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142-652: The Ediacaran ( / ˌ iː d i ˈ æ k ər ə n , ˌ ɛ d i -/ EE -dee- AK -ər-ən, ED -ee- ) is a geological period of the Neoproterozoic Era that spans 96 million years from the end of the Cryogenian Period at 635 Mya to the beginning of the Cambrian Period at 538.8 Mya. It is the last period of the Proterozoic Eon as well as the last of the so-called " Precambrian supereon", before

284-471: A jigsaw puzzle . Rocks normally form relatively horizontal layers, with each layer younger than the one underneath it. If a fossil is found between two layers whose ages are known, the fossil's age must lie between the two known ages. Because rock sequences are not continuous, but may be broken up by faults or periods of erosion , it is very difficult to match up rock beds that are not directly next to one another. However, fossils of species that survived for

426-510: A chemically distinctive carbonate layer that is referred to as a " cap carbonate ", because it caps glacial deposits. This bed is characterized by an unusual depletion of C that indicates a sudden climatic change at the end of the Marinoan ice age . The lower global boundary stratotype section (GSSP) of the Ediacaran is at the base of the cap carbonate (Nuccaleena Formation), immediately above

568-544: A collision that formed the Moon about 40 million years later, may have cooled quickly enough to have oceans and an atmosphere about 4,440  million years ago . There is evidence on the Moon of a Late Heavy Bombardment by asteroids from 4,000 to 3,800 million years ago . If, as seems likely, such a bombardment struck Earth at the same time, the first atmosphere and oceans may have been stripped away. Paleontology traces

710-602: A common ancestor. Ideally the "family tree" has only two branches leading from each node ("junction"), but sometimes there is too little information to achieve this, and paleontologists have to make do with junctions that have several branches. The cladistic technique is sometimes fallible, as some features, such as wings or camera eyes , evolved more than once, convergently  – this must be taken into account in analyses. Evolutionary developmental biology , commonly abbreviated to "Evo Devo", also helps paleontologists to produce "family trees", and understand fossils. For example,

852-451: A constant rate. These " molecular clocks ", however, are fallible, and provide only a very approximate timing: for example, they are not sufficiently precise and reliable for estimating when the groups that feature in the Cambrian explosion first evolved, and estimates produced by different techniques may vary by a factor of two. Earth formed about 4,570  million years ago and, after

994-403: A data source that is not limited to animals with easily fossilised hard parts, and they reflect organisms' behaviours. Also many traces date from significantly earlier than the body fossils of animals that are thought to have been capable of making them. Whilst exact assignment of trace fossils to their makers is generally impossible, traces may for example provide the earliest physical evidence of

1136-577: A formal proposal to the ICS for the establishment of the Anthropocene Series/Epoch. Nevertheless, the definition of the Anthropocene as a geologic time period rather than a geologic event remains controversial and difficult. An international working group of the ICS on pre-Cryogenian chronostratigraphic subdivision have outlined a template to improve the pre-Cryogenian geologic time scale based on

1278-567: A fortunate accident during other research. For example, the 1980 discovery by Luis and Walter Alvarez of iridium , a mainly extraterrestrial metal, in the Cretaceous – Paleogene boundary layer made asteroid impact the most favored explanation for the Cretaceous–Paleogene extinction event – although debate continues about the contribution of volcanism. A complementary approach to developing scientific knowledge, experimental science ,

1420-461: A geochronologic unit can be changed (and is more often subject to change) when refined by geochronometry while the equivalent chronostratigraphic unit (the revision of which is less frequent) remains unchanged. For example, in early 2022, the boundary between the Ediacaran and Cambrian periods (geochronologic units) was revised from 541 Ma to 538.8 Ma but the rock definition of the boundary (GSSP) at

1562-741: A known geological context. The geological history of Mars has been divided into two alternate time scales. The first time scale for Mars was developed by studying the impact crater densities on the Martian surface. Through this method four periods have been defined, the Pre-Noachian (~4,500–4,100 Ma), Noachian (~4,100–3,700 Ma), Hesperian (~3,700–3,000 Ma), and Amazonian (~3,000 Ma to present). Paleontologist Paleontology ( / ˌ p eɪ l i ɒ n ˈ t ɒ l ə dʒ i , ˌ p æ l i -, - ən -/ PAY -lee-on- TOL -ə-jee, PAL -ee-, -⁠ən- ), also spelled palaeontology or palæontology ,

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1704-617: A machine-readable Resource Description Framework / Web Ontology Language representation of the time scale, which is available through the Commission for the Management and Application of Geoscience Information GeoSciML project as a service and at a SPARQL end-point. Some other planets and satellites in the Solar System have sufficiently rigid structures to have preserved records of their own histories, for example, Venus , Mars and

1846-814: A minor group until the first jawed fish appeared in the Late Ordovician . The spread of animals and plants from water to land required organisms to solve several problems, including protection against drying out and supporting themselves against gravity . The earliest evidence of land plants and land invertebrates date back to about 476  million years ago and 490  million years ago respectively. Those invertebrates, as indicated by their trace and body fossils, were shown to be arthropods known as euthycarcinoids . The lineage that produced land vertebrates evolved later but very rapidly between 370  million years ago and 360  million years ago ; recent discoveries have overturned earlier ideas about

1988-641: A name that was earlier, in 1952, proposed by Russian geologist and paleontologist Boris Sokolov . The Vendian concept was formed stratigraphically top-down, and the lower boundary of the Cambrian became the upper boundary of the Vendian. Paleontological substantiation of this boundary was worked out separately for the siliciclastic basin (base of the Baltic Stage of the Eastern European Platform ) and for

2130-555: A rapid increase in knowledge about the history of life on Earth and to progress in the definition of the geologic time scale , largely based on fossil evidence. Although she was rarely recognised by the scientific community, Mary Anning was a significant contributor to the field of palaeontology during this period; she uncovered multiple novel Mesozoic reptile fossils and deducted that what were then known as bezoar stones are in fact fossilised faeces . In 1822 Henri Marie Ducrotay de Blainville , editor of Journal de Physique , coined

2272-471: A relative interval of geologic time. A chronostratigraphic unit is a body of rock, layered or unlayered, that is defined between specified stratigraphic horizons which represent specified intervals of geologic time. They include all rocks representative of a specific interval of geologic time, and only this time span. Eonothem, erathem, system, series, subseries, stage, and substage are the hierarchical chronostratigraphic units. A geochronologic unit

2414-535: A relatively short time can be used to link up isolated rocks: this technique is called biostratigraphy . For instance, the conodont Eoplacognathus pseudoplanus has a short range in the Middle Ordovician period. If rocks of unknown age are found to have traces of E. pseudoplanus , they must have a mid-Ordovician age. Such index fossils must be distinctive, be globally distributed and have a short time range to be useful. However, misleading results are produced if

2556-429: A specific and reliable order. This allows for a correlation of strata even when the horizon between them is not continuous. The geologic time scale is divided into chronostratigraphic units and their corresponding geochronologic units. The subdivisions Early and Late are used as the geochronologic equivalents of the chronostratigraphic Lower and Upper , e.g., Early Triassic Period (geochronologic unit)

2698-604: A steady increase in brain size after about 3  million years ago . There is a long-running debate about whether modern humans are descendants of a single small population in Africa , which then migrated all over the world less than 200,000 years ago and replaced previous hominine species, or arose worldwide at the same time as a result of interbreeding . Life on earth has suffered occasional mass extinctions at least since 542  million years ago . Despite their disastrous effects, mass extinctions have sometimes accelerated

2840-481: A system/series (early/middle/late); however, the International Commission on Stratigraphy advocates for all new series and subseries to be named for a geographic feature in the vicinity of its stratotype or type locality . The name of stages should also be derived from a geographic feature in the locality of its stratotype or type locality. Informally, the time before the Cambrian is often referred to as

2982-458: A wider sense, correlating strata across national and continental boundaries based on their similarity to each other. Many of the names below erathem/era rank in use on the modern ICC/GTS were determined during the early to mid-19th century. During the 19th century, the debate regarding Earth's age was renewed, with geologists estimating ages based on denudation rates and sedimentary thicknesses or ocean chemistry, and physicists determining ages for

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3124-494: Is a numeric-only, chronologic reference point used to define the base of geochronologic units prior to the Cryogenian. These points are arbitrarily defined. They are used where GSSPs have not yet been established. Research is ongoing to define GSSPs for the base of all units that are currently defined by GSSAs. The standard international units of the geologic time scale are published by the International Commission on Stratigraphy on

3266-479: Is a subdivision of geologic time. It is a numeric representation of an intangible property (time). These units are arranged in a hierarchy: eon, era, period, epoch, subepoch, age, and subage. Geochronology is the scientific branch of geology that aims to determine the age of rocks, fossils, and sediments either through absolute (e.g., radiometric dating ) or relative means (e.g., stratigraphic position , paleomagnetism , stable isotope ratios ). Geochronometry

3408-413: Is a way of representing deep time based on events that have occurred throughout Earth's history , a time span of about 4.54 ± 0.05 Ga (4.54 billion years). It chronologically organises strata, and subsequently time, by observing fundamental changes in stratigraphy that correspond to major geological or paleontological events. For example, the Cretaceous–Paleogene extinction event , marks

3550-584: Is composed only of eukaryotic cells, and the earliest evidence for it is the Francevillian Group Fossils from 2,100  million years ago , although specialisation of cells for different functions first appears between 1,430  million years ago (a possible fungus) and 1,200  million years ago (a probable red alga ). Sexual reproduction may be a prerequisite for specialisation of cells, as an asexual multicellular organism might be at risk of being taken over by rogue cells that retain

3692-477: Is hard to decide at what level to place a new higher-level grouping, e.g. genus or family or order ; this is important since the Linnaean rules for naming groups are tied to their levels, and hence if a group is moved to a different level it must be renamed. Paleontologists generally use approaches based on cladistics , a technique for working out the evolutionary "family tree" of a set of organisms. It works by

3834-443: Is often said to work by conducting experiments to disprove hypotheses about the workings and causes of natural phenomena. This approach cannot prove a hypothesis, since some later experiment may disprove it, but the accumulation of failures to disprove is often compelling evidence in favor. However, when confronted with totally unexpected phenomena, such as the first evidence for invisible radiation , experimental scientists often use

3976-594: Is one that contained an extinct "crocodile-like" marine reptile, which eventually came to be known as the mosasaurid Mosasaurus of the Cretaceous period. The first half of the 19th century saw geological and paleontological activity become increasingly well organised with the growth of geologic societies and museums and an increasing number of professional geologists and fossil specialists. Interest increased for reasons that were not purely scientific, as geology and paleontology helped industrialists to find and exploit natural resources such as coal. This contributed to

4118-401: Is our only means of giving rocks greater than about 50 million years old an absolute age, and can be accurate to within 0.5% or better. Although radiometric dating requires very careful laboratory work, its basic principle is simple: the rates at which various radioactive elements decay are known, and so the ratio of the radioactive element to the element into which it decays shows how long ago

4260-568: Is prolonged and is estimated to last for ~9.0 Myrs. As to the Treptichnus pedum , a reference ichnofossil for the lower boundary of the Cambrian, its usage for the stratigraphic detection of this boundary is always risky, because of the occurrence of very similar trace fossils belonging to the Treptichnids group well below the level of T. pedum in Namibia , Spain and Newfoundland , and possibly, in

4402-401: Is the field of geochronology that numerically quantifies geologic time. A Global Boundary Stratotype Section and Point (GSSP) is an internationally agreed-upon reference point on a stratigraphic section that defines the lower boundaries of stages on the geologic time scale. (Recently this has been used to define the base of a system) A Global Standard Stratigraphic Age (GSSA)

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4544-663: Is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely define global chronostratigraphic units of the International Chronostratigraphic Chart (ICC) that are used to define divisions of geologic time. The chronostratigraphic divisions are in turn used to define geochronologic units. The geologic time scale

4686-451: Is the scientific study of life that existed prior to the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossils to classify organisms and study their interactions with each other and their environments (their paleoecology ). Paleontological observations have been documented as far back as the 5th century BC. The science became established in

4828-503: Is thought to have been propelled by coevolution with pollinating insects. Social insects appeared around the same time and, although they account for only small parts of the insect "family tree", now form over 50% of the total mass of all insects. Humans evolved from a lineage of upright-walking apes whose earliest fossils date from over 6  million years ago . Although early members of this lineage had chimp -sized brains, about 25% as big as modern humans', there are signs of

4970-573: Is used in place of Lower Triassic System (chronostratigraphic unit). Rocks representing a given chronostratigraphic unit are that chronostratigraphic unit, and the time they were laid down in is the geochronologic unit, e.g., the rocks that represent the Silurian System are the Silurian System and they were deposited during the Silurian Period. This definition means the numeric age of

5112-646: The Anthropocene is a proposed epoch/series for the most recent time in Earth's history. While still informal, it is a widely used term to denote the present geologic time interval, in which many conditions and processes on Earth are profoundly altered by human impact. As of April 2022 the Anthropocene has not been ratified by the ICS; however, in May 2019 the Anthropocene Working Group voted in favour of submitting

5254-529: The Brothers of Purity , who wrote on the processes of stratification over the passage of time in their treatises . Their work likely inspired that of the 11th-century Persian polymath Avicenna (Ibn Sînâ, 980–1037) who wrote in The Book of Healing (1027) on the concept of stratification and superposition, pre-dating Nicolas Steno by more than six centuries. Avicenna also recognised fossils as "petrifications of

5396-567: The Middle Ages the Persian naturalist Ibn Sina , known as Avicenna in Europe, discussed fossils and proposed a theory of petrifying fluids on which Albert of Saxony elaborated in the 14th century. The Chinese naturalist Shen Kuo (1031–1095) proposed a theory of climate change based on the presence of petrified bamboo in regions that in his time were too dry for bamboo. In early modern Europe ,

5538-528: The Neogene - Quaternary . In deeper-level deposits in western Europe are early-aged mammals such as the palaeothere perissodactyl Palaeotherium and the anoplotheriid artiodactyl Anoplotherium , both of which were described earliest after the former two genera, which today are known to date to the Paleogene period. Cuvier figured out that even older than the two levels of deposits with extinct large mammals

5680-637: The Permian–Triassic extinction event . Amphibians Extinct Synapsids Mammals Extinct reptiles Lizards and snakes Extinct Archosaurs Crocodilians Extinct Dinosaurs Birds Naming groups of organisms in a way that is clear and widely agreed is important, as some disputes in paleontology have been based just on misunderstandings over names. Linnaean taxonomy is commonly used for classifying living organisms, but runs into difficulties when dealing with newly discovered organisms that are significantly different from known ones. For example: it

5822-513: The Permian–Triassic extinction event . A relatively recent discipline, molecular phylogenetics , compares the DNA and RNA of modern organisms to re-construct the "family trees" of their evolutionary ancestors. It has also been used to estimate the dates of important evolutionary developments, although this approach is controversial because of doubts about the reliability of the " molecular clock ". Techniques from engineering have been used to analyse how

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5964-530: The Precambrian or pre-Cambrian (Supereon). While a modern geological time scale was not formulated until 1911 by Arthur Holmes , the broader concept that rocks and time are related can be traced back to (at least) the philosophers of Ancient Greece . Xenophanes of Colophon (c. 570–487  BCE ) observed rock beds with fossils of shells located above the sea-level, viewed them as once living organisms, and used this to imply an unstable relationship in which

6106-653: The carbonate basin (base of the Tommotian stage of the Siberian Platform ). The lower boundary of the Vendian was suggested to be defined at the base of the Varanger ( Laplandian ) tillites . The Vendian in its type area consists of large subdivisions such as Laplandian, Redkino , Kotlin and Rovno regional stages with the globally traceable subdivisions and their boundaries, including its lower one. The Redkino, Kotlin and Rovno regional stages have been substantiated in

6248-454: The embryological development of some modern brachiopods suggests that brachiopods may be descendants of the halkieriids , which became extinct in the Cambrian period. Paleontology seeks to map out how living things have changed through time. A substantial hurdle to this aim is the difficulty of working out how old fossils are. Beds that preserve fossils typically lack the radioactive elements needed for radiometric dating . This technique

6390-577: The western United States . The stratigraphic range of T. pedum overlaps the range of the Ediacaran fossils in Namibia, and probably in Spain. The Ediacaran Period is not yet formally subdivided, but a proposed scheme recognises an Upper Ediacaran whose base corresponds with the Gaskiers glaciation , a Terminal Ediacaran Stage starting around 550  million years ago , a preceding stage beginning around 575 Ma with

6532-521: The " jigsaw puzzles " of biostratigraphy (arrangement of rock layers from youngest to oldest). Classifying ancient organisms is also difficult, as many do not fit well into the Linnaean taxonomy classifying living organisms, and paleontologists more often use cladistics to draw up evolutionary "family trees". The final quarter of the 20th century saw the development of molecular phylogenetics , which investigates how closely organisms are related by measuring

6674-463: The 18th century as a result of Georges Cuvier 's work on comparative anatomy , and developed rapidly in the 19th century. The term has been used since 1822 formed from Greek παλαιός ( 'palaios' , "old, ancient"), ὄν ( 'on' , ( gen. 'ontos' ), "being, creature"), and λόγος ( 'logos' , "speech, thought, study"). Paleontology lies on the border between biology and geology , but it differs from archaeology in that it excludes

6816-504: The Avalon explosion 575 million years ago and died out during the End-Ediacaran extinction event 539 million years ago. Forerunners of some modern animal phyla also appeared during this period, including cnidarians and early bilaterians , as well as mollusc -like Kimberella . Hard-bodied organisms with mineralized shells also began their fossil record in the last few million years of

6958-577: The C-isotope data are concerned, due to primary lateral variations of δ C carb in the upper layer of the ocean. Furthermore, Oman presents in its stratigraphic record a large negative carbon isotope excursion, within the Shuram Formation that is clearly away from any glacial evidence strongly questioning systematic association of negative δ C carb excursion and glacial events. Also, the Shuram excursion

7100-668: The Commission on Stratigraphy (applied in 1965) to become a member commission of IUGS led to the founding of the ICS. One of the primary objectives of the ICS is "the establishment, publication and revision of the ICS International Chronostratigraphic Chart which is the standard, reference global Geological Time Scale to include the ratified Commission decisions". Following on from Holmes, several A Geological Time Scale books were published in 1982, 1989, 2004, 2008, 2012, 2016, and 2020. However, since 2013,

7242-548: The Early Cambrian , along with several "weird wonders" that bear little obvious resemblance to any modern animals. There is a long-running debate about whether this Cambrian explosion was truly a very rapid period of evolutionary experimentation; alternative views are that modern-looking animals began evolving earlier but fossils of their precursors have not yet been found, or that the "weird wonders" are evolutionary "aunts" and "cousins" of modern groups. Vertebrates remained

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7384-464: The Earth's Moon . Dominantly fluid planets, such as the giant planets , do not comparably preserve their history. Apart from the Late Heavy Bombardment , events on other planets probably had little direct influence on the Earth, and events on Earth had correspondingly little effect on those planets. Construction of a time scale that links the planets is, therefore, of only limited relevance to

7526-411: The Earth's organic and inorganic past". William Whewell (1794–1866) classified paleontology as one of the historical sciences, along with archaeology , geology, astronomy , cosmology , philology and history itself: paleontology aims to describe phenomena of the past and to reconstruct their causes. Hence it has three main elements: description of past phenomena; developing a general theory about

7668-468: The Earth's time scale, except in a Solar System context. The existence, timing, and terrestrial effects of the Late Heavy Bombardment are still a matter of debate. The geologic history of Earth's Moon has been divided into a time scale based on geomorphological markers, namely impact cratering , volcanism , and erosion . This process of dividing the Moon's history in this manner means that

7810-897: The Ediacara Hills fossil site. The Ediacaran marks the first widespread appearance of complex multicellular fauna following the end of the Cryogenian global glaciation known as the Snowball Earth . The relatively sudden evolutionary radiation event, known as the Avalon Explosion , is represented by now-extinct, relatively simple soft-bodied animal phyla such as Proarticulata ( bilaterians with simple articulation , e.g. Dickinsonia and Spriggina ), Petalonamae ( sea pen -like animals, e.g. Charnia ), Aspidella (radial-shaped animals, e.g. Cyclomedusa ) and Trilobozoa (animals with tri-radial symmetry , e.g. Tribrachidium ). Most of these organisms appeared during or after

7952-461: The Ediacaran. The supercontinent Pannotia formed and broke apart by the end of the period. The Ediacaran also witnessed several glaciation events , such as the Gaskiers and Baykonurian glaciations . The Shuram excursion also occurred during this period, but its glacial origin is unlikely. The Ediacaran Period overlaps but is shorter than the Vendian Period (650 to 543 million years ago),

8094-712: The Elatina diamictite in the Enorama Creek section, Brachina Gorge, Flinders Ranges, South Australia. The GSSP of the upper boundary of the Ediacaran is the lower boundary of the Cambrian on the SE coast of Newfoundland approved by the International Commission on Stratigraphy as a preferred alternative to the base of the Tommotian Stage in Siberia which was selected on the basis of the ichnofossil Treptichnus pedum (Seilacher, 1955). In

8236-447: The ICS has taken responsibility for producing and distributing the ICC citing the commercial nature, independent creation, and lack of oversight by the ICS on the prior published GTS versions (GTS books prior to 2013) although these versions were published in close association with the ICS. Subsequent Geologic Time Scale books (2016 and 2020 ) are commercial publications with no oversight from

8378-404: The ICS, and do not entirely conform to the chart produced by the ICS. The ICS produced GTS charts are versioned (year/month) beginning at v2013/01. At least one new version is published each year incorporating any changes ratified by the ICS since the prior version. The following five timelines show the geologic time scale to scale. The first shows the entire time from the formation of the Earth to

8520-415: The ICS. While some regional terms are still in use, the table of geologic time conforms to the nomenclature , ages, and colour codes set forth by the International Commission on Stratigraphy in the official International Chronostratigraphic Chart. The International Commission on Stratigraphy also provide an online interactive version of this chart. The interactive version is based on a service delivering

8662-665: The International Chronostratigraphic Chart; however, regional terms are still in use in some areas. The numeric values on the International Chronostratigrahpic Chart are represented by the unit Ma (megaannum, for 'million years '). For example, 201.4 ± 0.2 Ma, the lower boundary of the Jurassic Period, is defined as 201,400,000 years old with an uncertainty of 200,000 years. Other SI prefix units commonly used by geologists are Ga (gigaannum, billion years), and ka (kiloannum, thousand years), with

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8804-607: The Nama biotic assemblage. There is evidence for a mass extinction during this period from early animals changing the environment, dating to the same time as the transition between the White Sea and the Nama-type biotas. Alternatively, this mass extinction has also been theorised to have been the result of an anoxic event . The relative proximity of the Moon at this time meant that tides were stronger and more rapid than they are now. The day

8946-412: The ability to reproduce. The earliest known animals are cnidarians from about 580  million years ago , but these are so modern-looking that they must be descendants of earlier animals. Early fossils of animals are rare because they had not developed mineralised , easily fossilized hard parts until about 548  million years ago . The earliest modern-looking bilaterian animals appear in

9088-445: The action of gravity. However, it is now known that not all sedimentary layers are deposited purely horizontally, but this principle is still a useful concept. The principle of lateral continuity that states layers of sediments extend laterally in all directions until either thinning out or being cut off by a different rock layer, i.e. they are laterally continuous. Layers do not extend indefinitely; their limits are controlled by

9230-505: The age of rocks). It is used primarily by Earth scientists (including geologists , paleontologists , geophysicists , geochemists , and paleoclimatologists ) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies , paleomagnetic properties, and fossils . The definition of standardised international units of geologic time

9372-506: The age range of 635 to 538.8 million years is based on correlations to other countries where dating has been possible. The base age of approximately 635 million years is based on U–Pb ( uranium – lead ) and Re–Os ( rhenium – osmium ) dating from Africa, China, North America, and Tasmania. The fossil record from the Ediacaran Period is sparse, as more easily fossilized hard-shelled animals had yet to evolve. The Ediacaran biota include

9514-409: The amount and type of sediment in a sedimentary basin , and the geometry of that basin. The principle of cross-cutting relationships that states a rock that cuts across another rock must be younger than the rock it cuts across. The law of included fragments that states small fragments of one type of rock that are embedded in a second type of rock must have formed first, and were included when

9656-515: The appearance of moderately complex animals (comparable to earthworms ). Geochemical observations may help to deduce the global level of biological activity at a certain period, or the affinity of certain fossils. For example, geochemical features of rocks may reveal when life first arose on Earth, and may provide evidence of the presence of eukaryotic cells, the type from which all multicellular organisms are built. Analyses of carbon isotope ratios may help to explain major transitions such as

9798-625: The atmosphere increased their effectiveness as nurseries of evolution. While eukaryotes , cells with complex internal structures, may have been present earlier, their evolution speeded up when they acquired the ability to transform oxygen from a poison to a powerful source of metabolic energy. This innovation may have come from primitive eukaryotes capturing oxygen-powered bacteria as endosymbionts and transforming them into organelles called mitochondria . The earliest evidence of complex eukaryotes with organelles (such as mitochondria) dates from 1,850  million years ago . Multicellular life

9940-456: The base of the Cambrian, and thus the boundary between the Ediacaran and Cambrian systems (chronostratigraphic units) has not been changed; rather, the absolute age has merely been refined. Chronostratigraphy is the element of stratigraphy that deals with the relation between rock bodies and the relative measurement of geological time. It is the process where distinct strata between defined stratigraphic horizons are assigned to represent

10082-565: The beginning of the subsequent Cambrian Period marks the start of the Phanerozoic Eon, where recognizable fossil evidence of life becomes common. The Ediacaran Period is named after the Ediacara Hills of South Australia , where trace fossils of a diverse community of previously unrecognized lifeforms (later named the Ediacaran biota ) were first discovered by geologist Reg Sprigg in 1946. Its status as an official geological period

10224-415: The bodies of ancient organisms might have worked, for example the running speed and bite strength of Tyrannosaurus , or the flight mechanics of Microraptor . It is relatively commonplace to study the internal details of fossils using X-ray microtomography . Paleontology, biology, archaeology, and paleoneurobiology combine to study endocranial casts (endocasts) of species related to humans to clarify

10366-518: The bodies of plants and animals", with the 13th-century Dominican bishop Albertus Magnus (c. 1200–1280) extending this into a theory of a petrifying fluid. These works appeared to have little influence on scholars in Medieval Europe who looked to the Bible to explain the origins of fossils and sea-level changes, often attributing these to the ' Deluge ', including Ristoro d'Arezzo in 1282. It

10508-430: The cap carbonates in a rather short distance but cap carbonates do not occur above every tillite elsewhere in the world. The C-isotope chemostratigraphic characteristics obtained for contemporaneous cap carbonates in different parts of the world may be variable in a wide range owing to different degrees of secondary alteration of carbonates, dissimilar criteria used for selection of the least altered samples, and, as far as

10650-401: The causes of various types of change; and applying those theories to specific facts. When trying to explain the past, paleontologists and other historical scientists often construct a set of one or more hypotheses about the causes and then look for a " smoking gun ", a piece of evidence that strongly accords with one hypothesis over any others. Sometimes researchers discover a "smoking gun" by

10792-763: The characteristics and evolution of humans as a species. When dealing with evidence about humans, archaeologists and paleontologists may work together – for example paleontologists might identify animal or plant fossils around an archaeological site , to discover the people who lived there, and what they ate; or they might analyze the climate at the time of habitation. In addition, paleontology often borrows techniques from other sciences, including biology, osteology , ecology, chemistry , physics and mathematics. For example, geochemical signatures from rocks may help to discover when life first arose on Earth, and analyses of carbon isotope ratios may help to identify climate changes and even to explain major transitions such as

10934-520: The chronological order in which rocks were formed, is useful to both paleontologists and geologists. Biogeography studies the spatial distribution of organisms, and is also linked to geology, which explains how Earth's geography has changed over time. Although paleontology became established around 1800, earlier thinkers had noticed aspects of the fossil record. The ancient Greek philosopher Xenophanes (570–480 BCE) concluded from fossil sea shells that some areas of land were once under water. During

11076-566: The cooling of the Earth or the Sun using basic thermodynamics or orbital physics. These estimations varied from 15,000 million years to 0.075 million years depending on method and author, but the estimations of Lord Kelvin and Clarence King were held in high regard at the time due to their pre-eminence in physics and geology. All of these early geochronometric determinations would later prove to be incorrect. The discovery of radioactive decay by Henri Becquerel , Marie Curie , and Pierre Curie laid

11218-445: The date when lineages first appeared. For instance, if fossils of B or C date to X million years ago and the calculated "family tree" says A was an ancestor of B and C, then A must have evolved more than X million years ago. It is also possible to estimate how long ago two living clades diverged – i.e. approximately how long ago their last common ancestor must have lived – by assuming that DNA mutations accumulate at

11360-586: The development of mammalian traits such as endothermy and hair. After the Cretaceous–Paleogene extinction event 66  million years ago killed off all the dinosaurs except the birds, mammals increased rapidly in size and diversity, and some took to the air and the sea. Fossil evidence indicates that flowering plants appeared and rapidly diversified in the Early Cretaceous between 130  million years ago and 90  million years ago . Their rapid rise to dominance of terrestrial ecosystems

11502-449: The development of the body plans of most animal phyla . The discovery of fossils of the Ediacaran biota and developments in paleobiology extended knowledge about the history of life back far before the Cambrian. Increasing awareness of Gregor Mendel 's pioneering work in genetics led first to the development of population genetics and then in the mid-20th century to the modern evolutionary synthesis , which explains evolution as

11644-453: The developments in mass spectrometry pioneered by Francis William Aston , Arthur Jeffrey Dempster , and Alfred O. C. Nier during the early to mid- 20th century would finally allow for the accurate determination of radiometric ages, with Holmes publishing several revisions to his geological time-scale with his final version in 1960. The establishment of the IUGS in 1961 and acceptance of

11786-404: The different layers of stone unless they had been upon the shore and had been covered over by earth newly thrown up by the sea which then became petrified? And if the above-mentioned Deluge had carried them to these places from the sea, you would find the shells at the edge of one layer of rock only, not at the edge of many where may be counted the winters of the years during which the sea multiplied

11928-477: The different levels of deposits represented different time periods in the early 19th century. The surface-level deposits in the Americas contained later mammals like the megatheriid ground sloth Megatherium and the mammutid proboscidean Mammut (later known informally as a "mastodon"), which were some of the earliest-named fossil mammal genera with official taxonomic authorities. They today are known to date to

12070-707: The earliest widespread Ediacaran biota fossils; two proposed schemes differ on whether the lower strata should be divided into an Early and Middle Ediacaran or not, because it is not clear whether the Shuram excursion (which would divide the Early and Middle) is a separate event from the Gaskiers, or whether the two events are correlated. The dating of the rock type section of the Ediacaran Period in South Australia has proven uncertain due to lack of overlying igneous material. Therefore,

12212-409: The end of the 20th century have been particularly important as they have provided new information about the earliest evolution of animals, early fish, dinosaurs and the evolution of birds. The last few decades of the 20th century saw a renewed interest in mass extinctions and their role in the evolution of life on Earth. There was also a renewed interest in the Cambrian explosion that apparently saw

12354-410: The evolution of the human brain. Paleontology even contributes to astrobiology , the investigation of possible life on other planets , by developing models of how life may have arisen and by providing techniques for detecting evidence of life. As knowledge has increased, paleontology has developed specialised subdivisions. Vertebrate paleontology concentrates on fossils from the earliest fish to

12496-466: The evolutionary history of life back to over 3,000  million years ago , possibly as far as 3,800  million years ago . The oldest clear evidence of life on Earth dates to 3,000  million years ago , although there have been reports, often disputed, of fossil bacteria from 3,400  million years ago and of geochemical evidence for the presence of life 3,800  million years ago . Some scientists have proposed that life on Earth

12638-555: The exceptional events that cause quick burial make it difficult to study the normal environments of the animals. The sparseness of the fossil record means that organisms are expected to exist long before and after they are found in the fossil record – this is known as the Signor–Lipps effect . Trace fossils consist mainly of tracks and burrows, but also include coprolites (fossil feces ) and marks left by feeding. Trace fossils are particularly significant because they represent

12780-504: The focus of paleontology shifted to understanding evolutionary paths, including human evolution , and evolutionary theory. The last half of the 19th century saw a tremendous expansion in paleontological activity, especially in North America. The trend continued in the 20th century with additional regions of the Earth being opened to systematic fossil collection. Fossils found in China near

12922-449: The following: At the end of the 18th century Georges Cuvier 's work established comparative anatomy as a scientific discipline and, by proving that some fossil animals resembled no living ones, demonstrated that animals could become extinct , leading to the emergence of paleontology. The expanding knowledge of the fossil record also played an increasing role in the development of geology, particularly stratigraphy . Cuvier proved that

13064-580: The fossil record: different environments are more favorable to the preservation of different types of organism or parts of organisms. Further, only the parts of organisms that were already mineralised are usually preserved, such as the shells of molluscs. Since most animal species are soft-bodied, they decay before they can become fossilised. As a result, although there are 30-plus phyla of living animals, two-thirds have never been found as fossils. Occasionally, unusual environments may preserve soft tissues. These lagerstätten allow paleontologists to examine

13206-414: The foundational principles of determining the correlation of strata relative to geologic time. Over the course of the 18th-century geologists realised that: The apparent, earliest formal division of the geologic record with respect to time was introduced during the era of Biblical models by Thomas Burnet who applied a two-fold terminology to mountains by identifying " montes primarii " for rock formed at

13348-401: The geologic time scale of Earth. This table is arranged with the most recent geologic periods at the top, and the oldest at the bottom. The height of each table entry does not correspond to the duration of each subdivision of time. As such, this table is not to scale and does not accurately represent the relative time-spans of each geochronologic unit. While the Phanerozoic Eon looks longer than

13490-492: The ground work for radiometric dating, but the knowledge and tools required for accurate determination of radiometric ages would not be in place until the mid-1950s. Early attempts at determining ages of uranium minerals and rocks by Ernest Rutherford , Bertram Boltwood , Robert Strutt , and Arthur Holmes, would culminate in what are considered the first international geological time scales by Holmes in 1911 and 1913. The discovery of isotopes in 1913 by Frederick Soddy , and

13632-675: The history and driving forces behind their evolution. Land plants were so successful that their detritus caused an ecological crisis in the Late Devonian , until the evolution of fungi that could digest dead wood. During the Permian period, synapsids , including the ancestors of mammals , may have dominated land environments, but this ended with the Permian–Triassic extinction event 251  million years ago , which came very close to wiping out all complex life. The extinctions were apparently fairly sudden, at least among vertebrates. During

13774-521: The history of Earth's climate and the mechanisms that have changed it  – which have sometimes included evolutionary developments, for example the rapid expansion of land plants in the Devonian period removed more carbon dioxide from the atmosphere, reducing the greenhouse effect and thus helping to cause an ice age in the Carboniferous period. Biostratigraphy , the use of fossils to work out

13916-487: The history of stratigraphy it was the first case of usage of bioturbations for the System boundary definition. Nevertheless, the definitions of the lower and upper boundaries of the Ediacaran on the basis of chemostratigraphy and ichnofossils are disputable. Cap carbonates generally have a restricted geographic distribution (due to specific conditions of their precipitation) and usually siliciclastic sediments laterally replace

14058-542: The immediate ancestors of modern mammals . Invertebrate paleontology deals with fossils such as molluscs , arthropods , annelid worms and echinoderms . Paleobotany studies fossil plants , algae , and fungi. Palynology , the study of pollen and spores produced by land plants and protists , straddles paleontology and botany , as it deals with both living and fossil organisms. Micropaleontology deals with microscopic fossil organisms of all kinds. Instead of focusing on individual organisms, paleoecology examines

14200-434: The index fossils turn out to have longer fossil ranges than first thought. Stratigraphy and biostratigraphy can in general provide only relative dating ( A was before B ), which is often sufficient for studying evolution. However, this is difficult for some time periods, because of the problems involved in matching up rocks of the same age across different continents . Family-tree relationships may also help to narrow down

14342-538: The interactions between different ancient organisms, such as their food chains , and the two-way interactions with their environments.   For example, the development of oxygenic photosynthesis by bacteria caused the oxygenation of the atmosphere and hugely increased the productivity and diversity of ecosystems . Together, these led to the evolution of complex eukaryotic cells, from which all multicellular organisms are built. Paleoclimatology , although sometimes treated as part of paleoecology, focuses more on

14484-463: The internal anatomy of animals that in other sediments are represented only by shells, spines, claws, etc. – if they are preserved at all. However, even lagerstätten present an incomplete picture of life at the time. The majority of organisms living at the time are probably not represented because lagerstätten are restricted to a narrow range of environments, e.g. where soft-bodied organisms can be preserved very quickly by events such as mudslides; and

14626-456: The investigation of evolutionary "family trees" by techniques derived from biochemistry , began to make an impact, particularly when it was proposed that the human lineage had diverged from apes much more recently than was generally thought at the time. Although this early study compared proteins from apes and humans, most molecular phylogenetics research is now based on comparisons of RNA and DNA . Fossils of organisms' bodies are usually

14768-913: The latter often represented in calibrated units ( before present ). The names of geologic time units are defined for chronostratigraphic units with the corresponding geochronologic unit sharing the same name with a change to the suffix (e.g. Phanerozoic Eonothem becomes the Phanerozoic Eon). Names of erathems in the Phanerozoic were chosen to reflect major changes in the history of life on Earth: Paleozoic (old life), Mesozoic (middle life), and Cenozoic (new life). Names of systems are diverse in origin, with some indicating chronologic position (e.g., Paleogene), while others are named for lithology (e.g., Cretaceous), geography (e.g., Permian ), or are tribal (e.g., Ordovician ) in origin. Most currently recognised series and subseries are named for their position within

14910-561: The layers of sand and mud brought down by the neighboring rivers and spread them over its shores. And if you wish to say that there must have been many deluges in order to produce these layers and the shells among them it would then become necessary for you to affirm that such a deluge took place every year. These views of da Vinci remained unpublished, and thus lacked influence at the time; however, questions of fossils and their significance were pursued and, while views against Genesis were not readily accepted and dissent from religious doctrine

15052-537: The litho- and biostratigraphic differences around the world in time equivalent rocks. The ICS has long worked to reconcile conflicting terminology by standardising globally significant and identifiable stratigraphic horizons that can be used to define the lower boundaries of chronostratigraphic units. Defining chronostratigraphic units in such a manner allows for the use of global, standardised nomenclature. The International Chronostratigraphic Chart represents this ongoing effort. Several key principles are used to determine

15194-409: The logic that, if groups B and C have more similarities to each other than either has to group A, then B and C are more closely related to each other than either is to A. Characters that are compared may be anatomical , such as the presence of a notochord , or molecular , by comparing sequences of DNA or proteins . The result of a successful analysis is a hierarchy of clades – groups that share

15336-555: The lower boundary of the Paleogene System/Period and thus the boundary between the Cretaceous and Paleogene systems/periods. For divisions prior to the Cryogenian , arbitrary numeric boundary definitions ( Global Standard Stratigraphic Ages , GSSAs) are used to divide geologic time. Proposals have been made to better reconcile these divisions with the rock record. Historically, regional geologic time scales were used due to

15478-409: The most informative type of evidence. The most common types are wood, bones, and shells. Fossilisation is a rare event, and most fossils are destroyed by erosion or metamorphism before they can be observed. Hence the fossil record is very incomplete, increasingly so further back in time. Despite this, it is often adequate to illustrate the broader patterns of life's history. There are also biases in

15620-628: The oldest definite multicellular organisms (with specialized tissues), the most common types of which resemble segmented worms, fronds, disks, or immobile bags. Auroralumina was a cnidarian . Most members of the Ediacaran biota bear little resemblance to modern lifeforms, and their relationship even with the immediately following lifeforms of the Cambrian explosion is rather difficult to interpret. More than 100 genera have been described, and well known forms include Arkarua , Charnia , Dickinsonia , Ediacaria , Marywadea , Cephalonega , Pteridinium , and Yorgia . However, despite

15762-414: The outcome of events such as mutations and horizontal gene transfer , which provide genetic variation , with genetic drift and natural selection driving changes in this variation over time. Within the next few years the role and operation of DNA in genetic inheritance were discovered, leading to what is now known as the "Central Dogma" of molecular biology . In the 1960s molecular phylogenetics ,

15904-484: The overall enigmaticness of most Ediacaran organisms, some fossils identifiable as hard-shelled agglutinated foraminifera (which are not classified as animals) are known from latest Ediacaran sediments of western Siberia. Sponges recognisable as such also lived during the Ediacaran. Four different biotic intervals are known in the Ediacaran, each being characterised by the prominence of a unique ecology and faunal assemblage. The first spanned from 635 to around 575 Ma and

16046-408: The pertinent time span. As of April 2022 these proposed changes have not been accepted by the ICS. The proposed changes (changes from the current scale [v2023/09]) are italicised: Proposed pre-Cambrian timeline (GTS2012), shown to scale: Current ICC pre-Cambrian timeline (v2023/09), shown to scale: The following table summarises the major events and characteristics of the divisions making up

16188-452: The present, but this gives little space for the most recent eon. The second timeline shows an expanded view of the most recent eon. In a similar way, the most recent era is expanded in the third timeline, the most recent period is expanded in the fourth timeline, and the most recent epoch is expanded in the fifth timeline. Horizontal scale is Millions of years (above timelines) / Thousands of years (below timeline) First suggested in 2000,

16330-452: The principal types of evidence about ancient life, and geochemical evidence has helped to decipher the evolution of life before there were organisms large enough to leave body fossils. Estimating the dates of these remains is essential but difficult: sometimes adjacent rock layers allow radiometric dating , which provides absolute dates that are accurate to within 0.5%, but more often paleontologists have to rely on relative dating by solving

16472-485: The principles of superposition, original horizontality, lateral continuity, and cross-cutting relationships. From this Steno reasoned that strata were laid down in succession and inferred relative time (in Steno's belief, time from Creation ). While Steno's principles were simple and attracted much attention, applying them proved challenging. These basic principles, albeit with improved and more nuanced interpretations, still form

16614-432: The radioactive element was incorporated into the rock. Radioactive elements are common only in rocks with a volcanic origin, and so the only fossil-bearing rocks that can be dated radiometrically are a few volcanic ash layers. Consequently, paleontologists must usually rely on stratigraphy to date fossils. Stratigraphy is the science of deciphering the "layer-cake" that is the sedimentary record, and has been compared to

16756-521: The relative relationships of rocks and thus their chronostratigraphic position. The law of superposition that states that in undeformed stratigraphic sequences the oldest strata will lie at the bottom of the sequence, while newer material stacks upon the surface. In practice, this means a younger rock will lie on top of an older rock unless there is evidence to suggest otherwise. The principle of original horizontality that states layers of sediments will originally be deposited horizontally under

16898-467: The rest, it merely spans ~539 million years (~12% of Earth's history), whilst the previous three eons collectively span ~3,461 million years (~76% of Earth's history). This bias toward the most recent eon is in part due to the relative lack of information about events that occurred during the first three eons compared to the current eon (the Phanerozoic). The use of subseries/subepochs has been ratified by

17040-630: The rock record to bring it in line with the post-Tonian geologic time scale. This work assessed the geologic history of the currently defined eons and eras of the pre-Cambrian, and the proposals in the "Geological Time Scale" books 2004, 2012, and 2020. Their recommend revisions of the pre-Cryogenian geologic time scale were (changes from the current scale [v2023/09] are italicised): Proposed pre-Cambrian timeline (Shield et al. 2021, ICS working group on pre-Cryogenian chronostratigraphy), shown to scale: Current ICC pre-Cambrian timeline (v2023/09), shown to scale: The book, Geologic Time Scale 2012,

17182-701: The same approach as historical scientists: construct a set of hypotheses about the causes and then look for a "smoking gun". Paleontology lies between biology and geology since it focuses on the record of past life, but its main source of evidence is fossils in rocks. For historical reasons, paleontology is part of the geology department at many universities: in the 19th and early 20th centuries, geology departments found fossil evidence important for dating rocks, while biology departments showed little interest. Paleontology also has some overlap with archaeology , which primarily works with objects made by humans and with human remains, while paleontologists are interested in

17324-474: The sea had at times transgressed over the land and at other times had regressed . This view was shared by a few of Xenophanes's contemporaries and those that followed, including Aristotle (384–322 BCE) who (with additional observations) reasoned that the positions of land and sea had changed over long periods of time. The concept of deep time was also recognised by Chinese naturalist Shen Kuo (1031–1095) and Islamic scientist -philosophers, notably

17466-544: The second rock was forming. The relationships of unconformities which are geologic features representing a gap in the geologic record. Unconformities are formed during periods of erosion or non-deposition, indicating non-continuous sediment deposition. Observing the type and relationships of unconformities in strata allows geologist to understand the relative timing the strata. The principle of faunal succession (where applicable) that states rock strata contain distinctive sets of fossils that succeed each other vertically in

17608-478: The similarity of the DNA in their genomes . Molecular phylogenetics has also been used to estimate the dates when species diverged, but there is controversy about the reliability of the molecular clock on which such estimates depend. The simplest definition of "paleontology" is "the study of ancient life". The field seeks information about several aspects of past organisms: "their identity and origin, their environment and evolution, and what they can tell us about

17750-465: The slow recovery from this catastrophe a previously obscure group, archosaurs , became the most abundant and diverse terrestrial vertebrates. One archosaur group, the dinosaurs, were the dominant land vertebrates for the rest of the Mesozoic , and birds evolved from one group of dinosaurs. During this time mammals' ancestors survived only as small, mainly nocturnal insectivores , which may have accelerated

17892-631: The study of anatomically modern humans . It now uses techniques drawn from a wide range of sciences, including biochemistry , mathematics , and engineering. Use of all these techniques has enabled paleontologists to discover much of the evolutionary history of life , almost back to when Earth became capable of supporting life, nearly 4 billion years ago. As knowledge has increased, paleontology has developed specialised sub-divisions, some of which focus on different types of fossil organisms while others study ecology and environmental history, such as ancient climates . Body fossils and trace fossils are

18034-617: The systematic study of fossils emerged as an integral part of the changes in natural philosophy that occurred during the Age of Reason . In the Italian Renaissance, Leonardo da Vinci made various significant contributions to the field as well as depicted numerous fossils. Leonardo's contributions are central to the history of paleontology because he established a line of continuity between the two main branches of paleontology – ichnology and body fossil paleontology. He identified

18176-537: The time during which the rocks were laid down, and the collection of rocks themselves (i.e., it was correct to say Tertiary rocks, and Tertiary Period). Only the Quaternary division is retained in the modern geologic time scale, while the Tertiary division was in use until the early 21st century. The Neptunism and Plutonism theories would compete into the early 19th century with a key driver for resolution of this debate being

18318-400: The time from the end of global Marinoan glaciation to the first appearance worldwide of somewhat complicated trace fossils ( Treptichnus pedum (Seilacher, 1955)). Although the Ediacaran Period does contain soft-bodied fossils , it is unusual in comparison to later periods because its beginning is not defined by a change in the fossil record. Rather, the beginning is defined at the base of

18460-730: The time of the 'Deluge', and younger " monticulos secundarios" formed later from the debris of the " primarii" . Anton Moro (1687–1784) also used primary and secondary divisions for rock units but his mechanism was volcanic. In this early version of the Plutonism theory, the interior of Earth was seen as hot, and this drove the creation of primary igneous and metamorphic rocks and secondary rocks formed contorted and fossiliferous sediments. These primary and secondary divisions were expanded on by Giovanni Targioni Tozzetti (1712–1783) and Giovanni Arduino (1713–1795) to include tertiary and quaternary divisions. These divisions were used to describe both

18602-562: The time scale boundaries do not imply fundamental changes in geological processes, unlike Earth's geologic time scale. Five geologic systems/periods ( Pre-Nectarian , Nectarian , Imbrian , Eratosthenian , Copernican ), with the Imbrian divided into two series/epochs (Early and Late) were defined in the latest Lunar geologic time scale. The Moon is unique in the Solar System in that it is the only other body from which humans have rock samples with

18744-465: The type area of the Vendian on the basis of the abundant organic-walled microfossils , megascopic algae, metazoan body fossils and ichnofossils . The lower boundary of the Vendian could have a biostratigraphic substantiation as well taking into consideration the worldwide occurrence of the Pertatataka assemblage of giant acanthomorph acritarchs . The Ediacaran Period (c. 635–538.8 Mya) represents

18886-406: The word "palaeontology" to refer to the study of ancient living organisms through fossils. As knowledge of life's history continued to improve, it became increasingly obvious that there had been some kind of successive order to the development of life. This encouraged early evolutionary theories on the transmutation of species . After Charles Darwin published Origin of Species in 1859, much of

19028-526: The work of James Hutton (1726–1797), in particular his Theory of the Earth , first presented before the Royal Society of Edinburgh in 1785. Hutton's theory would later become known as uniformitarianism , popularised by John Playfair (1748–1819) and later Charles Lyell (1797–1875) in his Principles of Geology . Their theories strongly contested the 6,000 year age of the Earth as suggested determined by James Ussher via Biblical chronology that

19170-454: Was "seeded" from elsewhere , but most research concentrates on various explanations of how life could have arisen independently on Earth. For about 2,000 million years microbial mats , multi-layered colonies of different bacteria, were the dominant life on Earth. The evolution of oxygenic photosynthesis enabled them to play the major role in the oxygenation of the atmosphere from about 2,400  million years ago . This change in

19312-533: Was 21.9 ± 0.4 hours, and there were 13.1 ± 0.1 synodic months/year and 400 ± 7 solar days/year. A few English language documentaries have featured the Ediacaran Period and biota: Geological period The geologic time scale or geological time scale ( GTS ) is a representation of time based on the rock record of Earth . It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (a scientific branch of geology that aims to determine

19454-429: Was accepted at the time by western religion. Instead, using geological evidence, they contested Earth to be much older, cementing the concept of deep time. During the early 19th century William Smith , Georges Cuvier , Jean d'Omalius d'Halloy , and Alexandre Brongniart pioneered the systematic division of rocks by stratigraphy and fossil assemblages. These geologists began to use the local names given to rock units in

19596-455: Was dominated by acritarchs known as large ornamented Ediacaran microfossils . The second spanned from around 575 to 560 Ma and was characterised by the Avalon biota. The third spanned from 560 to 550 Ma; its biota has been dubbed the White Sea biota due to many fossils from this time being found along the coasts of the White Sea . The fourth lasted from 550 to 539 Ma and is known as the interval of

19738-415: Was in some places unwise, scholars such as Girolamo Fracastoro shared da Vinci's views, and found the attribution of fossils to the 'Deluge' absurd. Niels Stensen, more commonly known as Nicolas Steno (1638–1686), is credited with establishing four of the guiding principles of stratigraphy. In De solido intra solidum naturaliter contento dissertationis prodromus Steno states: Respectively, these are

19880-535: Was not until the Italian Renaissance when Leonardo da Vinci (1452–1519) would reinvigorate the relationships between stratification, relative sea-level change, and time, denouncing attribution of fossils to the 'Deluge': Of the stupidity and ignorance of those who imagine that these creatures were carried to such places distant from the sea by the Deluge...Why do we find so many fragments and whole shells between

20022-774: Was ratified in 2004 by the International Union of Geological Sciences (IUGS), making it the first new geological period declared in 120 years. Although the period took namesake from the Ediacara Hills in the Nilpena Ediacara National Park , the type section is actually located in the bed of the Enorama Creek within the Brachina Gorge in the Ikara-Flinders Ranges National Park , at 31°19′53.8″S 138°38′0.1″E  /  31.331611°S 138.633361°E  / -31.331611; 138.633361 , approximately 55 km (34 mi) southeast of

20164-476: Was the last commercial publication of an international chronostratigraphic chart that was closely associated with the ICS. It included a proposal to substantially revise the pre-Cryogenian time scale to reflect important events such as the formation of the Solar System and the Great Oxidation Event , among others, while at the same time maintaining most of the previous chronostratigraphic nomenclature for

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