192-496: The coyote ( Canis latrans ), also known as the American jackal , prairie wolf , or brush wolf , is a species of canine native to North America . It is smaller than its close relative, the gray wolf , and slightly smaller than the closely related eastern wolf and red wolf . It fills much of the same ecological niche as the golden jackal does in Eurasia ; however, the coyote
384-530: A mammal adapted to desert life. It minimizes its water loss by producing concentrated urine and dry dung , and is able to lose 40% of its body weight through water loss without dying of dehydration. Carnivores can obtain much of their water needs from the body fluids of their prey. Many other hot desert animals are nocturnal , seeking out shade during the day or dwelling underground in burrows. At depths of more than 50 cm (20 in), these remain at between 30 and 32 °C (86 and 90 °F) regardless of
576-403: A melanocortin 1 receptor mutation inherited from Golden Retrievers . Coyotes have hybridized with wolves to varying degrees, particularly in eastern North America . The so-called " eastern coyote " of northeastern North America probably originated in the aftermath of the extermination of gray and eastern wolves in the northeast, thus allowing coyotes to colonize former wolf ranges and mix with
768-455: A mosaic of smooth stones, often forming desert pavements , and little further erosion takes place. Other desert features include rock outcrops , exposed bedrock and clays once deposited by flowing water. Temporary lakes may form and salt pans may be left when waters evaporate. There may be underground sources of water, in the form of springs and seepages from aquifers . Where these are found, oases can occur. Plants and animals living in
960-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
1152-456: A sand blasting mechanism which grinds away solid objects in its path as the kinetic energy of the wind is transferred to the ground. The sand eventually ends up deposited in level areas known as sand-fields or sand-seas, or piled up in dunes. Many people think of deserts as consisting of extensive areas of billowing sand dunes because that is the way they are often depicted on TV and in films, but deserts do not always look like this. Across
1344-428: A semi-arid climate and are less extreme than regular deserts. Like arid deserts, temperatures can vary greatly in semi deserts. They share some characteristics of a true desert and are usually located at the edge of deserts and continental dry areas. They usually receive precipitation from 250 to 500 mm (9.8 to 19.7 in) but this can vary due to evapotranspiration and soil nutrition. Semi-deserts can be found in
1536-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
1728-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
1920-637: A brightening of 'ochraceous' tones – deep orange or brown – towards the Pacific coast ( C. l. ochropus , C. l. umpquensis ), a reduction in size in Aridoamerica ( C. l. microdon , C. l. mearnsi ) and a general trend towards dark reddish colors and short muzzles in Mexican and Central American populations. [REDACTED] Coyotes occasionally mate with domestic dogs , sometimes producing crosses colloquially known as " coydogs ". Such matings are rare in
2112-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
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#17328482305322304-513: A deep sea green colour small and piercing. Their [claws] are rather longer than those of the ordinary wolf or that common to the Atlantic states, none of which are to be found in this quarter, nor I believe above the river Plat. The coyote was first scientifically described by naturalist Thomas Say in September ;1819, on the site of Lewis and Clark's Council Bluffs, 24 km (15 mi) up
2496-438: A desert conglomerate . In time, bacteria that live on the surface of the stones accumulate a film of minerals and clay particles, forming a shiny brown coating known as desert varnish . Other non-sandy deserts consist of exposed outcrops of bedrock , dry soils or aridisols , and a variety of landforms affected by flowing water , such as alluvial fans , sinks or playas , temporary or permanent lakes , and oases. A hamada
2688-505: A destructive effect on the exposed rocky surfaces. The repeated fluctuations put a strain on exposed rock and the flanks of mountains crack and shatter. Fragmented strata slide down into the valleys where they continue to break into pieces due to the relentless sun by day and chill by night. Successive strata are exposed to further weathering. The relief of the internal pressure that has built up in rocks that have been underground for aeons can cause them to shatter. Exfoliation also occurs when
2880-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
3072-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
3264-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On
3456-596: A local priest, noted that the "wolves" encountered there were smaller and less daring than European wolves. Another account from the early 1800s in Edwards County mentioned wolves howling at night, though these were likely coyotes. This species was encountered several times during the Lewis and Clark Expedition (1804–1806), though it was already well known to European traders on the upper Missouri . Meriwether Lewis , writing on 5 May 1805, in northeastern Montana , described
3648-415: A low basal metabolic rate and by remaining underground during the heat of the day, reducing loss of water through its skin and respiratory system when at rest. Herbivorous mammals obtain moisture from the plants they eat. Species such as the addax antelope , dik-dik , Grant's gazelle and oryx are so efficient at doing this that they apparently never need to drink. The camel is a superb example of
3840-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
4032-420: A more slender skull and skeleton than the modern coyote. Ronald Nowak found that the early populations had small, delicate, narrowly proportioned skulls that resemble small coyotes and appear to be ancestral to C. latrans . C. lepophagus was similar in weight to modern coyotes, but had shorter limb bones that indicate a less cursorial lifestyle. The coyote represents a more primitive form of Canis than
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#17328482305324224-484: A number of ways, generally combining total precipitation, number of days on which this falls, temperature, and humidity, and sometimes additional factors. For example, Phoenix, Arizona , receives less than 250 mm (9.8 in) of precipitation per year, and is immediately recognized as being located in a desert because of its aridity-adapted plants. The North Slope of Alaska's Brooks Range also receives less than 250 mm (9.8 in) of precipitation per year and
4416-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
4608-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that
4800-433: A person approaches them they frequently bark, their note being precisely that of the small dog. They are of an intermediate size between that of the fox and dog, very active fleet and delicately formed; the ears large erect and pointed the head long and pointed more like that of the fox; tale long ... the hair and fur also resembles the fox, tho' is much coarser and inferior. They are of a pale reddish-brown colour. The eye of
4992-536: A reference to the many vocalizations they produce. ᒣᐢᒐᒑᑲᓂᐢ ( Mescacâkanis ) Perro de monte Isapaippü Itsappü Sedet Domestic dog [REDACTED] Gray wolf [REDACTED] Coyote [REDACTED] African wolf [REDACTED] Golden jackal [REDACTED] Ethiopian wolf [REDACTED] Dhole [REDACTED] African wild dog [REDACTED] Side-striped jackal [REDACTED] Black-backed jackal [REDACTED] Xiaoming Wang and Richard H. Tedford , one of
5184-423: A series of long, linear dunes known as seif dunes may form. These also occur parallel to a strong wind that blows in one general direction. Transverse dunes run at a right angle to the prevailing wind direction. Star dunes are formed by variable winds, and have several ridges and slip faces radiating from a central point. They tend to grow vertically; they can reach a height of 500 m (1,600 ft), making them
5376-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
5568-461: A sparse trophic network . Plant growth is limited by rainfall , temperature extremes and desiccating winds. Deserts have strong temporal variability in the availability of resources due to the total amount of annual rainfall and the size of individual rainfall events. Resources are often ephemeral or episodic, and this triggers sporadic animal movements and 'pulse and reserve' or 'boom-bust' ecosystem dynamics. Erosion and sedimentation are high due to
5760-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
5952-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
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6144-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
6336-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
6528-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
6720-439: A steady wind begins to blow, fine particles lying on the exposed ground begin to vibrate. At greater wind speeds, some particles are lifted into the air stream. When they land, they strike other particles which may be jerked into the air in their turn, starting a chain reaction . Once ejected, these particles move in one of three possible ways, depending on their size, shape and density; suspension , saltation or creep. Suspension
6912-571: A storm that may be many kilometers away. Most deserts are in basins with no drainage to the sea but some are crossed by exotic rivers sourced in mountain ranges or other high rainfall areas beyond their borders. The River Nile , the Colorado River and the Yellow River do this, losing much of their water through evaporation as they pass through the desert and raising groundwater levels nearby. There may also be underground sources of water in deserts in
7104-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
7296-485: A tolerable level. In many ways, birds are the ablest to do this of the higher animals. They can move to areas of greater food availability as the desert blooms after local rainfall and can fly to faraway waterholes. In hot deserts, gliding birds can remove themselves from the over-heated desert floor by using thermals to soar in the cooler air at great heights. In order to conserve energy, other desert birds run rather than fly. The cream-colored courser flits gracefully across
7488-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
7680-632: A type of cold desert . While they do not lack water, having a persistent cover of snow and ice, this is merely due to marginal evaporation rates and low precipitation. The McMurdo dry valleys of Antarctica, which lack water (whether rain, ice, or snow) much like a non-polar desert and even have such desert features as hypersaline lakes and intermittent streams that resemble (except for being frozen at their surfaces) hot or cold deserts for extreme aridity and lack of precipitation of any kind. Extreme winds and not seasonal heat desiccate these nearly-lifeless terrains. The concept of "biological desert" redefines
7872-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
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8064-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
8256-400: A year; they have no annual seasonal cycle of precipitation and experience twelve-month periods with no rainfall at all. Arid deserts receive between 25 and 200 mm (1 and 8 in) in a year and semiarid deserts between 200 and 500 mm (8 and 20 in). However, such factors as the temperature, humidity, rate of evaporation and evapotranspiration, and the moisture storage capacity of
8448-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
8640-465: Is 150 km (93 mi) long and is the largest oasis in the Libyan Desert. A lake occupied this depression in ancient times and thick deposits of sandy-clay resulted. Wells are dug to extract water from the porous sandstone that lies underneath. Seepages may occur in the walls of canyons and pools may survive in deep shade near the dried up watercourse below. Lakes may form in basins where there
8832-563: Is a howl made by solitary individuals. Humans are the coyote's greatest threat, followed by cougars and gray wolves. Despite predation by gray wolves, coyotes sometimes mate with them, and with eastern, or red wolves, producing " coywolf " hybrids. In the northeastern regions of North America, the eastern coyote (a larger subspecies, though still smaller than wolves) is the result of various historical and recent matings with various types of wolves. Genetic studies show that most North American wolves contain some level of coyote DNA. The coyote
9024-563: Is a Spanish borrowing of its Nahuatl name coyōtl pronunciation ) comes from the historian Francisco Javier Clavijero 's Historia de México in 1780. The first time it was used in English occurred in William Bullock 's Six months' residence and travels in Mexico (1824), where it is variously transcribed as cayjotte and cocyotie . The word's spelling was standardized as "coyote" by
9216-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
9408-525: Is a jumble of eroded sandstone outcrops , canyons , blocks, pinnacles , fissures , slabs and ravines . In some places the wind has carved holes or arches, and in others, it has created mushroom-like pillars narrower at the base than the top. On the Colorado Plateau , it is water that has been the prevailing eroding force. Here, rivers, such as the Colorado , have cut their way over the millennia through
9600-452: Is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use
9792-579: Is a prominent character in Native American folklore, mainly in Aridoamerica , usually depicted as a trickster that alternately assumes the form of an actual coyote or a man. As with other trickster figures, the coyote uses deception and humor to rebel against social conventions. The animal was especially respected in Mesoamerican cosmology as a symbol of military might. After the European colonization of
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#17328482305329984-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
10176-532: Is a type of desert landscape consisting of a high rocky plateau where the sand has been removed by aeolian processes . Other landforms include plains largely covered by gravels and angular boulders, from which the finer particles have been stripped by the wind. These are called "reg" in the western Sahara, "serir" in the eastern Sahara, "gibber plains" in Australia and "saï" in central Asia. The Tassili Plateau in Algeria
10368-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
10560-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,
10752-573: Is continuous above 5,600 m (18,400 ft). Nevertheless, there is some plant life in the Atacama, in the form of specialist plants that obtain moisture from dew and the fogs that blow in from the Pacific. When rain falls in deserts, as it occasionally does, it is often with great violence. The desert surface is evidence of this with dry stream channels known as arroyos or wadis meandering across its surface. These can experience flash floods , becoming raging torrents with surprising rapidity after
10944-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
11136-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
11328-535: Is generally larger. The coyote is listed as least concern by the International Union for Conservation of Nature , due to its wide distribution and abundance throughout North America. The species is versatile, able to adapt to and expand into environments modified by humans; urban coyotes are common in many cities. The coyote was sighted in eastern Panama (across the Panama Canal from their home range) for
11520-576: Is gray-brown, with reddish legs, ears, and flanks. No significant differences exist between eastern and western coyotes in aggression and fighting, though eastern coyotes tend to fight less, and are more playful. Unlike western coyote pups, in which fighting precedes play behavior, fighting among eastern coyote pups occurs after the onset of play. Eastern coyotes tend to reach sexual maturity at two years of age, much later than in western coyotes. Eastern and red wolves are also products of varying degrees of wolf-coyote hybridization. The eastern wolf probably
11712-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
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#173284823053211904-479: Is higher than that of forests or the sea. A desert is a region of land that is very dry because it receives low amounts of precipitation (usually in the form of rain, but it may be snow, mist or fog), often has little coverage by plants, and in which streams dry up unless they are supplied by water from outside the area. Deserts generally receive less than 250 mm (10 in) of precipitation each year. The potential evapotranspiration may be large but (in
12096-494: Is longer and denser than in southern forms, with the fur of some Mexican and Central American forms being almost hispid (bristly). Generally, adult coyotes (including coywolf hybrids) have a sable coat color, dark neonatal coat color, bushy tail with an active supracaudal gland , and a white facial mask. Albinism is extremely rare in coyotes. Out of a total of 750,000 coyotes killed by federal and cooperative hunters between March 1938 and June 1945, only two were albinos. The coyote
12288-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
12480-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
12672-551: Is often classified as a cold desert. Other regions of the world have cold deserts, including areas of the Himalayas and other high-altitude areas in other parts of the world. Polar deserts cover much of the ice-free areas of the Arctic and Antarctic. A non-technical definition is that deserts are those parts of Earth's surface that have insufficient vegetation cover to support a human population. Potential evapotranspiration supplements
12864-434: Is often less than 40 mm or 1.5 in) to being very far from the nearest available sources of moisture and are often in the lee of mountain ranges. Montane deserts are normally cold, or may be scorchingly hot by day and very cold by night as is true of the northeastern slopes of Mount Kilimanjaro . Polar deserts such as McMurdo Dry Valleys remain ice-free because of the dry katabatic winds that flow downhill from
13056-421: Is only possible for particles less than 0.1 mm (0.0039 in) in diameter. In a dust storm, these fine particles are lifted up and wafted aloft to heights of up to 6 km (3.7 mi). They reduce visibility and can remain in the atmosphere for days on end, conveyed by the trade winds for distances of up to 6,000 km (3,700 mi). Denser clouds of dust can be formed in stronger winds, moving across
13248-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
13440-538: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes
13632-486: Is sufficient precipitation or meltwater from glaciers above. They are usually shallow and saline, and wind blowing over their surface can cause stress, moving the water over nearby low-lying areas. When the lakes dry up, they leave a crust or hardpan behind. This area of deposited clay, silt or sand is known as a playa . The deserts of North America have more than one hundred playas, many of them relics of Lake Bonneville which covered parts of Utah, Nevada and Idaho during
13824-657: Is the Atacama Desert . It is virtually devoid of life because it is blocked from receiving precipitation by the Andes mountains to the east and the Chilean Coast Range to the west. The cold Humboldt Current and the anticyclone of the Pacific are essential to keep the dry climate of the Atacama. The average precipitation in the Chilean region of Antofagasta is just 1 mm (0.039 in) per year. Some weather stations in
14016-450: Is the key to plant growth. It can only take place during the day as energy from the sun is required, but during the day, many deserts become very hot. Opening stomata to allow in the carbon dioxide necessary for the process causes evapotranspiration , and conservation of water is a top priority for desert vegetation. Some plants have resolved this problem by adopting crassulacean acid metabolism , allowing them to open their stomata during
14208-431: Is the specializations of mammalian kidneys shown by desert-inhabiting species. Many examples of convergent evolution have been identified in desert organisms, including between cacti and Euphorbia , kangaroo rats and jerboas , Phrynosoma and Moloch lizards. Deserts present a very challenging environment for animals. Not only do they require food and water but they also need to keep their body temperature at
14400-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
14592-508: Is typically smaller than the gray wolf, but has longer ears and a relatively larger braincase , as well as a thinner frame, face, and muzzle. The scent glands are smaller than the gray wolf's, but are the same color. Its fur color variation is much less varied than that of a wolf. The coyote also carries its tail downwards when running or walking, rather than horizontally as the wolf does. Coyote tracks can be distinguished from those of dogs by their more elongated, less rounded shape. Unlike dogs,
14784-528: Is very cold and carries little moisture so little precipitation occurs and what does fall, usually snow, is carried along in the often strong wind and may form blizzards, drifts and dunes similar to those caused by dust and sand in other desert regions. In Antarctica , for example, the annual precipitation is about 50 mm (2 in) on the central plateau and some ten times that amount on some major peninsulas. Based on precipitation alone, hyperarid deserts receive less than 25 mm (1 in) of rainfall
14976-669: The Atacama Desert . Deserts of this type are often long and narrow and bounded to the east by mountain ranges. They occur in Namibia , Chile , southern California and Baja California . Other coastal deserts influenced by cold currents are found in Western Australia , the Arabian Peninsula and Horn of Africa , and the western fringes of the Sahara. In 1961, Peveril Meigs divided desert regions on Earth into three categories according to
15168-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
15360-521: The Quaternary extinction event . Compared to their modern Holocene counterparts, Pleistocene coyotes ( C. l. orcutti ) were larger and more robust, likely in response to larger competitors and prey. Pleistocene coyotes were likely more specialized carnivores than their descendants, as their teeth were more adapted to shearing meat, showing fewer grinding surfaces suited for processing vegetation. Their reduction in size occurred within 1,000 years of
15552-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
15744-616: The horse latitudes at 30° to 35° North and South. These belts are associated with the subtropical anticyclone and the large-scale descent of dry air. The Sahara Desert is of this type. Mid-latitude deserts occur between 30° and 50° North and South. They are mostly in areas remote from the sea where most of the moisture has already precipitated from the prevailing winds. They include the Tengger and Sonoran Deserts . Monsoon deserts are similar. They occur in regions where large temperature differences occur between sea and land. Moist warm air rises over
15936-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
16128-414: The rocks , which consequently break in pieces. Although rain seldom occurs in deserts, there are occasional downpours that can result in flash floods. Rain falling on hot rocks can cause them to shatter, and the resulting fragments and rubble strewn over the desert floor are further eroded by the wind. This picks up particles of sand and dust, which can remain airborne for extended periods – sometimes causing
16320-417: The windward slope of the mountain range . When they descend on the leeward side, they warm and their capacity to hold moisture increases so an area with relatively little precipitation occurs. The Taklamakan Desert is an example, lying in the rain shadow of the Himalayas and receiving less than 38 mm (1.5 in) precipitation annually. Other areas are arid by virtue of being a very long way from
16512-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
16704-594: The 1880s. The English pronunciation is heard both as a two-syllable word (with the final "e" silent) and as three-syllables (with the final "e" pronounced), with a tendency for the three-syllable pronunciation in eastern states and near the Mexican border, and outside the United States, with two syllables in western and central states. Alternative English names for the coyote include "prairie wolf", "brush wolf", "cased wolf", "little wolf" and "American jackal". Its binomial name Canis latrans translates to "barking dog",
16896-562: The Americas , it was seen in Anglo-American culture as a cowardly and untrustworthy animal. Unlike wolves, which have seen their public image improve, attitudes towards the coyote remain largely negative. Coyote males average 8 to 20 kg (18 to 44 lb) in weight, while females average 7 to 18 kg (15 to 40 lb), though size varies geographically. Northern subspecies, which average 18 kg (40 lb), tend to grow larger than
17088-434: The Atacama have never received rain. Evidence suggests that the Atacama may not have had any significant rainfall from 1570 to 1971. It is so arid that mountains that reach as high as 6,885 m (22,589 ft) are completely free of glaciers and, in the southern part from 25°S to 27°S, may have been glacier-free throughout the Quaternary , though permafrost extends down to an altitude of 4,400 m (14,400 ft) and
17280-805: The Eurasian golden jackal, the coyote is gregarious, but not as dependent on conspecifics as more social canid species like wolves are. This is likely because the coyote is not a specialized hunter of large prey as the latter species is. The basic social unit of a coyote pack is a family containing a reproductive female. However, unrelated coyotes may join forces for companionship, or to bring down prey too large to attack on their own. Such "nonfamily" packs are only temporary, and may consist of bachelor males, nonreproductive females and subadult young. Families are formed in midwinter, when females enter estrus . Pair bonding can occur 2–3 months before actual copulation takes place. Species A species ( pl. : species)
17472-556: The Missouri River from the mouth of the Platte during a government-sponsored expedition with Major Stephen Long . He had the first edition of the Lewis and Clark journals in hand, which contained Biddle's edited version of Lewis's observations dated 5 May 1805. His account was published in 1823. Say was the first person to document the difference between a " prairie wolf " (coyote) and on
17664-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
17856-467: The Quaternary extinction event, when their large prey died out. Furthermore, Pleistocene coyotes were unable to exploit the big-game hunting niche left vacant after the extinction of the dire wolf ( Aenocyon dirus ), as it was rapidly filled by gray wolves, which likely actively killed off the large coyotes, with natural selection favoring the modern gracile morph. In 1993, a study proposed that
18048-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
18240-517: The absence of available water) the actual evapotranspiration may be close to zero. Semi-deserts are regions which receive between 250 and 500 mm (10 and 20 in) and when clad in grass, these are known as steppes . Most deserts on Earth such as the Sahara Desert , Grand Australian Desert and the Great Basin Desert , occur in low altitudes . One of the driest places on Earth
18432-411: The age of 35 days, with eastern coyote pups having longer legs than their western counterparts. Differences in dental development also occurs, with tooth eruption being later, and in a different order in the eastern coyote. Aside from its size, the eastern coyote is physically similar to the western coyote. The four color phases range from dark brown to blond or reddish blond, though the most common phase
18624-401: The air but soon fall back to earth, ejecting other particles in the process. Their weight prevents them from being airborne for long and most only travel a distance of a few meters (yards). The sand streams along above the surface of the ground like a fluid, often rising to heights of about 30 cm (12 in). In a really severe steady blow, 2 m (6 ft 7 in) is about as high as
18816-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
19008-722: The amount of precipitation they received. In this now widely accepted system, extremely arid lands have at least twelve consecutive months without precipitation, arid lands have less than 250 mm (9.8 in) of annual precipitation, and semiarid lands have a mean annual precipitation of between 250 and 500 mm (9.8 and 19.7 in). Both extremely arid and arid lands are considered to be deserts while semiarid lands are generally referred to as steppes when they are grasslands. Deserts are also classified, according to their geographical location and dominant weather pattern, as trade wind, mid-latitude, rain shadow, coastal, monsoon, or polar deserts . Trade wind deserts occur either side of
19200-409: The anterior ones near the wrist; tail bushy, fusiform, straight, varied with gray and cinnamon, a spot near the base above, and tip black; the tip of the trunk of the tail, attains the tip of the os calcis , when the leg is extended; beneath white, immaculate, tail cinnamon towards the tip, tip black; posterior feet four toed, anterior five toed. The first published usage of the word "coyote" (which
19392-492: The assumptions made, that all of the North American wolves and coyotes diverged from a common ancestor about 51,000 years ago. However, the proposed timing of the wolf / coyote divergence conflicts with the discovery of a coyote-like specimen in strata dated to 1 Mya. The study also indicated that all North American wolves have a significant amount of coyote ancestry and all coyotes some degree of wolf ancestry, and that
19584-411: The barren rock is to be found in the so-called Dry Valleys of Antarctica that almost never get snow, which can have ice-encrusted saline lakes that suggest evaporation far greater than the rare snowfall due to the strong katabatic winds that even evaporate ice. Deserts, both hot and cold, play a part in moderating Earth's temperature, because they reflect more of the incoming light and their albedo
19776-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
19968-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
20160-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be
20352-457: The capture of large quantities of solar energy . English desert and its Romance cognates (including Italian and Portuguese deserto , French désert and Spanish desierto ) all come from the ecclesiastical Latin dēsertum (originally "an abandoned place"), a participle of dēserere , "to abandon". The correlation between aridity and sparse population is complex and dynamic, varying by culture, era, and technologies; thus
20544-516: The cellular structure of which has been modified to allow them to store water. When rain falls, the water is rapidly absorbed by the shallow roots and retained to allow them to survive until the next downpour, which may be months or years away. The giant saguaro cacti of the Sonoran Desert form "forests", providing shade for other plants and nesting places for desert birds. Saguaro grows slowly but may live for up to two hundred years. The surface of
20736-615: The coldest part of the year. Animals adapted to live in deserts are called xerocoles . There is no evidence that body temperature of mammals and birds is adaptive to the different climates, either of great heat or cold. In fact, with a very few exceptions, their basal metabolic rate is determined by body size, irrespective of the climate in which they live. Many desert animals (and plants) show especially clear evolutionary adaptations for water conservation or heat tolerance and so are often studied in comparative physiology , ecophysiology , and evolutionary physiology . One well-studied example
20928-597: The collision between airborne particles and by the impacts of saltating sand grains landing on the ground. The mechanism is little understood but the particles usually have a negative charge when their diameter is under 250 μm and a positive one when they are over 500 μm. Deserts and semi-deserts are home to ecosystems with low or very low biomass and primary productivity in arid or semi-arid climates. They are mostly found in subtropical high-pressure belts and major continental rain shadows . Primary productivity depends on low densities of small photoautotrophs that sustain
21120-586: The color of the underlying surface. The desert lark takes frequent dust baths which ensures that it matches its environment. Water and carbon dioxide are metabolic end products of oxidation of fats, proteins, and carbohydrates. Oxidising a gram of carbohydrate produces 0.60 grams of water; a gram of protein produces 0.41 grams of water; and a gram of fat produces 1.07 grams of water, making it possible for xerocoles to live with little or no access to drinking water. The kangaroo rat for example makes use of this water of metabolism and conserves water both by having
21312-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
21504-468: The concept of desert, without the characteristic of aridity, not lacking water, but instead lacking life. Such places can be so-called "ocean deserts", which are mostly at the centers of gyres , but also hypoxic or anoxic waters such as dead zones . Deserts usually have a large diurnal and seasonal temperature range, with high daytime temperatures falling sharply at night. The diurnal range may be as much as 20 to 30 °C (36 to 54 °F) and
21696-416: The concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part
21888-506: The coyote in these terms: The small wolf or burrowing dog of the prairies are the inhabitants almost invariably of the open plains; they usually associate in bands of ten or twelve sometimes more and burrow near some pass or place much frequented by game; not being able alone to take deer or goat they are rarely ever found alone but hunt in bands; they frequently watch and seize their prey near their burrows; in these burrows, they raise their young and to them they also resort when pursued; when
22080-580: The coyote's usual characteristics . F 1 hybrids tend to be intermediate in form between dogs and coyotes, while F 2 hybrids (second generation) are more varied. Both F 1 and F 2 hybrids resemble their coyote parents in terms of shyness and intrasexual aggression. Hybrids are fertile and can be successfully bred through four generations. Melanistic coyotes owe their black pelts to a mutation that first arose in domestic dogs. A population of non-albino white coyotes in Newfoundland owe their coloration to
22272-452: The desert floor to scarify the seed coat. The seed of the mesquite tree, which grows in deserts in the Americas, is hard and fails to sprout even when planted carefully. When it has passed through the gut of a pronghorn it germinates readily, and the little pile of moist dung provides an excellent start to life well away from the parent tree. The stems and leaves of some plants lower the surface velocity of sand-carrying winds and protect
22464-500: The desert need special adaptations to survive in the harsh environment. Plants tend to be tough and wiry with small or no leaves, water-resistant cuticles , and often spines to deter herbivory . Some annual plants germinate , bloom and die in the course of a few weeks after rainfall, while other long-lived plants survive for years and have deep root systems able to tap underground moisture. Animals need to keep cool and find enough food and water to survive. Many are nocturnal , and stay in
22656-587: The driest season, and others curl their leaves up to reduce transpiration. Others, such as aloes , store water in succulent leaves or stems or in fleshy tubers. Desert plants maximize water uptake by having shallow roots that spread widely, or by developing long taproots that reach down to deep rock strata for ground water. The saltbush in Australia has succulent leaves and secretes salt crystals, enabling it to live in saline areas. In common with cacti, many have developed spines to ward off browsing animals. Some desert plants produce seed which lies dormant in
22848-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
23040-455: The external temperature. Jerboas , desert rats , kangaroo rats and other small rodents emerge from their burrows at night and so do the foxes, coyotes, jackals and snakes that prey on them. Kangaroos keep cool by increasing their respiration rate, panting, sweating and moistening the skin of their forelegs with saliva . Mammals living in cold deserts have developed greater insulation through warmer body fur and insulating layers of fat beneath
23232-702: The first time in 2013. The coyote has 19 recognized subspecies . The average male weighs 8 to 20 kg (18 to 44 lb) and the average female 7 to 18 kg (15 to 40 lb). Their fur color is predominantly light gray and red or fulvous interspersed with black and white, though it varies somewhat with geography. It is highly flexible in social organization, living either in a family unit or in loosely knit packs of unrelated individuals. Primarily carnivorous , its diet consists mainly of deer , rabbits , hares , rodents , birds , reptiles , amphibians , fish , and invertebrates , though it may also eat fruits and vegetables on occasion. Its characteristic vocalization
23424-526: The foremost authorities on carnivore evolution, proposed that the genus Canis was the descendant of the coyote-like Eucyon davisi and its remains first appeared in the Miocene 6 million years ago (Mya) in the southwestern US and Mexico. By the Pliocene (5 Mya), the larger Canis lepophagus appeared in the same region and by the early Pleistocene (1 Mya) C. latrans (the coyote)
23616-561: The form of springs , aquifers , underground rivers or lakes. Where these lie close to the surface, wells can be dug and oases may form where plant and animal life can flourish. The Nubian Sandstone Aquifer System under the Sahara Desert is the largest known accumulation of fossil water . The Great Man-Made River is a scheme launched by Libya's Muammar Gaddafi to tap this aquifer and supply water to coastal cities. Kharga Oasis in Egypt
23808-407: The formation of sand storms or dust storms . Wind-blown sand grains striking any solid object in their path can abrade the surface. Rocks are smoothed down, and the wind sorts sand into uniform deposits. The grains end up as level sheets of sand or are piled high in billowing sand dunes . Other deserts are flat, stony plains where all the fine material has been blown away and the surface consists of
24000-458: The fragmentation of harder granites , limestone and sandstone . There is a certain critical size (about 0.5 mm) below which further temperature-induced weathering of rocks does not occur and this provides a minimum size for sand grains. As the mountains are eroded, more and more sand is created. At high wind speeds, sand grains are picked up off the surface and blown along, a process known as saltation . The whirling airborne grains act as
24192-403: The gray wolf, as shown by its relatively small size and its comparatively narrow skull and jaws, which lack the grasping power necessary to hold the large prey in which wolves specialize. This is further corroborated by the coyote's sagittal crest , which is low or totally flattened, thus indicating a weaker bite than the wolves. The coyote is not a specialized carnivore as the wolf is, as shown by
24384-439: The ground from erosion. Even small fungi and microscopic plant organisms found on the soil surface (so-called cryptobiotic soil ) can be a vital link in preventing erosion and providing support for other living organisms. Cold deserts often have high concentrations of salt in the soil. Grasses and low shrubs are the dominant vegetation here and the ground may be covered with lichens . Most shrubs have spiny leaves and shed them in
24576-474: The ground have a marked effect on the degree of aridity and the plant and animal life that can be sustained. Rain falling in the cold season may be more effective at promoting plant growth, and defining the boundaries of deserts and the semiarid regions that surround them on the grounds of precipitation alone is problematic. A semi-arid desert or a steppe is a version of the arid desert with much more rainfall, vegetation and higher humidity. These regions feature
24768-432: The ground on its long legs, stopping periodically to snatch up insects. Like other desert birds, it is well-camouflaged by its coloring and can merge into the landscape when stationary. The sandgrouse is an expert at this and nests on the open desert floor dozens of kilometers (miles) away from the waterhole it needs to visit daily. Some small diurnal birds are found in very restricted localities where their plumage matches
24960-427: The ground rises quickly through a small pocket of cooler, low-pressure air above forming a whirling column of particles, a dust devil . Sandstorms occur with much less frequency than dust storms. They are often preceded by severe dust storms and occur when the wind velocity increases to a point where it can lift heavier particles. These grains of sand, up to about 0.5 mm (0.020 in) in diameter are jerked into
25152-429: The high desert floor, creating canyons that are over a mile (6,000 feet or 1,800 meters) deep in places, exposing strata that are over two billion years old. Sand and dust storms are natural events that occur in arid regions where the land is not protected by a covering of vegetation. Dust storms usually start in desert margins rather than the deserts themselves where the finer materials have already been blown away. As
25344-746: The high elevations of the Tabernas Desert (and some parts of the Spanish Plateu ), The Sahel , The Eurasian Steppe , most of Central Asia , the Western US , most of Northern Mexico , portions of South America (especially in Argentina ) and the Australian Outback . They usually feature BSh (hot steppe) or BSk (temperate steppe) in the Köppen climate classification . Coastal deserts are mostly found on
25536-450: The import of water from an outside source. Many trade routes have been forged across deserts, especially across the Sahara , and traditionally were used by caravans of camels carrying salt, gold, ivory and other goods. Large numbers of slaves were also taken northwards across the Sahara. Some mineral extraction also takes place in deserts, and the uninterrupted sunlight gives potential for
25728-428: The lachrymal sac black-brown; rostrum cinnamon, tinctured with grayish on the nose; lips white, edged with black, three series of black seta; head between the ears intermixed with gray, and dull cinnamon, hairs dusky plumbeous at base; sides paler than the back, obsoletely fasciate with black above the legs; legs cinnamon on the outer side, more distinct on the posterior hair: a dilated black abbreviated line on
25920-523: The lack of heat to aid in the evaporation process. Deserts are sometimes classified as "hot" or "cold", "semiarid" or "coastal". The characteristics of hot deserts include high temperatures in summer; greater evaporation than precipitation, usually exacerbated by high temperatures, strong winds and lack of cloud cover; considerable variation in the occurrence of precipitation, its intensity and distribution; and low humidity. Winter temperatures vary considerably between different deserts and are often related to
26112-479: The land with a billowing leading edge. The sunlight can be obliterated and it may become as dark as night at ground level. In a study of a dust storm in China in 2001, it was estimated that 6.5 million tons of dust were involved, covering an area of 134,000,000 km (52,000,000 sq mi). The mean particle size was 1.44 μm. A much smaller scale, short-lived phenomenon can occur in calm conditions when hot air near
26304-469: The land, deposits its water content and circulates back to sea. Further inland, areas receive very little precipitation. The Thar Desert near the India/Pakistan border is of this type. In some parts of the world, deserts are created by a rain shadow effect. Orographic lift occurs as air masses rise to pass over high ground. In the process they cool and lose much of their moisture by precipitation on
26496-727: The larger chewing surfaces on the molars , reflecting the species' relative dependence on vegetable matter. In these respects, the coyote resembles the fox-like progenitors of the genus more so than the wolf. The oldest fossils that fall within the range of the modern coyote date to 0.74–0.85 Ma (million years) in Hamilton Cave, West Virginia; 0.73 Ma in Irvington, California; 0.35–0.48 Ma in Porcupine Cave, Colorado, and in Cumberland Cave, Pennsylvania. Modern coyotes arose 1,000 years after
26688-499: The largest dataset of nuclear genome sequences against the wolf reference genome. The study supports the findings of previous studies that North American gray wolves and wolf-like canids were the result of complex gray wolf and coyote mixing. A polar wolf from Greenland and a coyote from Mexico represented the purest specimens. The coyotes from Alaska, California, Alabama, and Quebec show almost no wolf ancestry. Coyotes from Missouri, Illinois, and Florida exhibit 5–10% wolf ancestry. There
26880-692: The last ice age when the climate was colder and wetter. These include the Great Salt Lake , Utah Lake , Sevier Lake and many dry lake beds. The smooth flat surfaces of playas have been used for attempted vehicle speed records at Black Rock Desert and Bonneville Speedway and the United States Air Force uses Rogers Dry Lake in the Mojave Desert as runways for aircraft and the Space Shuttle . Deserts have been defined and classified in
27072-510: The life processes of plants. Potential evapotranspiration , then, is the amount of water that could evaporate in any given region. As an example, Tucson, Arizona receives about 300 mm (12 in) of rain per year, however about 2,500 mm (98 in) of water could evaporate over the course of a year. In other words, about eight times more water could evaporate from the region than actually falls as rain. Rates of evapotranspiration in cold regions such as Alaska are much lower because of
27264-412: The location of the desert on the continental landmass and the latitude. Daily variations in temperature can be as great as 22 °C (40 °F) or more, with heat loss by radiation at night being increased by the clear skies. Cold deserts, sometimes known as temperate deserts, occur at higher latitudes than hot deserts, and the aridity is caused by the dryness of the air. Some cold deserts are far from
27456-404: The measurement of precipitation in providing a scientific measurement-based definition of a desert. The water budget of an area can be calculated using the formula P − PE ± S , wherein P is precipitation, PE is potential evapotranspiration rates and S is the amount of surface storage of water. Evapotranspiration is the combination of water loss through atmospheric evaporation and through
27648-597: The nearest available sources of moisture. Montane deserts are arid places with a very high altitude ; the most prominent example is found north of the Himalayas, in the Kunlun Mountains and the Tibetan Plateau . Many locations within this category have elevations exceeding 3,000 m (9,800 ft) and the thermal regime can be hemiboreal . These places owe their profound aridity (the average annual precipitation
27840-663: The next page of his journal a wolf which he named Canis nubilus ( Great Plains wolf ). Say described the coyote as: Canis latrans . Cinereous or gray, varied with black above, and dull fulvous, or cinnamon; hair at base dusky plumbeous, in the middle of its length dull cinnamon, and at tip gray or black, longer on the vertebral line; ears erect, rounded at tip, cinnamon behind, the hair dark plumbeous at base, inside lined with gray hair; eyelids edged with black, superior eyelashes black beneath, and at tip above; supplemental lid margined with black-brown before, and edged with black brown behind; iris yellow; pupil black-blue; spot upon
28032-409: The night to allow CO 2 to enter, and close them during the day, or by using C4 carbon fixation . Many desert plants have reduced the size of their leaves or abandoned them altogether. Cacti are present in both North and South America with a post-Gondwana origin. The genus is desert specialist, and in most species, the leaves have been dispensed with and the chlorophyll displaced into the trunks,
28224-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
28416-514: The ocean and others are separated by mountain ranges from the sea, and in both cases, there is insufficient moisture in the air to cause much precipitation. The largest of these deserts are found in Central Asia. Others occur on the eastern side of the Rocky Mountains , the eastern side of the southern Andes and in southern Australia. Polar deserts are a particular class of cold desert. The air
28608-426: The outer surfaces of rocks split off in flat flakes. This is believed to be caused by the stresses put on the rock by repeated thermal expansions and contractions which induces fracturing parallel to the original surface. Chemical weathering processes probably play a more important role in deserts than was previously thought. The necessary moisture may be present in the form of dew or mist. Ground water may be drawn to
28800-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
28992-466: The pavement is formed. It may be that after the sand and dust is blown away by the wind the stones jiggle themselves into place; alternatively, stones previously below ground may in some way work themselves to the surface. Very little further erosion takes place after the formation of a pavement, and the ground becomes stable. Evaporation brings moisture to the surface by capillary action and calcium salts may be precipitated, binding particles together to form
29184-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
29376-595: The red wolf and eastern wolf are highly admixed with different proportions of gray wolf and coyote ancestry. Genetic studies relating to wolves or dogs have inferred phylogenetic relationships based on the only reference genome available, that of the Boxer dog. In 2017, the first reference genome of the wolf Canis lupus lupus was mapped to aid future research. In 2018, a study looked at the genomic structure and admixture of North American wolves, wolf-like canids, and coyotes using specimens from across their entire range that mapped
29568-519: The remnant wolf populations. This hybrid is smaller than either the gray or eastern wolf, and holds smaller territories, but is in turn larger and holds more extensive home ranges than the typical western coyote. As of 2010, the eastern coyote's genetic makeup is fairly uniform, with minimal influence from eastern wolves or western coyotes. Adult eastern coyotes are larger than western coyotes, with female eastern coyotes weighing 21% more than male western coyotes. Physical differences become more apparent by
29760-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
29952-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
30144-438: The rock surface experiences even greater temperature differentials. During the day the sky is usually clear and most of the sun 's radiation reaches the ground, but as soon as the sun sets, the desert cools quickly by radiating heat into space. In hot deserts, the temperature during daytime can exceed 45 °C (113 °F) in summer and plunge below freezing point at night during winter. Such large temperature variations have
30336-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
30528-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
30720-503: The sand stream can rise as the largest sand grains do not become airborne at all. They are transported by creep, being rolled along the desert floor or performing short jumps. During a sandstorm, the wind-blown sand particles become electrically charged . Such electric fields , which range in size up to 80 kV/m, can produce sparks and cause interference with telecommunications equipment. They are also unpleasant for humans and can cause headaches and nausea. The electric fields are caused by
30912-435: The shade or underground during the heat of the day. They tend to be efficient at conserving water, extracting most of their needs from their food and concentrating their urine . Some animals remain in a state of dormancy for long periods, ready to become active again during the rare rainfall. They then reproduce rapidly while conditions are favorable before returning to dormancy. People have struggled to live in deserts and
31104-441: The sheet consists of thin horizontal layers of coarse silt and very fine to medium grain sand, separated by layers of coarse sand and pea-gravel which are a single grain thick. These larger particles anchor the other particles in place and may also be packed together on the surface so as to form a miniature desert pavement. Small ripples form on the sand sheet when the wind exceeds 24 km/h (15 mph). They form perpendicular to
31296-412: The skin. The arctic weasel has a metabolic rate that is two or three times as high as would be expected for an animal of its size. Birds have avoided the problem of losing heat through their feet by not attempting to maintain them at the same temperature as the rest of their bodies, a form of adaptive insulation. The emperor penguin has dense plumage, a downy under layer, an air insulation layer next to
31488-466: The soil until sparked into growth by rainfall. With annuals , such plants grow with great rapidity and may flower and set seed within weeks, aiming to complete their development before the last vestige of water dries up. For perennial plants, reproduction is more likely to be successful if the seed germinates in a shaded position, but not so close to the parent plant as to be in competition with it. Some seed will not germinate until it has been blown about on
31680-520: The southern subspecies of Mexico, which average 11.5 kg (25 lb). Total length ranges on average from 1.0 to 1.35 m (3 ft 3 in to 4 ft 5 in); comprising a tail length of 40 cm (16 in), with females being shorter in both body length and height. The largest coyote on record was a male killed near Afton, Wyoming , on November 19, 1937, which measured 1.5 m (4 ft 11 in) from nose to tail, and weighed 34 kg (75 lb). Scent glands are located at
31872-464: The sparse vegetation cover and the activities of large mammals and people. Plants and animals in deserts are mostly adapted to extreme and prolonged water deficits , but their reproductive phenology often responds to short episodes of surplus. Competitive interactions are weak. Plants face severe challenges in arid environments. Problems they need to solve include how to obtain enough water, how to avoid being eaten and how to reproduce. Photosynthesis
32064-502: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
32256-440: The surface by evaporation and the formation of salt crystals may dislodge rock particles as sand or disintegrate rocks by exfoliation. Shallow caves are sometimes formed at the base of cliffs by this means. As the desert mountains decay, large areas of shattered rock and rubble occur. The process continues and the end products are either dust or sand. Dust is formed from solidified clay or volcanic deposits whereas sand results from
32448-464: The surface of the ground. Dunes are sometimes solitary, but they are more often grouped together in dune fields. When these are extensive, they are known as sand seas or ergs . The shape of the dune depends on the characteristics of the prevailing wind. Barchan dunes are produced by strong winds blowing across a level surface and are crescent-shaped with the concave side away from the wind. When there are two directions from which winds regularly blow,
32640-561: The surrounding mountains. Former desert areas presently in non-arid environments, such as the Sandhills in Nebraska , are known as paleodeserts. In the Köppen climate classification system, deserts are classed as BWh (hot desert) or BWk (temperate desert). In the Thornthwaite climate classification system , deserts would be classified as arid megathermal climates. Polar deserts are
32832-597: The surrounding semi-arid lands for millennia. Nomads have moved their flocks and herds to wherever grazing is available, and oases have provided opportunities for a more settled way of life. The cultivation of semi-arid regions encourages erosion of soil and is one of the causes of increased desertification. Desert farming is possible with the aid of irrigation , and the Imperial Valley in California provides an example of how previously barren land can be made productive by
33024-399: The tallest type of dune. Rounded mounds of sand without a slip face are the rare dome dunes, found on the upwind edges of sand seas. In deserts where large amounts of limestone mountains surround a closed basin , such as at White Sands National Park in south-central New Mexico , occasional storm runoff transports dissolved limestone and gypsum into a low-lying pan within the basin where
33216-407: The trunk is folded like a concertina , allowing it to expand, and a large specimen can hold eight tons of water after a good downpour. Other xerophytic plants have developed similar strategies by a process known as convergent evolution . They limit water loss by reducing the size and number of stomata, by having waxy coatings and hairy or tiny leaves. Some are deciduous, shedding their leaves in
33408-614: The unprotected surface of the ground to denudation . About one-third of the land surface of the Earth is arid or semi-arid . This includes much of the polar regions , where little precipitation occurs, and which are sometimes called polar deserts or "cold deserts". Deserts can be classified by the amount of precipitation that falls, by the temperature that prevails, by the causes of desertification or by their geographical location. Deserts are formed by weathering processes as large variations in temperature between day and night put strains on
33600-510: The upper canines of coyotes extend past the mental foramina . At the time of the European colonization of the Americas, coyotes were largely confined to open plains and arid regions of the western half of the continent. In early post-Columbian historical records, determining whether the writer is describing coyotes or wolves is often difficult. One record from 1750 in Kaskaskia, Illinois , written by
33792-527: The upper side of the base of the tail and are a bluish-black color. The color and texture of the coyote's fur vary somewhat geographically. The hair's predominant color is light gray and red or fulvous , interspersed around the body with black and white. Coyotes living at high elevations tend to have more black and gray shades than their desert -dwelling counterparts, which are more fulvous or whitish-gray. The coyote's fur consists of short, soft underfur and long, coarse guard hairs. The fur of northern subspecies
33984-496: The use of the word desert can cause confusion. In English before the 20th century, desert was often used in the sense of "unpopulated area", without specific reference to aridity ; but today the word is most often used in its climate-science sense (an area of low precipitation). Phrases such as " desert island " and " Great American Desert ", or Shakespeare 's "deserts of Bohemia " ( The Winter's Tale ) in previous centuries did not necessarily imply sand or aridity; their focus
34176-416: The water evaporates, depositing the gypsum and forming crystals known as selenite . The crystals left behind by this process are eroded by the wind and deposited as vast white dune fields that resemble snow-covered landscapes. These types of dune are rare, and only form in closed arid basins that retain the highly soluble gypsum that would otherwise be washed into the sea. A large part of the surface area of
34368-476: The western edges of continental land masses in regions where cold currents approach the land or cold water upwellings rise from the ocean depths. The cool winds crossing this water pick up little moisture and the coastal regions have low temperatures and very low rainfall, the main precipitation being in the form of fog and dew. The range of temperatures on a daily and annual scale is relatively low, being 11 °C (20 °F) and 5 °C (9 °F) respectively in
34560-631: The wild, as the mating cycles of dogs and coyotes do not coincide, and coyotes are usually antagonistic towards dogs. Hybridization usually only occurs when coyotes are expanding into areas where conspecifics are few, and dogs are the only alternatives. Even then, pup survival rates are lower than normal, as dogs do not form pair bonds with coyotes, thus making the rearing of pups more difficult. In captivity, F 1 hybrids (first generation) tend to be more mischievous and less manageable as pups than dogs, and are less trustworthy on maturity than wolf-dog hybrids . Hybrids vary in appearance, but generally retain
34752-537: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
34944-404: The wind blows, saltation and creep take place on the windward side of the dune and individual grains of sand move uphill. When they reach the crest, they cascade down the far side. The upwind slope typically has a gradient of 10° to 20° while the lee slope is around 32°, the angle at which loose dry sand will slip. As this wind-induced movement of sand grains takes place, the dune moves slowly across
35136-511: The wind direction and gradually move across the surface as the wind continues to blow. The distance between their crests corresponds to the average length of jumps made by particles during saltation. The ripples are ephemeral and a change in wind direction causes them to reorganise. Sand dunes are accumulations of windblown sand piled up in mounds or ridges. They form downwind of copious sources of dry, loose sand and occur when topographic and climatic conditions cause airborne particles to settle. As
35328-497: The wolf is proposed to be due to the coyote retaining more of the mitochondrial genome from the unknown extinct canid. As of 2005, 19 subspecies are recognized. Geographic variation in coyotes is not great, though taken as a whole, the eastern subspecies ( C. l. thamnos and C. l. frustor ) are large, dark-colored animals, with a gradual paling in color and reduction in size westward and northward ( C. l. texensis , C. l. latrans , C. l. lestes , and C. l. incolatus ),
35520-600: The wolves of North America display skull traits more similar to the coyote than wolves from Eurasia. In 2010, a study found that the coyote was a basal member of the clade that included the Tibetan wolf , the domestic dog , the Mongolian wolf and the Eurasian wolf , with the Tibetan wolf diverging early from wolves and domestic dogs. In 2016, a whole-genome DNA study proposed, based on
35712-450: The world's deserts consists of flat, stone-covered plains dominated by wind erosion. In " eolian deflation ", the wind continually removes fine-grained material, which becomes wind-blown sand. This exposes coarser-grained material, mainly pebbles with some larger stones or cobbles , leaving a desert pavement , an area of land overlaid by closely packed smooth stones forming a tessellated mosaic. Different theories exist as to how exactly
35904-470: The world, around 20% of desert is sand, varying from only 2% in North America to 30% in Australia and over 45% in Central Asia. Where sand does occur, it is usually in large quantities in the form of sand sheets or extensive areas of dunes . A sand sheet is a near-level, firm expanse of partially consolidated particles in a layer that varies from a few centimeters to a few meters thick. The structure of
36096-811: Was 40% wolf to 60% coyote ancestry in red wolves, 60% wolf to 40% coyote in Eastern timber wolves, and 75% wolf to 25% coyote in the Great Lakes wolves. There was 10% coyote ancestry in Mexican wolves and the Atlantic Coast wolves, 5% in Pacific Coast and Yellowstone wolves, and less than 3% in Canadian archipelago wolves. If a third canid had been involved in the admixture of the North American wolf-like canids, then its genetic signature would have been found in coyotes and wolves, which it has not. In 2018, whole genome sequencing
36288-482: Was a result of a wolf-coyote admixture, combined with extensive backcrossing with parent gray wolf populations. The red wolf may have originated during a time of declining wolf populations in the Southeastern Woodlands , forcing a wolf-coyote hybridization, as well as backcrossing with local parent coyote populations to the extent that about 75–80% of the modern red wolf's genome is of coyote derivation. Like
36480-592: Was in existence. They proposed that the progression from Eucyon davisi to C. lepophagus to the coyote was linear evolution. C. latrans and C. aureus are closely related to C. edwardii , a species that appeared earliest spanning the mid- Blancan ( late Pliocene ) to the close of the Irvingtonian (late Pleistocene), and coyote remains indistinguishable from C. latrans were contemporaneous with C. edwardii in North America. Johnston describes C. lepophagus as having
36672-459: Was the sparse population. Deserts occupy about one third of Earth's land surface. Bottomlands may be salt -covered flats. Eolian processes are major factors in shaping desert landscapes. Polar deserts (also seen as "cold deserts") have similar features, except the main form of precipitation is snow rather than rain. Antarctica is the world's largest cold desert (composed of about 98% thick continental ice sheet and 2% barren rock). Some of
36864-456: Was used to compare members of the genus Canis . The study indicates that the common ancestor of the coyote and gray wolf has genetically admixed with a ghost population of an extinct, unidentified canid. The "ghost" canid was genetically close to the dhole , and had evolved after the divergence of the African wild dog from the other canid species. The basal position of the coyote compared to
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