25-620: Geraniaceae is a family of flowering plants placed in the order Geraniales . The family name is derived from the genus Geranium . The family includes both the genus Geranium (the cranesbills, or true geraniums) and the garden plants called geraniums, which modern botany classifies as genus Pelargonium , along with other related genera. The family comprises 830 species in five to seven genera. The largest genera are Geranium (430 species), Pelargonium (280 species) and Erodium (80 species). Geraniaceae are herbs or subshrubs. The Sarcocaulon are succulent , but other members of
50-1845: A molecular phylogeny of plants placed the flowering plants in their evolutionary context: Bryophytes [REDACTED] Lycophytes [REDACTED] Ferns [REDACTED] [REDACTED] [REDACTED] The main groups of living angiosperms are: Amborellales [REDACTED] 1 sp. New Caledonia shrub Nymphaeales [REDACTED] c. 80 spp. water lilies & allies Austrobaileyales [REDACTED] c. 100 spp. woody plants Magnoliids [REDACTED] c. 10,000 spp. 3-part flowers, 1-pore pollen, usu. branch-veined leaves Chloranthales [REDACTED] 77 spp. Woody, apetalous Monocots [REDACTED] c. 70,000 spp. 3-part flowers, 1 cotyledon , 1-pore pollen, usu. parallel-veined leaves Ceratophyllales [REDACTED] c. 6 spp. aquatic plants Eudicots [REDACTED] c. 175,000 spp. 4- or 5-part flowers, 3-pore pollen, usu. branch-veined leaves Amborellales Melikyan, Bobrov & Zaytzeva 1999 Nymphaeales Salisbury ex von Berchtold & Presl 1820 Austrobaileyales Takhtajan ex Reveal 1992 Chloranthales Mart. 1835 Canellales Cronquist 1957 Piperales von Berchtold & Presl 1820 Magnoliales de Jussieu ex von Berchtold & Presl 1820 Laurales de Jussieu ex von Berchtold & Presl 1820 Acorales Link 1835 Alismatales Brown ex von Berchtold & Presl 1820 Petrosaviales Takhtajan 1997 Dioscoreales Brown 1835 Pandanales Brown ex von Berchtold & Presl 1820 Liliales Perleb 1826 Asparagales Link 1829 Arecales Bromhead 1840 Poales Small 1903 Zingiberales Grisebach 1854 Commelinales de Mirbel ex von Berchtold & Presl 1820 Francoaceae See text The Francoaceae are
75-522: A small family of flowering plants in the order Geraniales , including the genera Francoa , commonly known as bridal wreaths. The Francoaceae are recognized as a family under various classification schemes. Under the 2009 APG III system the Francoaceae were included within the Melianthaceae. In the 2016 APG IV system the Francoaceae are again recognized as a family, with Melianthaceae included in
100-610: A total of 64 angiosperm orders and 416 families. The diversity of flowering plants is not evenly distributed. Nearly all species belong to the eudicot (75%), monocot (23%), and magnoliid (2%) clades. The remaining five clades contain a little over 250 species in total; i.e. less than 0.1% of flowering plant diversity, divided among nine families. The 25 most species-rich of 443 families, containing over 166,000 species between them in their APG circumscriptions, are: The botanical term "angiosperm", from Greek words angeíon ( ἀγγεῖον 'bottle, vessel') and spérma ( σπέρμα 'seed'),
125-858: Is starting to impact plants and is likely to cause many species to become extinct by 2100. Angiosperms are terrestrial vascular plants; like the gymnosperms, they have roots , stems , leaves , and seeds . They differ from other seed plants in several ways. The largest angiosperms are Eucalyptus gum trees of Australia, and Shorea faguetiana , dipterocarp rainforest trees of Southeast Asia, both of which can reach almost 100 metres (330 ft) in height. The smallest are Wolffia duckweeds which float on freshwater, each plant less than 2 millimetres (0.08 in) across. Considering their method of obtaining energy, some 99% of flowering plants are photosynthetic autotrophs , deriving their energy from sunlight and using it to create molecules such as sugars . The remainder are parasitic , whether on fungi like
150-524: Is a large diversity in floral architecture. Flowers are usually grouped in cymes (e.g. in Geranium ), umbels (e.g. in Pelargonium ) or, more rarely, spikes . Geraniaceae are normally pollinated by insects , but self-pollination is not uncommon. A number of species are pollinated by the fly Moegistorhynchus longirostris . The fruit is a unique schizocarp made of five (or three) achenes , in
175-468: Is a striking vegetative and floral variation. Flowering plant Basal angiosperms Core angiosperms Flowering plants are plants that bear flowers and fruits , and form the clade Angiospermae ( / ˌ æ n dʒ i ə ˈ s p ər m iː / ). The term 'angiosperm' is derived from the Greek words ἀγγεῖον / angeion ('container, vessel') and σπέρμα / sperma ('seed'), meaning that
200-485: The Alismatales grow in marine environments, spreading with rhizomes that grow through the mud in sheltered coastal waters. Some specialised angiosperms are able to flourish in extremely acid or alkaline habitats. The sundews , many of which live in nutrient-poor acid bogs , are carnivorous plants , able to derive nutrients such as nitrate from the bodies of trapped insects. Other flowers such as Gentiana verna ,
225-517: The inverted repeat . Recent comparison of DNA-fragments resulted in the following phylogenetic tree. Hypseocharitaceae Pelargonium Monsonia including Sarcocaulon California Erodium Geranium Most species are found in temperate or warm temperate regions, though some are tropical. Pelargonium has its centre for diversity in the Cape region in South Africa , where there
250-460: The orchids for part or all of their life-cycle, or on other plants , either wholly like the broomrapes, Orobanche , or partially like the witchweeds, Striga . In terms of their environment, flowering plants are cosmopolitan, occupying a wide range of habitats on land, in fresh water and in the sea. On land, they are the dominant plant group in every habitat except for frigid moss-lichen tundra and coniferous forest . The seagrasses in
275-494: The pistil consists of five (less commonly three) merged carpels . The linear stigmas are free, and the ovary is superior. The nectaries are localised at the bases of the antesepalous stamens and are formed by the receptacle. Pelargonium has only one nectary gland on the adaxial side of the flower. It is hidden in a tube-like cavity which is formed by the receptacle. Flower morphology is conserved within Geraniaceae, but there
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#1732859409821300-481: The seeds are enclosed within a fruit. The group was formerly called Magnoliophyta . Angiosperms are by far the most diverse group of land plants with 64 orders , 416 families , approximately 13,000 known genera and 300,000 known species . They include all forbs (flowering plants without a woody stem ), grasses and grass-like plants, a vast majority of broad-leaved trees , shrubs and vines , and most aquatic plants . Angiosperms are distinguished from
325-883: The "Big Five" extinction events in Earth's history, only the Cretaceous–Paleogene extinction event had occurred while angiosperms dominated plant life on the planet. Today, the Holocene extinction affects all kingdoms of complex life on Earth, and conservation measures are necessary to protect plants in their habitats in the wild ( in situ ), or failing that, ex situ in seed banks or artificial habitats like botanic gardens . Otherwise, around 40% of plant species may become extinct due to human actions such as habitat destruction , introduction of invasive species , unsustainable logging , land clearing and overharvesting of medicinal or ornamental plants . Further, climate change
350-813: The dominant group of plants across the planet. Agriculture is almost entirely dependent on angiosperms, and a small number of flowering plant families supply nearly all plant-based food and livestock feed. Rice , maize and wheat provide half of the world's staple calorie intake, and all three plants are cereals from the Poaceae family (colloquially known as grasses). Other families provide important industrial plant products such as wood , paper and cotton , and supply numerous ingredients for beverages , sugar production , traditional medicine and modern pharmaceuticals . Flowering plants are also commonly grown for decorative purposes , with certain flowers playing significant cultural roles in many societies. Out of
375-691: The family generally are not. Leaves are usually lobed or otherwise divided, sometimes peltate , opposite or alternate and usually have stipules . The flowers are generally regular, or symmetrical. They are hermaphroditic , actinomorphic (radially symmetrical, like in Geranium ) or slightly zygomorphic (with a bilateral symmetry, like in Pelargonium ). The calyx and the corolla are both pentamerous (with five lobes), petals and sepals are free and distinct. The androecium consists in two whorls of five stamens each, some of which can be unfertile;
400-635: The flowering plants as an unranked clade without a formal Latin name (angiosperms). A formal classification was published alongside the 2009 revision in which the flowering plants rank as the subclass Magnoliidae. From 1998, the Angiosperm Phylogeny Group (APG) has reclassified the angiosperms, with updates in the APG II system in 2003, the APG III system in 2009, and the APG IV system in 2016. In 2019,
425-567: The flowerstalk. Geraniaceae and Francoaceae are the two families included in the order Geraniales under the Angiosperm Phylogeny Group (APG) classification (APG IV). There has been some uncertainty in the number of genera to be included. Stevens gives seven genera listed here, while Christenhusz and Byng state five genera. Stevens also lists four synonyms of Geranium : Geraniopsis Chrtek Neurophyllodes (A. Gray) O. Degener Robertianum Picard Robertiella Hypseocharis , with between one and three species, which comes from
450-430: The lower half of the five fertile stamens is made much wider by a wing with a rounded top on each side of the narrow higher part of the filament that carries an anther. Geranium only has ten fertile stamens without wings and lacks staminodes, except for G. pusillum that only has five stamens. Monsonia only has fifteen fertile stamens, which are merged at their base into a ring or merged at their base in trios with
475-412: The lower part the achenes are inside the calyx, while the upper part (the stylar beak) is the style of the flower, looking like a kind of long beak over the achenes. When the fruit is mature the style breaks into five (or three) hygroscopically active (ready to absorb water) bristles that curl, causing the achenes to be released. California lacks filaments without anthers (called staminodes), but
500-518: The manner of vines or lianas . The number of species of flowering plants is estimated to be in the range of 250,000 to 400,000. This compares to around 12,000 species of moss and 11,000 species of pteridophytes . The APG system seeks to determine the number of families , mostly by molecular phylogenetics . In the 2009 APG III there were 415 families. The 2016 APG IV added five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, for
525-444: The middle filament longer than the others, except for M. brevirostrata with only five stamens. Erodium has five staminodes and five fertile stamens, without wings. Pelargonium has ten filaments without wings, between two and seven of which are topped by anthers, while the remaining three to eight are staminodes lacking anthers, but it can easily be distinguished by having only one narrow tube-like nectary inside what looks like
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#1732859409821550-554: The other major seed plant clade, the gymnosperms , by having flowers , xylem consisting of vessel elements instead of tracheids , endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous , over 300 million years ago. In the Cretaceous , angiosperms diversified explosively , becoming
575-599: The south-west Andean region of South America , is considered the sister to the rest of the family. Some authors separate Hyspeocharis as a monogeneric family Hypseocharitaceae , while older sources placed it in the Oxalidaceae . The genus Rhynchotheca has also been separated into the Vivianiaceae . The Geraniaceae have a number of genetic features unique amongst angiosperms, including highly rearranged plastid genomes differing in gene content, order and expansion of
600-568: The spring gentian, are adapted to the alkaline conditions found on calcium -rich chalk and limestone , which give rise to often dry topographies such as limestone pavement . As for their growth habit , the flowering plants range from small, soft herbaceous plants , often living as annuals or biennials that set seed and die after one growing season, to large perennial woody trees that may live for many centuries and grow to many metres in height. Some species grow tall without being self-supporting like trees by climbing on other plants in
625-483: Was coined in the form "Angiospermae" by Paul Hermann in 1690, including only flowering plants whose seeds were enclosed in capsules. The term angiosperm fundamentally changed in meaning in 1827 with Robert Brown , when angiosperm came to mean a seed plant with enclosed ovules. In 1851, with Wilhelm Hofmeister 's work on embryo-sacs, Angiosperm came to have its modern meaning of all the flowering plants including Dicotyledons and Monocotyledons. The APG system treats
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