serves as a reservoir for stored carbon in the soil, effectively sequestering carbon from the atmosphere. Additionally, soil microbes contribute to the formation of stable soil organic matter through

2409-545: Is usually saturated with water". More precisely, wetlands are areas where "water covers the soil , or is present either at or near the surface of the soil all year or for varying periods of time during the year, including during the growing season". A patch of land that develops pools of water after a rain storm would not necessarily be considered a "wetland", even though the land is wet. Wetlands have unique characteristics: they are generally distinguished from other water bodies or landforms based on their water level and on

2482-525: Is wide spread within the phylum Bacillota , especially in the Clostridia . The pathway is also used by methanogens , which are mainly Euryarchaeota , and several anaerobic chemolithoautotrophs, such as sulfate-reducing bacteria and archaea. It is probably performed also by the Brocadiales, an order of Planctomycetota that oxidize ammonia in anaerobic condition. Hydrogenotrophic methanogenesis , which

2555-588: The Calvin cycle or the reductive citric acid cycle. The Calvin cycle accounts for 90% of biological carbon fixation. Consuming adenosine triphosphate (ATP) and nicotinamide adenine dinucleotide phosphate (NADPH), the Calvin cycle in plants accounts for the predominance of carbon fixation on land. In algae and cyanobacteria, it accounts for the dominance of carbon fixation in the oceans. The Calvin cycle converts carbon dioxide into sugar, as triose phosphate (TP), which

2628-460: The Campylobacterota . This feature allows primary production in the ocean's aphotic environments , or "dark primary production." Without it, there would be no primary production in aphotic environments, which would lead to habitats without life. The cycle involves the biosynthesis of acetyl-CoA from two molecules of CO 2 . The key steps of the reverse Krebs cycle are: This pathway

2701-492: The Millennium Ecosystem Assessment from 2005. Methods exist for assessing wetland ecological health . These methods have contributed to wetland conservation by raising public awareness of the functions that wetlands can provide. Since 1971, work under an international treaty seeks to identify and protect " wetlands of international importance ." A simplified definition of wetland is "an area of land that

2774-586: The Nunavut , which are federal Crown land. They are demarcated to prevent hunting within their boundaries, a policy which is enforced. Hunting is permitted in adjacent land. Southern James Bay lies within the flat sedimentary basin of the Hudson Bay lowland in the Hudson Plains ecozone , and its coast is "characterised by a sequence of mudflats , intertidal marshes and supertidal meadow-marshes, which grade through

2847-666: The Pantanal in South America, and the Sundarbans in the Ganges - Brahmaputra delta. Wetlands contribute many ecosystem services that benefit people. These include for example water purification , stabilization of shorelines, storm protection and flood control . In addition, wetlands also process and condense carbon (in processes called carbon fixation and sequestration ), and other nutrients and water pollutants . Wetlands can act as

2920-444: The atmosphere , their water usually has low mineral ionic composition. In contrast, wetlands fed by groundwater or tides have a higher concentration of dissolved nutrients and minerals. Fen peatlands receive water both from precipitation and ground water in varying amounts so their water chemistry ranges from acidic with low levels of dissolved minerals to alkaline with high accumulation of calcium and magnesium . Salinity has

2993-576: The beavers , coypu , swamp rabbit , Florida panther , jaguar , and moose . Wetlands attract many mammals due to abundant seeds, berries, and other vegetation as food for herbivores, as well as abundant populations of invertebrates, small reptiles and amphibians as prey for predators. Invertebrates of wetlands include aquatic insects such as dragonflies , aquatic bugs and beetles , midges, mosquitos , crustaceans such as crabs, crayfish, shrimps, microcrustaceans, mollusks like clams, mussels, snails and worms. Invertebrates comprise more than half of

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3066-517: The cyanobacteria . It also fixes carbon in the anoxygenic photosynthesis in one type of Pseudomonadota called purple bacteria , and in some non-phototrophic Pseudomonadota. Of the other autotrophic pathways, two are known only in bacteria (the reductive citric acid cycle and the 3-hydroxypropionate cycle ), two only in archaea (two variants of the 3-hydroxypropionate cycle), and one in both bacteria and archaea (the reductive acetyl CoA pathway ). Sulfur- and hydrogen-oxidizing bacteria often use

3139-619: The Okavango River inland delta, the Kafue River flood plain, the Lake Bangweulu flood plain (Africa), Mississippi River (US), Amazon River (South America), Yangtze River (China), Danube River (Central Europe) and Murray-Darling River (Australia). Groundwater replenishment can be achieved for example by marsh , swamp , and subterranean karst and cave hydrological systems. The surface water visibly seen in wetlands only represents

3212-511: The Southeastern US, mallines of Argentina, Mediterranean seasonal ponds of Europe and California, turloughs of Ireland, billabongs of Australia, among many others. Wetlands are found throughout the world in different climates. Temperatures vary greatly depending on the location of the wetland. Many of the world's wetlands are in the temperate zones , midway between the North or South Poles and

3285-494: The US, the best known classifications are the Cowardin classification system and the hydrogeomorphic (HGM) classification system. The Cowardin system includes five main types of wetlands: marine (ocean-associated), estuarine (mixed ocean- and river-associated), riverine (within river channels), lacustrine (lake-associated) and palustrine (inland nontidal habitats). Peatlands are

3358-610: The United States' commercial fish and shellfish stocks depend solely on estuaries to survive. Amphibians such as frogs and salamanders need both terrestrial and aquatic habitats in which to reproduce and feed. Because amphibians often inhabit depressional wetlands like prairie potholes and Carolina bays, the connectivity among these isolated wetlands is an important control of regional populations. While tadpoles feed on algae, adult frogs forage on insects. Frogs are sometimes used as an indicator of ecosystem health because their thin skin permits absorption of nutrients and toxins from

3431-434: The accumulation of atmospheric CO2 and mitigate climate change but also enhances soil fertility, water retention, and nutrient cycling , thereby supporting plant growth and ecosystem productivity. Consequently, understanding the role of soil microbes in biological carbon fixation is essential for managing soil health , mitigating climate change, and promoting sustainable land management practices. Biological carbon fixation

3504-404: The activity of microorganisms, such as bacteria and fungi. These soil microbes play a crucial role in the global carbon cycle by sequestering carbon from decomposed organic matter and recycling it back into the soil, thereby contributing to soil fertility and ecosystem productivity.   In soil environments, organic matter derived from dead plant and animal material undergoes decomposition ,

3577-665: The carbon fixed by autotrophs or other heterotrophs. Six natural or autotrophic carbon fixation pathways are currently known. They are the: i) Calvin-Benson-Bassham (Calvin Cycle), ii) Reverse Krebs (rTCA) cycle, iii) the reductive acetyl-CoA (Wood-Ljungdahl pathway), iv) 3-hydroxy propionate [3-HP] bicycle , v) 3-hydroypropionate/4- hydroxybutyrate (3-HP/4-HB) cycle, and vi) the dicarboxylate/ 4-hydroxybutyrate (DC/4-HB) cycle. "Fixed carbon," "reduced carbon," and "organic carbon" may all be used interchangeably to refer to various organic compounds. The primary form of fixed inorganic carbon

3650-444: The carboxylation of propionyl-CoA to methylamalonyl-CoA. From this point a series of reactions lead to the formation of glyoxylate which will thus become part of the second cycle. In the second cycle, glyoxylate is approximately one equivalent of propionyl-CoA forming methylamalonyl-CoA. This, in turn, is then converted through a series of reactions into citramalyl-CoA. The citramalyl-CoA is split into pyruvate and Acetyl-CoA thanks to

3723-701: The complex: the 14.6 square kilometres (5.6 sq mi) Moose River Bird Sanctuary is at the mouth of the Moose River , and the larger 238.3 square kilometres (92.0 sq mi) Hannah Bay Bird Sanctuary on the eastern coast of Hannah Bay at the mouth of the Harricana River . As many as 75,000 geese of various species may simultaneously use the staging area in autumn, and large populations of ducks are also easily observed. "Substantial numbers of diving sea ducks occur offshore." The sites are located on Ontario Crown land, except for outlying islands that are part of

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3796-422: The enzyme MMC lyase. At this point the pyruvate is released, while the Acetyl-CoA is reused and carboxylated again at Malonyl-CoA thus reconstituting the cycle. A total of 19 reactions are involved in 3-hydroxypropionate bicycle and 13 multifunctional enzymes are used. The multifunctionality of these enzymes is an important feature of this pathway which thus allows the fixation of three bicarbonate molecules. It

3869-622: The equator. In these zones, summers are warm and winters are cold, but temperatures are not extreme. In subtropical zone wetlands, such as along the Gulf of Mexico , average temperatures might be 11 °C (52 °F). Wetlands in the tropics are subjected to much higher temperatures for a large portion of the year. Temperatures for wetlands on the Arabian Peninsula can exceed 50 °C (122 °F) and these habitats would therefore be subject to rapid evaporation. In northeastern Siberia , which has

3942-494: The following areas: According to the Ramsar Convention: The economic worth of the ecosystem services provided to society by intact, naturally functioning wetlands is frequently much greater than the perceived benefits of converting them to 'more valuable' intensive land use – particularly as the profits from unsustainable use often go to relatively few individuals or corporations, rather than being shared by society as

4015-464: The interface between truly terrestrial ecosystems and aquatic systems, making them inherently different from each other, yet highly dependent on both." In environmental decision-making, there are subsets of definitions that are agreed upon to make regulatory and policy decisions. Under the Ramsar international wetland conservation treaty , wetlands are defined as follows: An ecological definition of

4088-462: The known animal species in wetlands, and are considered the primary food web link between plants and higher animals (such as fish and birds). Depending on a wetland's geographic and topographic location, the functions it performs can support multiple ecosystem services , values, or benefits. United Nations Millennium Ecosystem Assessment and Ramsar Convention described wetlands as a whole to be of biosphere significance and societal importance in

4161-522: The lighter carbon stable isotope carbon-12 over the heavier carbon-13 . This is known as carbon isotope discrimination and results in carbon-12 to carbon-13 ratios in the plant that are higher than in the free air. Measurement of this ratio is important in the evaluation of water use efficiency in plants, and also in assessing the possible or likely sources of carbon in global carbon cycle studies. In addition to photosynthetic and chemosynthetic processes, biological carbon fixation occurs in soil through

4234-424: The main choice for chemolithoautotrophs limited in energy and living in anaerobic conditions. The 3-Hydroxypropionate bicycle , also known as 3-HP/malyl-CoA cycle, discovered only in 1989, is utilized by green non-sulfur phototrophs of Chloroflexaceae family, including the maximum exponent of this family Chloroflexus auranticus by which this way was discovered and demonstrated. The 3-Hydroxipropionate bicycle

4307-588: The origin of life. Six autotrophic carbon fixation pathways are known: the Calvin Cycle, the Reverse Krebs Cycle, the reductive acetyl-CoA, the 3-HP bicycle, the 3-HP/4-HB cycle, and the DC/4-HB cycles. The organisms the Calvin cycle is found in are plants, algae, cyanobacteria , aerobic proteobacteria, and purple bacteria. The Calvin cycle fixes carbon in the chloroplasts of plants and algae, and in

4380-832: The plants and animals controlled by the wetland hydrology are often additional components of the definitions. Wetlands can be tidal (inundated by tides) or non-tidal. The water in wetlands is either freshwater , brackish , saline , or alkaline . There are four main kinds of wetlands – marsh , swamp , bog , and fen (bogs and fens being types of peatlands or mires ). Some experts also recognize wet meadows and aquatic ecosystems as additional wetland types. Sub-types include mangrove forests , carrs , pocosins , floodplains , peatlands, vernal pools , sinks , and many others. The following three groups are used within Australia to classify wetland by type: Marine and coastal zone wetlands, inland wetlands and human-made wetlands. In

4453-409: The reduction of CO 2 to a methyl residue bound to a cofactor. The intermediates are formate for bacteria and formyl-methanofuran for archaea, and also the carriers, tetrahydrofolate and tetrahydropterins respectively in bacteria and archaea, are different, such as the enzymes forming the cofactor-bound methyl group. Otherwise, the carbonyl branch is homologous between the two domains and consists of

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4526-493: The reduction of another molecule of CO 2 to a carbonyl residue bound to an enzyme, catalyzed by the CO dehydrogenase/acetyl-CoA synthase. This key enzyme is also the catalyst for the formation of acetyl-CoA starting from the products of the previous reactions, the methyl and the carbonyl residues. This carbon fixation pathway requires only one molecule of ATP for the production of one molecule of pyruvate, which makes this process one of

4599-429: The soil and underlying rock into aquifers which are the source of much of the world's drinking water . Wetlands can also act as recharge areas when the surrounding water table is low and as a discharge zone when it is high. Mangroves , coral reefs , salt marsh can help with shoreline stabilization and storm protection. Tidal and inter-tidal wetland systems protect and stabilize coastal zones. Coral reefs provide

4672-537: The soil of wetlands. Anaerobic and aerobic respiration in the soil influences the nutrient cycling of carbon, hydrogen, oxygen, and nitrogen, and the solubility of phosphorus thus contributing to the chemical variations in its water. Wetlands with low pH and saline conductivity may reflect the presence of acid sulfates and wetlands with average salinity levels can be heavily influenced by calcium or magnesium. Biogeochemical processes in wetlands are determined by soils with low redox potential. The life forms of

4745-412: The soils and plants at higher elevations. Plants and animals may vary within a wetland seasonally or in response to flood regimes. There are four main groups of hydrophytes that are found in wetland systems throughout the world. Submerged wetland vegetation can grow in saline and fresh-water conditions. Some species have underwater flowers, while others have long stems to allow the flowers to reach

4818-504: The source of the water. For example, marshes are wetlands dominated by emergent herbaceous vegetation such as reeds , cattails and sedges . Swamps are dominated by woody vegetation such as trees and shrubs (although reed swamps in Europe are dominated by reeds, not trees). Mangrove forest are wetlands with mangroves , halophytic woody plants that have evolved to tolerate salty water . Examples of wetlands classified by

4891-515: The sources of water include tidal wetlands , where the water source is ocean tides ); estuaries , water source is mixed tidal and river waters; floodplains , water source is excess water from overflowed rivers or lakes; and bogs and vernal ponds , water source is rainfall or meltwater . The world's largest wetlands include the Amazon River basin , the West Siberian Plain ,

4964-406: The spatial and temporal dispersion, flow, and physio-chemical attributes of surface and ground waters. Sources of hydrological flows into wetlands are predominantly precipitation , surface water (saltwater or freshwater), and groundwater. Water flows out of wetlands by evapotranspiration , surface flows and tides , and subsurface water outflow. Hydrodynamics (the movement of water through and from

5037-709: The surface of the water. When trees and shrubs comprise much of the plant cover in saturated soils, those areas in most cases are called swamps . The upland boundary of swamps is determined partly by water levels. This can be affected by dams Some swamps can be dominated by a single species, such as silver maple swamps around the Great Lakes . Others, like those of the Amazon basin , have large numbers of different tree species. Other examples include cypress ( Taxodium ) and mangrove swamps. Many species of fish are highly dependent on wetland ecosystems. Seventy-five percent of

5110-490: The surface. Submerged species provide a food source for native fauna, habitat for invertebrates, and also possess filtration capabilities. Examples include seagrasses and eelgrass . Floating water plants or floating vegetation are usually small, like those in the Lemnoideae subfamily (duckweeds). Emergent vegetation like the cattails ( Typha spp.), sedges ( Carex spp.) and arrow arum ( Peltandra virginica ) rise above

5183-457: The surrounding environment resulting in increased extinction rates in unfavorable and polluted environmental conditions. Reptiles such as snakes , lizards , turtles , alligators and crocodiles are common in wetlands of some regions. In freshwater wetlands of the Southeastern US, alligators are common and a freshwater species of crocodile occurs in South Florida. The Florida Everglades

5256-451: The synthesis of extracellular polymers , enzymes, and other biochemical compounds . These substances help bind soil particles together, forming aggregates that protect organic carbon from microbial decomposition and physical erosion . Over time, these aggregates accumulate in the soil, resulting in the formation of soil organic matter, which can persist for centuries to millennia. The sequestration of carbon in soil not only helps mitigate

5329-440: The types of plants that live within them. Specifically, wetlands are characterized as having a water table that stands at or near the land surface for a long enough period each year to support aquatic plants . A more concise definition is a community composed of hydric soil and hydrophytes . Wetlands have also been described as ecotones , providing a transition between dry land and water bodies. Wetlands exist "...at

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