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90-622: The Garfagnina is a cattle breed from Tuscany in central Italy. It is one of the 16 minor Italian cattle breeds of limited diffusion recognised and protected by the Ministero delle Politiche Agricole Alimentari e Forestali , the Italian ministry of agriculture. This cattle -related article is a stub . You can help Misplaced Pages by expanding it . Cattle Cattle ( Bos taurus ) are large, domesticated , bovid ungulates widely kept as livestock . They are prominent modern members of

180-515: A dominance hierarchy . This is maintained in several ways. Cattle often engage in mock fights where they test each other's strength in a non-aggressive way. Licking is primarily performed by subordinates and received by dominant animals. Mounting is a playful behavior shown by calves of both sexes and by bulls and sometimes by cows in estrus, however, this is not a dominance related behavior as has been found in other species. Dominance-associated aggressiveness does not correlate with rank position, but

270-449: A feedlot for "45 days or more in a 12-month period". Historically, the cattle population of Britain rose from 9.8 million in 1878 to 11.7 million in 1908, but beef consumption rose much faster. Britain became the "stud farm of the world" exporting livestock to countries where there were no indigenous cattle. In 1929 80% of the meat trade of the world was products of what were originally English breeds. There were nearly 70 million cattle in

360-415: A cow is about nine months long. The ratio of male to female offspring at birth is approximately 52:48. A cow's udder has two pairs of mammary glands or teats. Farms often use artificial insemination , the artificial deposition of semen in the female's genital tract ; this allows farmers to choose from a wide range of bulls to breed their cattle. Estrus too may be artificially induced to facilitate

450-414: A dairy, which may be onsite at the farm or the milk may be shipped to a dairy plant for eventual sale of a dairy product. Lactation is induced in heifers and spayed cows by a combination of physical and psychological stimulation, by drugs, or by a combination of those methods. For mother cows to continue producing milk, they give birth to one calf per year. If the calf is male, it is generally slaughtered at

540-404: A diurnal pattern, with a stronger preference for clover in the morning, and the proportion of grass increasing towards the evening. When grazing, cattle vary several aspects of their bite, i.e. tongue and jaw movements, depending on characteristics of the plant they are eating. Bite area decreases with the density of the plants but increases with their height. Bite area is determined by the sweep of

630-619: A learning experiment had higher heart rates, indicating an emotional reaction to their own learning. After separation from their mothers, Holstein calves react, indicating low mood. Similarly, after hot-iron dehorning , calves react to the post-operative pain. The position of the ears has been used as an indicator of emotional state. Cattle can tell when other cattle are stressed by the chemicals in their urine. Cattle are gregarious , and even short-term isolation causes psychological stress . When heifers are isolated, vocalizations, heart rate and plasma cortisol all increase. When visual contact

720-504: A longer period of time than horses . Oxen are used worldwide, especially in developing countries . There are some 11 million draft oxen in sub-Saharan Africa, while in 1998 India had over 65 million oxen. At the start of the 21st century, about half the world's crop production depended on land preparation by draft animals. Cattle are not often kept solely for hides, and they are usually a by-product of beef production. Hides are used mainly for leather products such as shoes. In 2012, India

810-633: A misfolded brain protein , in contaminated meat. Among the intestinal parasites of cattle are Paramphistomum flukes, affecting the rumen, and hookworms in the small intestine. Climate change is expected to exacerbate heat stress in cattle, and for longer periods. Heat-stressed cattle may experience accelerated breakdown of adipose tissue by the liver, causing lipidosis . Cattle eat less when heat stressed, resulting in ruminal acidosis , which can lead to laminitis . Cattle can attempt to deal with higher temperatures by panting more often ; this rapidly decreases carbon dioxide concentrations at

900-668: A mix of taurine cattle, zebu, and yak. The aurochs originally ranged throughout Europe, North Africa, and much of Asia. In historical times, its range became restricted to Europe, and the last known individual died in Mazovia , Poland, around 1627. Breeders have attempted to recreate a similar appearance to the aurochs by crossing traditional types of domesticated cattle, producing the Heck breed . A group of taurine-type cattle exist in Africa; they either represent an independent domestication event or were

990-768: A powerful greenhouse gas, as a byproduct of enteric fermentation , with each cow belching out 100kg a year. Additional methane is produced by anaerobic fermentation of stored manure . The FAO estimates that in 2015 around 7% of global greenhouse gas emissions were due to cattle, but this is uncertain. Reducing methane emissions quickly helps limit climate change . Concentrated animal feeding operations in particular produce substantial amounts of wastewater and manure, which can cause environmental harms such as soil erosion, human and animal exposure to toxic chemicals, development of antibiotic resistant bacteria and an increase in E. coli contamination. In many world regions, overgrazing by cattle has reduced biodiversity of

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1080-497: A quarter of the world's meat comes from cattle. World cattle meat production in 2021 was 72.3 million tons. Certain breeds of cattle, such as the Holstein-Friesian , are used to produce milk , much of which is processed into dairy products such as butter , cheese , and yogurt . Dairy cattle are usually kept on specialized dairy farms designed for milk production. Most cows are milked twice per day, with milk processed at

1170-428: A range of production and behavioral characteristics for both dairy and beef cattle. Cattle have played a key role in human history , having been domesticated since at least the early neolithic age. Archaeozoological and genetic data indicate that cattle were first domesticated from wild aurochs ( Bos primigenius ) approximately 10,500 years ago. There were two major areas of domestication: one in central Anatolia ,

1260-559: A similar aquatic lifestyle. Hippopotamuses appeared in the late Miocene and occupied Africa and Asia—they never got to the Americas. The camels ( Tylopoda ) were, during large parts of the Cenozoic , limited to North America; early forms like Cainotheriidae occupied Europe. Among the North American camels were groups like the stocky, short-legged Merycoidodontidae . They first appeared in

1350-477: A simple stomach that digests food. Thus, they were grouped together as non-ruminants (Porcine). All other even-toed ungulates have molars with a selenodont construction (crescent-shaped cusps) and have the ability to ruminate , which requires regurgitating food and re-chewing it. Differences in stomach construction indicated that rumination evolved independently between tylopods and ruminants ; therefore, tylopods were excluded from Ruminantia . The taxonomy that

1440-469: A slim build, lanky legs, and a long tail. Their hind legs were much longer than their front legs. The early to middle Eocene saw the emergence of the ancestors of most of today's mammals. Two formerly widespread, but now extinct, families of even-toed ungulates were Entelodontidae and Anthracotheriidae . Entelodonts existed from the middle Eocene to the early Miocene in Eurasia and North America. They had

1530-468: A stocky body with short legs and a massive head, which was characterized by two humps on the lower jaw bone. Anthracotheres had a large, porcine ( pig -like) build, with short legs and an elongated muzzle . This group appeared in the middle Eocene up until the Pliocene , and spread throughout Eurasia, Africa, and North America. Anthracotheres are thought to be the ancestors of hippos, and, likewise, probably led

1620-461: A third independent domestication from the North African aurochs. Whether there have been two or three domestications, European, African, and Asian cattle share much of their genomes both through their species ancestry and through repeated migrations of livestock and genetic material between species, as shown in the diagram. Cattle were originally identified as three separate species: Bos taurus ,

1710-424: A very flexible body, contributing to their speed by increasing their stride length. Many even-toed ungulates have a relatively large head. The skull is elongated and rather narrow; the frontal bone is enlarged near the back and displaces the parietal bone , which forms only part of the side of the cranium (especially in ruminants). Four families of even-toed ungulates have cranial appendages. These Pecora (with

1800-456: A wider study mapped a further 4.2% of the cattle genome. Behavioral traits of cattle can be as heritable as some production traits, and often, the two can be related. The heritability of temperament (response to isolation during handling) has been calculated as 0.36 and 0.46 for habituation to handling. Rangeland assessments show that the heritability of aggressiveness in cattle is around 0.36. Quantitative trait loci have been found for

1890-506: A young age to produce veal . Cows produce milk until three weeks before birth. Over the last fifty years, dairy farming has become more intensive to increase the yield of milk produced by each cow. The Holstein-Friesian is the breed of dairy cow most common in the UK, Europe and the United States. It has been bred selectively to produce the highest yields of milk of any cow. The average in the UK

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1980-725: Is a diurnal rhythm in suckling, peaking at roughly 6am, 11:30am, and 7pm. Under natural conditions, calves stay with their mother until weaning at 8 to 11 months. Heifer and bull calves are equally attached to their mothers in the first few months of life. Cattle have a variety of cognitive abilities. They can memorize the locations of multiple food sources, and can retain memories for at least 48 days. Young cattle learn more quickly than adults, and calves are capable of discrimination learning, distinguishing familiar and unfamiliar animals, and between humans, using faces and other cues. Calves prefer their own mother's vocalizations to those of an unfamiliar cow. Vocalizations provide information on

2070-471: Is around 22 litres per day. Dairy is a large industry worldwide. In 2023, the 27 European Union countries produced 143 million tons of cow's milk; the United States 104.1 million tons; and India 99.5 million tons. India further produces 94.4 million tons of buffalo milk , making it (in 2023) the world's largest milk producer; its dairy industry employs some 80 million people. Oxen are cattle trained as draft animals . Oxen can pull heavier loads and for

2160-474: Is better than that of horses, but worse at localising sounds than goats, and much worse than dogs or humans. They can distinguish between live and recorded human speech. Olfaction probably plays a large role in their social life, indicating social and reproductive status. Cattle can tell when other animals are stressed by smelling the alarm chemicals in their urine. Cattle can be trained to recognise conspecific individuals using olfaction only. Cattle live in

2250-428: Is closely related to rank distance between individuals. The horns of cattle are honest signals used in mate selection. Horned cattle attempt to keep greater distances between themselves and have fewer physical interactions than hornless cattle, resulting in more stable social relationships. In calves, agonistic behavior becomes less frequent as space allowance increases, but not as group size changes, whereas in adults,

2340-496: Is contentious, and authorities such as the American Society of Mammalogists treat these taxa as separate species. Complicating the matter is the ability of cattle to interbreed with other closely related species. Hybrid individuals and even breeds exist, not only between taurine cattle and zebu (such as the sanga cattle ( Bos taurus africanus x Bos indicus ), but also between one or both of these and some other members of

2430-471: Is more adapted to a squeezing mastication , which is characteristic of omnivores . Camels and ruminants have fewer teeth; there is often a yawning diastema , a designated gap in the teeth where the molars are aligned for crushing plant matter. The incisors are often reduced in ruminants, and are completely absent in the upper jaw . The canines are enlarged and tusk-like in the Suina, and are used for digging in

2520-488: Is now northern Syria. Although European cattle are largely descended from the taurine lineage, gene flow from African cattle (partially of indicine origin) contributed substantial genomic components to both southern European cattle breeds and their New World descendants. A study on 134 breeds showed that modern taurine cattle originated from Africa, Asia, North and South America, Australia, and Europe. Some researchers have suggested that African taurine cattle are derived from

2610-496: Is re-instated, vocalizations rapidly decline; heart rate decreases more rapidly if the returning cattle are familiar to the previously isolated individual. Mirrors have been used to reduce stress in isolated cattle. The average sleep time of a domestic cow is about 4 hours a day. Cattle do have a stay apparatus , but do not sleep standing up; they lie down to sleep deeply. In 2009, the National Institutes of Health and

2700-499: Is some 25–30 years. Beef cattle go to slaughter at around 18 months, and dairy cows at about five years. Cattle are ruminants , meaning their digestive system is highly specialized for processing plant material such as grass rich in cellulose , a tough carbohydrate polymer which many animals cannot digest. They do this in symbiosis with micro-organisms – bacteria , fungi , and protozoa – that possess cellulases , enzymes that split cellulose into its constituent sugars . Among

2790-409: Is spread by midges . Psoroptic mange is a disabling skin condition caused by mites . Bovine tuberculosis is caused by a bacterium; it causes disease in humans and in wild animals such as deer and badgers. Foot-and-mouth disease is caused by a virus, affects a range of hoofed livestock and is highly contagious. Bovine spongiform encephalopathy is a neurodegenerative disease spread by a prion ,

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2880-624: Is the number of animals within a specified area. High stocking density can affect cattle health, welfare, productivity, and feeding behaviour. Densely-stocked cattle feed more rapidly and lie down sooner, increasing the risk of teat infection, mastitis, and embryo loss . The stress and negative health impacts induced by high stocking density such as in concentrated animal feeding operations or feedlots , auctions, and transport may be detrimental to cattle welfare. To produce milk from dairy cattle , most calves are separated from their mothers soon after birth and fed milk replacement in order to retain

2970-704: Is therefore focused on anthracotheres (family Anthracotheriidae); one dating from the Eocene to Miocene was declared to be "hippo-like" upon discovery in the 19th century. A study from 2005 showed that the anthracotheres and hippopotamuses had very similar skulls , but differed in the adaptations of their teeth. It was nevertheless believed that cetaceans and anthracothereres descended from a common ancestor, and that hippopotamuses developed from anthracotheres. A study published in 2015 confirmed this, but also revealed that hippopotamuses were derived from older anthracotherians. The newly introduced genus Epirigenys from Eastern Africa

3060-525: Is thus the sister group of hippos. Linnaeus postulated a close relationship between camels and ruminants as early as the mid-1700s. Henri de Blainville recognized the similar anatomy of the limbs of pigs and hippos, and British zoologist Richard Owen coined the term "even-toed ungulates" and the scientific name "Artiodactyla" in 1848. Internal morphology (mainly the stomach and the molars) were used for classification. Suines (including pigs ) and hippopotamuses have molars with well-developed roots and

3150-448: The Levant and Western Iran , giving rise to the taurine line, and a second in the area that is now Pakistan, resulting in the indicine line. Modern mitochondrial DNA variation indicates the taurine line may have arisen from as few as 80 aurochs tamed in the upper reaches of Mesopotamia near the villages of Çayönü Tepesi in what is now southeastern Turkey, and Dja'de el-Mughara in what

3240-914: The Levant and Western Iran . A separate domestication event occurred in the Indian subcontinent , which gave rise to zebu. There were over 940 million cattle in the world by 2022. Cattle are responsible for around 7% of global greenhouse gas emissions . They were one of the first domesticated animals to have a fully-mapped genome . The term cattle was borrowed from Anglo-Norman catel (replacing native Old English terms like kine , now considered archaic, poetic, or dialectal), itself from Medieval Latin capitale 'principal sum of money, capital', itself derived in turn from Latin caput 'head'. Cattle originally meant movable personal property , especially livestock of any kind, as opposed to real property (the land, which also included wild or small free-roaming animals such as chickens—they were sold as part of

3330-674: The Old World , exist today only in the Americas . South America was settled by even-toed ungulates only in the Pliocene, after the land bridge at the Isthmus of Panama formed some three million years ago. With only the peccaries, lamoids (or llamas ), and various species of capreoline deer , South America has comparatively fewer artiodactyl families than other continents, except Australia, which has no native species. The classification of artiodactyls

3420-459: The US Department of Agriculture reported having mapped the bovine genome . Cattle have some 22,000 genes, of which 80% are shared with humans; they have about 1000 genes that they share with dogs and rodents, but not with humans. Using this bovine "HapMap", researchers can track the differences between breeds that affect meat and milk yields. Early research focused on Hereford genetic sequences;

3510-520: The genus Bos – yaks (the dzo or yattle ), banteng , and gaur . Hybrids such as the beefalo breed can even occur between taurine cattle and either species of bison , leading some authors to consider them part of the genus Bos , as well. The hybrid origin of some types may not be obvious – for example, genetic testing of the Dwarf Lulu breed, the only taurine-type cattle in Nepal, found them to be

3600-507: The hind legs have a reduction in the number of toes to three. The central axis of the leg is between the third and fourth toe. The first toe is missing in modern artiodactyls, and can only be found in now-extinct genera. The second and fifth toes are adapted differently between species: When camels have only two toes present, the claws are transformed into nails (while both are made of keratin , claws are curved and pointed while nails are flat and dull). These claws consist of three parts:

3690-587: The miniature Zebu are kept as pets . Taurine cattle are widely distributed across Europe and temperate areas of Asia, the Americas, and Australia. Zebus are found mainly in India and tropical areas of Asia, America, and Australia. Sanga cattle are found primarily in sub-Saharan Africa . These types, sometimes classified as separate species or subspecies, are further divided into over 1,000 recognized breeds . Around 10,500 years ago, taurine cattle were domesticated from wild aurochs progenitors in central Anatolia ,

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3780-413: The rumen , reticulum , omasum , and abomasum . The rumen is the largest compartment and it harbours the most important parts of the microbiome. The reticulum, the smallest compartment, is known as the "honeycomb". The omasum's main function is to absorb water and nutrients from the digestible feed. The abomasum has a similar function to the human stomach. Cattle regurgitate and re-chew their food in

3870-412: The ruminants as the sister group of cetaceans. Subsequent studies established the close relationship between hippopotamuses and cetaceans; these studies were based on casein genes , SINEs , fibrinogen sequences, cytochrome and rRNA sequences, IRBP (and vWF ) gene sequences, adrenergic receptors , and apolipoproteins . In 2001, the fossil limbs of a Pakicetus (amphibioid cetacean

3960-529: The European or "taurine" cattle (including similar types from Africa and Asia); Bos indicus , the Indicine or "zebu" ; and the extinct Bos primigenius , the aurochs . The aurochs is ancestral to both zebu and taurine cattle. They were later reclassified as one species, Bos taurus , with the aurochs ( B. t. primigenius ), zebu ( B. t. indicus ), and taurine ( B. t. taurus ) cattle as subspecies. However, this taxonomy

4050-452: The US by the early 1930s. Cattle have the largest biomass of any animal species on Earth, at roughly 400 million tonnes, followed closely by Antarctic krill at 379 million tonnes and humans at 373 million tonnes. In 2023, the countries with the most cattle were India with 307.5 million (32.6% of the total), Brazil with 194.4 million, and China with 101.5 million, out of a total of 942.6 million in

4140-457: The age, sex, dominance status and reproductive status of the caller, and may indicate estrus in cows and competitive display in bulls. Cows can categorize images as familiar and unfamiliar individuals. Cloned calves from the same donor form subgroups, suggesting that kin discrimination may be a basis of grouping behaviour. Cattle use visual/brain lateralisation when scanning novel and familiar stimuli. They prefer to view novel stimuli with

4230-670: The animals. In Spain, the Running of the bulls faces opposition due to the stress and injuries incurred by the bulls during the event. Artiodactyl Cetartiodactyla Montgelard et al. 1997 Artiodactyls are placental mammals belonging to the order Artiodactyla ( / ˌ ɑːr t i oʊ ˈ d æ k t ɪ l ə / AR -tee-oh- DAK -tih-lə , from Ancient Greek ἄρτιος , ártios  'even' and δάκτυλος , dáktylos  'finger, toe'). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in

4320-676: The available food; foraging velocity decreases and intake rate increases in areas of abundant palatable forage. Cattle avoid grazing areas contaminated by the faeces of other cattle more strongly than they avoid areas contaminated by sheep, but they do not avoid pasture contaminated by rabbits. In cattle, temperament or behavioral disposition can affect productivity, overall health, and reproduction. Five underlying categories of temperament traits have been proposed: shyness–boldness, exploration–avoidance, activity, aggressiveness , and sociability. There are many indicators of emotion in cattle. Holstein–Friesian heifers that had made clear improvements in

4410-474: The bulls weigh 1,000–1,200 kg (2,200–2,600 lb). Before 1790, beef cattle averaged only 160 kg (350 lb) net. Thereafter, weights climbed steadily. Cattle breeds vary widely in size; the tallest and heaviest is the Chianina , where a mature bull may be up to 1.8 m (5 ft 11 in) at the shoulder, and may reach 1,280 kg (2,820 lb) in weight. The natural life of domestic cattle

4500-481: The climate warms, increasing the risk of tick-borne diseases. Both beef and milk production are likely to experience declines due to climate change. Cattle health is at once a veterinary issue (for animal welfare and productivity), a public health issue (to limit the spread of disease), and a food safety issue (to ensure meat and dairy products are safe to eat). These concerns are reflected in farming regulations. These rules can become political matters, as when it

4590-500: The cows' milk for human consumption. Animal welfare advocates are critical of this practice , stating that this breaks the natural bond between the mother and her calf. The welfare of veal calves is also a concern. Two sports involving cattle are thought to be cruel by animal welfare groups: rodeos and bullfighting . Such groups oppose rodeo activities including bull riding , calf roping and steer roping , stating that rodeos are unnecessary and cause stress, injury, and death to

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4680-458: The early Eocene (about 53 million years ago). Since these findings almost simultaneously appeared in Europe , Asia , and North America , it is very difficult to accurately determine the origin of artiodactyls. The fossils are classified as belonging to the family Diacodexeidae ; their best-known and best-preserved member is Diacodexis . These were small animals, some as small as a hare , with

4770-609: The early Cenozoic ( Paleocene and Eocene ), which had hooves instead of claws on their feet. Their molars were adapted to a carnivorous diet, resembling the teeth in modern toothed whales , and, unlike other mammals, had a uniform construction. The suspected relations can be shown as follows:   Artiodactyla [REDACTED]   Mesonychia † [REDACTED]   Cetacea [REDACTED] Molecular findings and morphological indications suggest that artiodactyls, as traditionally defined, are paraphyletic with respect to cetaceans. Cetaceans are deeply nested within

4860-419: The exception of the musk deer ), have one of four types of cranial appendages: true horns, antlers , ossicones , or pronghorns . True horns have a bone core that is covered in a permanent sheath of keratin, and are found only in the bovids . Antlers are bony structures that are shed and replaced each year; they are found in deer (members of the family Cervidae ). They grow from a permanent outgrowth of

4950-633: The form of a hoof ). The other three toes are either present, absent, vestigial , or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two orders is that many artiodactyls (except for Suina ) digest plant cellulose in one or more stomach chambers rather than in their intestine (as perissodactyls do). Molecular biology, along with new fossil discoveries, has found that cetaceans ( whales , dolphins , and porpoises ) fall within this taxonomic branch, being most closely related to hippopotamuses . Some modern taxonomists thus apply

5040-1015: The former; the two groups together form a monophyletic taxon, for which the name Cetartiodactyla is sometimes used. Modern nomenclature divides Artiodactyla (or Cetartiodactyla) in four subordinate taxa: camelids (Tylopoda), pigs and peccaries (Suina), ruminants (Ruminantia), and hippos plus cetaceans (Whippomorpha). The presumed lineages within Artiodactyla can be represented in the following cladogram :   Tylopoda (camels) [REDACTED]   Suina (pigs) [REDACTED]   Tragulidae (mouse deer) [REDACTED]   Pecora (horn bearers) [REDACTED]   Hippopotamidae (hippopotamuses) [REDACTED]   Cetacea (whales) [REDACTED] The four summarized Artiodactyla taxa are divided into ten extant families: Although deer, musk deer, and pronghorns have traditionally been summarized as cervids (Cervioidea), molecular studies provide different—and inconsistent—results, so

5130-668: The frontal bone called the pedicle and can be branched, as in the white-tailed deer ( Odocoileus virginianus ), or palmate , as in the moose ( Alces alces ). Ossicones are permanent bone structures that fuse to the frontal or parietal bones during an animal's life and are found only in the Giraffidae . Pronghorns , while similar to horns in that they have keratinous sheaths covering permanent bone cores, are deciduous. All these cranial appendages can serve for posturing, battling for mating privilege, and for defense. In almost all cases, they are sexually dimorphic, and are often found only on

5220-489: The grazed plants and of animals at different trophic levels in the ecosystem . A well documented consequence of overgrazing is woody plant encroachment in rangelands, which significantly reduces the carrying capacity of the land over time. Cattle husbandry practices including branding , castration , dehorning , ear tagging , nose ringing , restraint, tail docking , the use of veal crates, and cattle prods have raised welfare concerns. Stocking density

5310-401: The ground and for defense. In ruminants, the males' upper canines are enlarged and used as a weapon in certain species (mouse deer, musk deer, water deer ); species with frontal weapons are usually missing the upper canines. The lower canines of ruminants resemble the incisors, so that these animals have eight uniform teeth in the frontal part of the lower jaw . The molars of porcine have only

5400-685: The land). The word is a variant of chattel (a unit of personal property) and closely related to capital in the economic sense. The word cow came via Anglo-Saxon cū (plural cȳ ), from Common Indo-European gʷōus ( genitive gʷowés ) 'a bovine animal', cf. Persian : gâv , Sanskrit : go- . In older English sources such as the King James Version of the Bible, cattle often means livestock, as opposed to deer , which are wild. Cattle are large artiodactyls , mammals with cloven hooves , meaning that they walk on two toes,

5490-584: The late Eocene and developed a great diversity of species in North America. Only in the late Miocene or early Pliocene did they migrate from North America into Eurasia. The North American varieties became extinct around 10,000 years ago. Suina (including pigs ) have been around since the Eocene. In the late Eocene or the Oligocene , two families stayed in Eurasia and Africa; the peccaries , which became extinct in

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5580-402: The left eye (using the right brain hemisphere), but the right eye for familiar stimuli. Individual cattle have also been observed to display different personality traits, such as fearfulness and sociability. Vision is the dominant sense; cattle obtain almost half of their information visually. Being prey animals, cattle evolved to look out for predators almost all around, with eyes that are on

5670-407: The limbs are predominantly localized, which ensures that artiodactyls often have very slender legs. A clavicle is never present, and the scapula is very agile and swings back and forth for added mobility when running. The special construction of the legs causes the legs to be unable to rotate, which allows for greater stability when running at high speeds. In addition, many smaller artiodactyls have

5760-634: The males are consistently larger and heavier than the females. In deer, only the males boast antlers, and the horns of bovines are usually small or not present in females. Male Indian antelopes have a much darker coat than females. Almost all even-toed ungulates have fur, with the exception being the nearly hairless hippopotamus. Fur varies in length and coloration depending on the habitat. Species in cooler regions can shed their coat. Camouflaged coats come in colors of yellow, gray, brown, or black tones. Even-toed ungulates bear their name because they have an even number of toes (two or four)—in some peccaries,

5850-459: The males. One exception is the species Rangifer tarandus , known as reindeer in Europe or caribou in North America, where both sexes can grow antlers yearly, though the females' antlers are typically smaller and not always present. There are two trends in terms of teeth within Artiodactyla. The Suina and hippopotamuses have a relatively large number of teeth (with some pigs having 44); their dentition

5940-440: The many bacteria that contribute are Fibrobacter succinogenes , Ruminococcus flavefaciens , and Ruminococcus albus . Cellulolytic fungi include several species of Neocallimastix , while the protozoa include the ciliates Eudiplodinium maggie and Ostracodinium album . If the animal's feed changes over time, the composition of this microbiome changes in response. Cattle have one large stomach with four compartments;

6030-421: The mouse deer, often reaches a body length of only 45 centimeters (18 in) and a weight of 1.5 kilograms (3.3 lb). The largest member, the hippopotamus, can grow up to 5 meters (16 ft) in length and weigh 4.5 metric tons (5 short tons), and the giraffe can grow to be 5.5 meters (18 ft) tall and 4.7 meters (15 ft) in body length. All even-toed ungulates display some form of sexual dimorphism :

6120-948: The name Cetartiodactyla ( / s ɪ ˌ t ɑːr t i oʊ ˈ d æ k t ɪ l ə / ) to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use " even-toed ungulates " to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature. The roughly 270 land-based even-toed ungulate species include pigs , peccaries , hippopotamuses , antelopes , deer , giraffes , camels , llamas , alpacas , sheep , goats and cattle . Many are herbivores, but suids are omnivorous, whereas cetaceans are entirely carnivorous. Artiodactyls are also known by many extinct groups such as anoplotheres , cainotheriids , merycoidodonts , entelodonts , anthracotheres , basilosaurids , and palaeomerycids . Many artiodactyls are of great dietary, economic, and cultural importance to humans. The oldest fossils of even-toed ungulates date back to

6210-647: The number of agonistic encounters increases with group size. Dominance relationships in semi-wild highland cattle are very firm, with few overt aggressive conflicts: most disputes are settled by agonistic (non-aggressive, competitive) behaviors with no physical contact between opponents, reducing the risk of injury. Dominance status depends on age and sex, with older animals usually dominant to young ones and males dominant to females. Young bulls gain superior dominance status over adult cows when they reach about 2 years of age. Cattle eat mixed diets, but prefer to eat approximately 70% clover and 30% grass. This preference has

6300-442: The plate (top and sides), the sole (bottom), and the bale (rear). In general, the claws of the forelegs are wider and blunter than those of the hind legs, and they are farther apart. Aside from camels, all even-toed ungulates put just the tip of the foremost phalanx on the ground. In even-toed ungulates, the bones of the stylopodium (upper arm or thigh bone) and zygopodiums (tibia and fibula) are usually elongated. The muscles of

6390-438: The price of increasing pH , respiratory alkalosis . To deal with this, cattle are forced to shed bicarbonate through urination , at the expense of rumen buffering. These two pathologies can both cause lameness . Another specific risk is mastitis . This worsens as Calliphora blowflies increase in number with continued warming, spreading mastitis-causing bacteria. Ticks too are likely to increase in temperate zones as

6480-437: The process of chewing the cud , like most ruminants. While feeding, cows swallow their food without chewing; it goes into the rumen for storage. Later, the food is regurgitated to the mouth, a mouthful at a time, where the cud is chewed by the molars, grinding down the coarse vegetation to small particles. The cud is then swallowed again and further digested by the micro-organisms in the cow's stomach. The gestation period for

6570-483: The process. Copulation lasts several seconds and consists of a single pelvic thrust . Cows seek secluded areas for calving. Semi-wild Highland cattle heifers first give birth at 2 or 3 years of age, and the timing of birth is synchronized with increases in natural food quality. Average calving interval is 391 days, and calving mortality within the first year of life is 5%. Beef calves suckle an average of 5 times per day, spending some 46 minutes suckling. There

6660-413: The question of phylogenetic systematics of infraorder Pecora (the horned ruminants) for the time being, cannot be answered. Artiodactyls are generally quadrupeds . Two major body types are known: suinids and hippopotamuses are characterized by a stocky body, short legs, and a large head; camels and ruminants, though, have a more slender build and lanky legs. Size varies considerably; the smallest member,

6750-472: The result of crossing taurines domesticated elsewhere with local aurochs, but they are genetically distinct; some authors name them as a separate subspecies, Bos taurus africanus . The only pure African taurine breeds remaining are the N'Dama , Kuri and some varieties of the West African Shorthorn. Feral cattle are those that have been allowed to go wild. Populations exist in many parts of

6840-472: The sequence with that of other living beings—the more similar they are, the more closely they are related. Comparison of even-toed ungulate and cetaceans genetic material has shown that the closest living relatives of whales and hippopotamuses is the paraphyletic group Artiodactyla. Dan Graur and Desmond Higgins were among the first to come to this conclusion, and included a paper published in 1994. However, they did not recognize hippopotamuses and classified

6930-495: The sides of their head rather than the front. This gives them a field of view of 330°, but limits binocular vision (and therefore stereopsis ) to some 30° to 50°, compared to 140° in humans. They are dichromatic , like most mammals. Cattle avoid bitter-tasting foods, selecting sweet foods for energy. Their sensitivity to sour-tasting foods helps them to maintain optimal ruminal pH . They seek out salty foods by taste and smell to maintain their electrolyte balance. Their hearing

7020-659: The size of a wolf) and Ichthyolestes (an early whale the size of a fox) were found in Pakistan. They were both archaeocetes ("ancient whales") from about 48 million years ago (in the Eocene). These findings showed that archaeocetes were more terrestrial than previously thought, and that the special construction of the talus (ankle bone) with a double-rolled joint surface, previously thought to be unique to even-toed ungulates, were also in early cetaceans. The mesonychians , another type of ungulate, did not show this special construction of

7110-583: The subfamily Bovinae and the most widespread species of the genus Bos . Mature female cattle are called cows and mature male cattle are bulls . Young female cattle are called heifers , young male cattle are oxen or bullocks , and castrated male cattle are known as steers . Cattle are commonly raised for meat , for dairy products , and for leather . As draft animals , they pull carts and farm implements . In India , cattle are sacred animals within Hinduism, and may not be killed. Small breeds such as

7200-516: The talus, and thus was concluded to not have the same ancestors as cetaceans. The oldest cetaceans date back to the early Eocene (53 million years ago), whereas the oldest known hippopotamus dates back only to the Miocene (15 million years ago). The hippopotamids are descended from the anthracotheres, a family of semiaquatic and terrestrial artiodactyls that appeared in the late Eocene, and are thought to have resembled small- or narrow-headed hippos. Research

7290-416: The third and fourth digits. Like all bovid species, they can have horns, which are unbranched and are not shed annually. Coloration varies with breed; common colors are black, white, and red/brown, and some breeds are spotted or have mixed colors. Bulls are larger than cows of the same breed by up to a few hundred kilograms. British Hereford cows, for example, weigh 600–800 kg (1,300–1,800 lb), while

7380-419: The tongue; in one study observing 750-kilogram (1,650 lb) steers, bite area reached a maximum of approximately 170 cm (30 sq in). Bite depth increases with the height of the plants. By adjusting their behavior, cattle obtain heavier bites in swards that are tall and sparse compared with short, dense swards of equal mass/area. Cattle adjust other aspects of their grazing behavior in relation to

7470-479: The traditional order Artiodactyla and infraorder Cetacea are sometimes subsumed into the more inclusive Cetartiodactyla taxon. An alternative approach is to include both land-dwelling even-toed ungulates and ocean-dwelling cetaceans in a revised Artiodactyla taxon. In the 1990s, biological systematics used not only morphology and fossils to classify organisms, but also molecular biology . Molecular biology involves sequencing an organism's DNA and RNA and comparing

7560-721: The world, sometimes on small islands. Some, such as Amsterdam Island cattle , Chillingham cattle , and Aleutian wild cattle have become sufficiently distinct to be described as breeds. Cattle are often raised by allowing herds to graze on the grasses of large tracts of rangeland . Raising cattle extensively in this manner allows the use of land that might be unsuitable for growing crops. The most common interactions with cattle involve daily feeding , cleaning and milking . Many routine husbandry practices involve ear tagging , dehorning , loading, medical operations , artificial insemination , vaccinations and hoof care, as well as training for agricultural shows and preparations. Around

7650-516: The world, Fulani husbandry rests on behavioural techniques, whereas in Europe, cattle are controlled primarily by physical means, such as fences . Breeders use cattle husbandry to reduce tuberculosis susceptibility by selective breeding and maintaining herd health to avoid concurrent disease. In the United States, many cattle are raised intensively, kept in concentrated animal feeding operations , meaning there are at least 700 mature dairy cows or at least 1000 other cattle stabled or confined in

7740-406: The world. Cattle are kept on farms to produce meat, milk, and leather, and sometimes to pull carts or farm implements. The meat of adult cattle is known as beef , and that of calves as veal . Other body parts are used as food products, including blood, liver , kidney , heart and oxtail . Approximately 300 million cattle, including dairy animals, are slaughtered each year for food. About

7830-426: Was hotly debated because ocean-dwelling cetaceans evolved from land-dwelling even-toed ungulates. Some semiaquatic even-toed ungulates ( hippopotamuses ) are more closely related to ocean-dwelling cetaceans than to other even-toed ungulates. Phylogenetic classification only recognizes monophyletic taxa; that is, groups that descend from a common ancestor and include all of its descendants. To address this problem,

7920-500: Was proposed in the UK in 2011 that milk from tuberculosis -infected cattle should be allowed to enter the food chain. Cattle disease attracted attention in the 1980s and 1990s when bovine spongiform encephalopathy (mad cow disease) broke out in the United Kingdom . BSE can cross into humans as the deadly variant Creutzfeldt–Jakob disease ; 178 people in the UK had died from it by 2010. The gut flora of cattle produce methane ,

8010-415: Was the world's largest producer of cattle hides. Cattle hides account for around 65% of the world's leather production. Cattle are subject to pests including arthropod parasites such as ticks (which can in turn transmit diseases caused by bacteria and protozoa), and diseases caused by pathogens including bacteria and viruses . Some viral diseases are spread by insects - i.e. bluetongue disease

8100-621: Was widely accepted by the end of the 20th century was:   Suidae [REDACTED]   Hippopotamidae [REDACTED]   Tylopoda [REDACTED]   Tragulidae [REDACTED]   Pecora [REDACTED] Modern cetaceans are highly adapted sea creatures which, morphologically, have little in common with land mammals; they are similar to other marine mammals , such as seals and sea cows , due to convergent evolution . However, they evolved from originally terrestrial mammals. The most likely ancestors were long thought to be mesonychians—large, carnivorous animals from

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