186-581: The Chinook salmon / ʃ ɪ ˈ n ʊ k / ( Oncorhynchus tshawytscha ) is the largest and most valuable species of Pacific salmon . Its common name is derived from the Chinookan peoples . Other vernacular names for the species include king salmon , Quinnat salmon , Tsumen , spring salmon , chrome hog , Blackmouth , and Tyee salmon . The scientific species name is based on the Russian common name chavycha (чавыча). Chinook are anadromous fish native to
372-423: A mutation–selection balance . It is predicted that a viral quasispecies at a low but evolutionarily neutral and highly connected (that is, flat) region in the fitness landscape will outcompete a quasispecies located at a higher but narrower fitness peak in which the surrounding mutants are unfit, "the quasispecies effect" or the "survival of the flattest". There is no suggestion that a viral quasispecies resembles
558-400: A "classical" method of determining species, such as with Linnaeus, early in evolutionary theory. However, different phenotypes are not necessarily different species (e.g. a four-winged Drosophila born to a two-winged mother is not a different species). Species named in this manner are called morphospecies . In the 1970s, Robert R. Sokal , Theodore J. Crovello and Peter Sneath proposed
744-424: A 'smallest clade' idea" (a phylogenetic species concept). Mishler and Wilkins and others concur with this approach, even though this would raise difficulties in biological nomenclature. Wilkins cited the ichthyologist Charles Tate Regan 's early 20th century remark that "a species is whatever a suitably qualified biologist chooses to call a species". Wilkins noted that the philosopher Philip Kitcher called this
930-504: A coastline is impossible to measure exactly . Canada borders approximately 5,200 miles (8,400 km) of coastline, while the remaining 5,300 miles (8,500 km) are bordered by the United States. Michigan has the longest shoreline of the United States, bordering roughly 3,288 miles (5,292 km) of lakes, followed by Wisconsin (820 miles (1,320 km)), New York (473 miles (761 km)), and Ohio (312 miles (502 km)). Traversing
1116-428: A connected series of neighbouring populations, each of which can sexually interbreed with adjacent related populations, but for which there exist at least two "end" populations in the series, which are too distantly related to interbreed, though there is a potential gene flow between each "linked" population. Such non-breeding, though genetically connected, "end" populations may co-exist in the same region thus closing
1302-502: A conservation advocacy group, are currently petitioning for an ESA listing on Chinook in several river basins in Washington state, including the Chehalis , Quinault, Queets, Hoh and Quillayute. The petition, filed in 2023, focuses on spring-run Chinook, the region's most vulnerable salmon population. In April 2008, commercial fisheries in both Oregon and California were closed in response to
1488-432: A different species from its ancestors. Viruses have enormous populations, are doubtfully living since they consist of little more than a string of DNA or RNA in a protein coat, and mutate rapidly. All of these factors make conventional species concepts largely inapplicable. A viral quasispecies is a group of genotypes related by similar mutations, competing within a highly mutagenic environment, and hence governed by
1674-508: A genetic boundary suitable for defining a species concept is present. DNA barcoding has been proposed as a way to distinguish species suitable even for non-specialists to use. One of the barcodes is a region of mitochondrial DNA within the gene for cytochrome c oxidase . A database, Barcode of Life Data System , contains DNA barcode sequences from over 190,000 species. However, scientists such as Rob DeSalle have expressed concern that classical taxonomy and DNA barcoding, which they consider
1860-523: A large watershed via many rivers and contain approximately 35,000 islands. There are also several thousand smaller lakes, often called "inland lakes", within the basin. The surface area of the five primary lakes combined is roughly equal to the size of the United Kingdom, while the surface area of the entire basin (the lakes and the land they drain) is about the size of the UK and France combined. Lake Michigan
2046-465: A lineage should be divided into multiple chronospecies , or when populations have diverged to have enough distinct character states to be described as cladistic species. Species and higher taxa were seen from the time of Aristotle until the 18th century as categories that could be arranged in a hierarchy, the great chain of being . In the 19th century, biologists grasped that species could evolve given sufficient time. Charles Darwin 's 1859 book On
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#17328523129122232-492: A misnomer, need to be reconciled, as they delimit species differently. Genetic introgression mediated by endosymbionts and other vectors can further make barcodes ineffective in the identification of species. A phylogenetic or cladistic species is "the smallest aggregation of populations (sexual) or lineages (asexual) diagnosable by a unique combination of character states in comparable individuals (semaphoronts)". The empirical basis – observed character states – provides
2418-405: A national basis. The total surface area of the lakes is approximately 94,250 square miles (244,100 km )—nearly the same size as the United Kingdom, and larger than the U.S. states of New York , New Jersey , Connecticut , Rhode Island , Massachusetts , Vermont , and New Hampshire combined. The Great Lakes coast measures approximately 10,500 miles (16,900 km), but the length of
2604-456: A nuisance to industries by clogging pipes. The U.S. Fish and Wildlife Service estimated in 2007 that the economic impact of the zebra mussel could be about $ 5 billion over the next decade. The alewife first entered the system west of Lake Ontario via 19th-century canals. By the 1960s, the small silver fish had become a familiar nuisance to beach goers across Lakes Michigan, Huron, and Erie. Periodic mass die-offs result in vast numbers of
2790-449: A particular species, including which genus (and higher taxa) it is placed in, is a hypothesis about the evolutionary relationships and distinguishability of that group of organisms. As further information comes to hand, the hypothesis may be corroborated or refuted. Sometimes, especially in the past when communication was more difficult, taxonomists working in isolation have given two distinct names to individual organisms later identified as
2976-400: A perfect model of life, it is still a useful tool to scientists and conservationists for studying life on Earth, regardless of the theoretical difficulties. If species were fixed and clearly distinct from one another, there would be no problem, but evolutionary processes cause species to change. This obliges taxonomists to decide, for example, when enough change has occurred to declare that
3162-681: A possible reason for a decrease in population of various whitefish , important because of their culinary desirability and, hence, economic consequence. Moreover, between 1879 and 1899, reported whitefish harvests declined from some 24.3 million pounds (11 million kg) to just over 9 million pounds (4 million kg). By 1900, commercial fishermen on Lake Michigan were hauling in an average of 41 million pounds of fish annually. By 1938, Wisconsin's commercial fishing operations were motorized and mechanized, generating jobs for more than 2,000 workers, and hauling 14 million pounds per year. The population of giant freshwater mussels
3348-425: A pronounced curve or hook called a " kype ." Studies have shown that larger and more dominant male salmon have a reproductive advantage as female Chinook are often more aggressive toward smaller males. Chinook spawn in larger and deeper waters than other salmon species and can be found on the spawning redds (nests) from September to December. The female salmon may lay her eggs in four to five nesting pockets within
3534-730: A redd. After laying eggs, females guard the redd from four to 25 days before dying, while males seek additional mates. Chinook eggs hatch 90 to 150 days after deposition, depending upon water temperature. Egg deposits are timed to ensure the young salmon fry emerge during an appropriate season for survival and growth. Fry and parr (young fish) usually stay in fresh water for 12 to 18 months before traveling downstream to estuaries , where they remain as smolts for several months. Some Chinook return to fresh water one or two years earlier than their counterparts and are referred to as "jack" salmon. "Jack" salmon are typically less than 60 cm (24 in) long but are sexually mature. The Yukon River has
3720-560: A relatively small area of southeast Lake Michigan where savanna or prairie occasionally intruded). Logging, urbanization, and agriculture uses have changed that relationship. In the early 21st century, Lake Superior's shores are 91% forested, Lake Huron 68%, Lake Ontario 49%, Lake Michigan 41%, and Lake Erie, where logging and urbanization has been most extensive, 21%. Some of these forests are second or third growth (i.e. they have been logged before, changing their composition). At least 13 wildlife species are documented as becoming extinct since
3906-620: A result, progress is slowly being made to help restore the salmon habitats along the river. Many tribes, including the Hoopa Valley Tribe, Karuk Tribe , Yurok Tribe , and the Shasta Indian Nation, voiced support for the removal of dams on the Klamath River. Representatives of the Shasta Indian Nation claimed that the construction of Copco No 1 Dam caused the submerging of sites significant to them, including burial grounds. Un-Dam
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#17328523129124092-483: A season, and can survive poor water quality conditions. The influx of parasitic lamprey populations after the development of the Erie Canal and the much later Welland Canal led to the two federal governments of the United States and Canada working on joint proposals to control it. By the mid-1950s, the lake trout populations of Lakes Michigan and Huron were reduced, with the lamprey deemed largely to blame. This led to
4278-707: A second fault line, the Saint Lawrence rift , formed approximately 570 million years ago, the basis for Lakes Ontario and Erie was created, along with what would become the Saint Lawrence River. The Great Lakes are estimated to have been formed at the end of the Last Glacial Period (the Wisconsin glaciation ended 10,000 to 12,000 years ago), when the Laurentide Ice Sheet receded. The retreat of
4464-525: A short period until they are strong enough to journey out to the ocean and acquire more food. Chinook juveniles divide into two types: ocean-type and stream-type. Ocean-type Chinook migrate to salt water in their first year. Stream-type salmon spend one full year in fresh water before migrating to the ocean. After a few years in the ocean, adult salmon, then large enough to escape most predators, return to their natal streambeds to mate. Chinook can have extended lifespans, in which some fish spend one to five years in
4650-400: A short way of saying that something applies to many species within a genus, but not to all. If scientists mean that something applies to all species within a genus, they use the genus name without the specific name or epithet. The names of genera and species are usually printed in italics . However, abbreviations such as "sp." should not be italicised. When a species' identity is not clear,
4836-632: A similar legend that has illustrated the importance of not messing up the Chinook salmon migration. The legend described three Skookums which can be related to the three dams on the Klamath river in California. It has been known that the creation of dams has negatively impacted the lives of many Native American Indians by disrupting their food supply and the flow of water. The impact on the salmon migration has been seen by not only tribal members but others as well, and as
5022-530: A single stocking of juveniles in the lower river around 1930. Sporadic efforts to introduce the fish to New Zealand waters in the late 19th century were largely failures and led to no evident establishments. Initially ova were imported from the Baird hatchery of the McCloud River in California. Further efforts in the early 20th century were more successful and subsequently led to the establishment of spawning runs in
5208-673: A small percid fish from Eurasia, became the most abundant fish species in Lake Superior's Saint Louis River within five years of its detection in 1986. Its range, which has expanded to Lake Huron, poses a significant threat to the lower lake fishery. Five years after first being observed in the St. Clair River, the round goby can now be found in all of the Great Lakes. The goby is considered undesirable for several reasons: it preys upon bottom-feeding fish, overruns optimal habitat, spawns multiple times
5394-404: A specialist may use "cf." before the epithet to indicate that confirmation is required. The abbreviations "nr." (near) or "aff." (affine) may be used when the identity is unclear but when the species appears to be similar to the species mentioned after. With the rise of online databases, codes have been devised to provide identifiers for species that are already defined, including: The naming of
5580-523: A species as groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups. It has been argued that this definition is a natural consequence of the effect of sexual reproduction on the dynamics of natural selection. Mayr's use of the adjective "potentially" has been a point of debate; some interpretations exclude unusual or artificial matings that occur only in captivity, or that involve animals capable of mating but that do not normally do so in
5766-400: A species as determined by a taxonomist. A typological species is a group of organisms in which individuals conform to certain fixed properties (a type), so that even pre-literate people often recognise the same taxon as do modern taxonomists. The clusters of variations or phenotypes within specimens (such as longer or shorter tails) would differentiate the species. This method was used as
Chinook salmon - Misplaced Pages Continue
5952-488: A species. All species definitions assume that an organism acquires its genes from one or two parents very like the "daughter" organism, but that is not what happens in HGT. There is strong evidence of HGT between very dissimilar groups of prokaryotes , and at least occasionally between dissimilar groups of eukaryotes , including some crustaceans and echinoderms . The evolutionary biologist James Mallet concludes that there
6138-685: A species. Generally the term includes the unknown element of a distinct act of creation. Many authors have argued that a simple textbook definition, following Mayr's concept, works well for most multi-celled organisms , but breaks down in several situations: Species identification is made difficult by discordance between molecular and morphological investigations; these can be categorised as two types: (i) one morphology, multiple lineages (e.g. morphological convergence , cryptic species ) and (ii) one lineage, multiple morphologies (e.g. phenotypic plasticity , multiple life-cycle stages). In addition, horizontal gene transfer (HGT) makes it difficult to define
6324-503: A sustainable manner. The planning and approval process for new salmon farms in New Zealand considers the farm's potential environmental effects, its effects on fishing activities (if it is a marine farm), and any possible cultural and social effects. In the interest of fish welfare, a number of New Zealand salmon farming operations anaesthetise salmon before slaughter using Aqui-S™, an organically based anaesthetic developed in New Zealand that
6510-519: A taxonomic decision at the discretion of cognizant specialists, is not governed by the Codes of Zoological or Botanical Nomenclature, in contrast to the PhyloCode , and contrary to what is done in several other fields, in which the definitions of technical terms, like geochronological units and geopolitical entities, are explicitly delimited. The nomenclatural codes that guide the naming of species, including
6696-506: A traditional biological species. The International Committee on Taxonomy of Viruses has since 1962 developed a universal taxonomic scheme for viruses; this has stabilised viral taxonomy. Most modern textbooks make use of Ernst Mayr 's 1942 definition, known as the Biological Species Concept as a basis for further discussion on the definition of species. It is also called a reproductive or isolation concept. This defines
6882-487: A trip to catch Chinook salmon, $ 90 for lake trout , and $ 180 for walleye . Should the Chinook salmon fishery collapse and be replaced with a native lake trout fishery, the economic value would decrease by 80%. Since the later 1970s, the size and age range of Chinook salmon have been declining according to studies along the northwest Pacific coast from Alaska to California for the years of 1977 to 2015 which examined about 1.5 million Chinook salmon. Ocean-5 Chinook (which means
7068-460: A value of NZ$ 61 million in export earnings. For the year ended March 2011, this amount had increased to NZ$ 85 million. New Zealand accounts for about half of the global production of Chinook salmon, and about half of New Zealand's production is exported. Japan is New Zealand's largest export market, with stock also being supplied to other countries of the Pacific Rim , including Australia. Farming of
7254-447: A variation on the morphological species concept, a phenetic species, defined as a set of organisms with a similar phenotype to each other, but a different phenotype from other sets of organisms. It differs from the morphological species concept in including a numerical measure of distance or similarity to cluster entities based on multivariate comparisons of a reasonably large number of phenotypic traits. A mate-recognition species
7440-515: A variety of reasons. Viruses are a special case, driven by a balance of mutation and selection , and can be treated as quasispecies . Biologists and taxonomists have made many attempts to define species, beginning from morphology and moving towards genetics . Early taxonomists such as Linnaeus had no option but to describe what they saw: this was later formalised as the typological or morphological species concept. Ernst Mayr emphasised reproductive isolation, but this, like other species concepts,
7626-438: Is "an entity composed of organisms which maintains its identity from other such entities through time and over space, and which has its own independent evolutionary fate and historical tendencies". This differs from the biological species concept in embodying persistence over time. Wiley and Mayden stated that they see the evolutionary species concept as "identical" to Willi Hennig 's species-as-lineages concept, and asserted that
Chinook salmon - Misplaced Pages Continue
7812-414: Is "new" water, originating from rivers, precipitation, and groundwater springs. In the post-glacial period, evaporation , and drainage have generally been balanced, making the levels of the lakes relatively constant. Intensive human population growth began in the region in the 20th century and continues today. At least two human water use activities have been identified as having the potential to affect
7998-400: Is a group of sexually reproducing organisms that recognise one another as potential mates. Expanding on this to allow for post-mating isolation, a cohesion species is the most inclusive population of individuals having the potential for phenotypic cohesion through intrinsic cohesion mechanisms; no matter whether populations can hybridise successfully, they are still distinct cohesion species if
8184-452: Is a population of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring , typically by sexual reproduction . It is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity . Other ways of defining species include their karyotype , DNA sequence, morphology , behaviour, or ecological niche . In addition, paleontologists use
8370-462: Is a prized and sought-after catch for a sporting angler. The flesh of the salmon is also highly valued for its dietary nutritional content, which includes high levels of important omega-3 fatty acids . Some populations are endangered; however, many are healthy. The Chinook salmon has not been assessed for the IUCN Red List . According to NOAA , the Chinook salmon population along the California coast
8556-458: Is a set of organisms adapted to a particular set of resources, called a niche, in the environment. According to this concept, populations form the discrete phenetic clusters that we recognise as species because the ecological and evolutionary processes controlling how resources are divided up tend to produce those clusters. A genetic species as defined by Robert Baker and Robert Bradley is a set of genetically isolated interbreeding populations. This
8742-873: Is also conducting efforts to revegetate certain areas in the watershed with trees and native grasses. The Oregon Department of Fish and Wildlife (ODFW), in cooperation with members of the Klamath and Yurok tribes, also finalized a two-phase plan to re-establish wild Chinook and other anadromous fishes in the Upper Klamath Basin. As of December 2022, the plan is in its monitoring phase, in which ODFW are studying 10,000 hatchery-born spring-run Chinook released in certain tributaries of Upper Klamath Lake . Introduced Chinook salmon in Lake Michigan are sought after by tourists enjoying chartered fishing trips . A 2016 survey of Wisconsin anglers found they would, on average, pay $ 140 for
8928-414: Is called speciation . Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book The Origin of Species . Speciation depends on a measure of reproductive isolation , a reduced gene flow. This occurs most easily in allopatric speciation, where populations are separated geographically and can diverge gradually as mutations accumulate. Reproductive isolation
9114-412: Is called the specific name or the specific epithet (in botanical nomenclature , also sometimes in zoological nomenclature ). For example, Boa constrictor is one of the species of the genus Boa , with constrictor being the species' epithet. While the definitions given above may seem adequate at first glance, when looked at more closely they represent problematic species concepts. For example,
9300-655: Is compromised, affected stock can decline. One Seattle Times article states, "Pacific salmon have disappeared from 40 percent of their historic range outside Alaska," and concludes it is imperative for people to realize the needs of salmon and try not to contribute to destructive practices that harm salmon runs. In the Pacific Northwest, the summer runs of especially large Chinook once common (before dams and overfishing led to declines) were known as June hogs . A Chinook's birthplace and later evolution can be tracked by looking at its otolith (ear) bone. The bone can record
9486-572: Is declining from factors such as overfishing, loss of freshwater and estuarine habitat, hydropower development, poor ocean conditions, and hatchery practices. Historically, the native distribution of Chinook salmon in North America ranged from the Ventura River in California in the south to Kotzebue Sound in Alaska in the north. Recent studies have shown that Chinook salmon are historically native to
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#17328523129129672-403: Is described formally, in a publication that assigns it a unique scientific name. The description typically provides means for identifying the new species, which may not be based solely on morphology (see cryptic species ), differentiating it from other previously described and related or confusable species and provides a validly published name (in botany) or an available name (in zoology) when
9858-485: Is essential for algae and other water-filtering agents not to be destroyed in the oceans because they contribute to the well-being of the food chain . With some populations endangered, precautions are necessary to prevent overfishing and habitat destruction , including appropriate management of hydroelectric and irrigation projects. If too few fish remain because of fishing and land management practices, salmon have more difficulty reproducing. When one of these factors
10044-621: Is flat according to the International Joint Commission. Reports about this issue on the U.S. side highlight five large municipal systems (those of Detroit, Cleveland, Buffalo, Milwaukee and Gary ) as being the largest current periodic sources of untreated discharges into the Great Lakes. The fish of the Great Lakes have anti-depressant drugs meant for humans in their brains, which has caused concerns. The number of American adults who take anti-depressant drugs rose from 7.7% of all American adults in 1999–2002 to 12.7% in 2011–2014. As
10230-671: Is further weakened by the existence of microspecies , groups of organisms, including many plants, with very little genetic variability, usually forming species aggregates . For example, the dandelion Taraxacum officinale and the blackberry Rubus fruticosus are aggregates with many microspecies—perhaps 400 in the case of the blackberry and over 200 in the dandelion, complicated by hybridisation , apomixis and polyploidy , making gene flow between populations difficult to determine, and their taxonomy debatable. Species complexes occur in insects such as Heliconius butterflies, vertebrates such as Hypsiboas treefrogs, and fungi such as
10416-726: Is hard or even impossible to test. Later biologists have tried to refine Mayr's definition with the recognition and cohesion concepts, among others. Many of the concepts are quite similar or overlap, so they are not easy to count: the biologist R. L. Mayden recorded about 24 concepts, and the philosopher of science John Wilkins counted 26. Wilkins further grouped the species concepts into seven basic kinds of concepts: (1) agamospecies for asexual organisms (2) biospecies for reproductively isolated sexual organisms (3) ecospecies based on ecological niches (4) evolutionary species based on lineage (5) genetic species based on gene pool (6) morphospecies based on form or phenotype and (7) taxonomic species,
10602-546: Is heavy. Pleasure boats can enter or exit the Great Lakes by way of the Erie Canal and Hudson River in New York. The Erie Canal connects to the Great Lakes at the east end of Lake Erie (at Buffalo, New York ) and at the south side of Lake Ontario (at Oswego, New York ). The lakes were originally fed by both precipitation and meltwater from glaciers which are no longer present. In modern times, only about 1% of volume per year
10788-554: Is in the Middle Fork and Upper Salmon River in Idaho. These fish travel over 2,100 m (7,000 ft) in elevation, and over 1,400 km (900 mi), in their migration through eight dams and reservoirs on the Columbia and Lower Snake Rivers. Chinook eat amphipods and other crustaceans and insects while young, and primarily other fish when older. Young salmon feed in streambeds for
10974-560: Is known locally in New Zealand) have established sizeable pelagic runs. The Chinook is blue-green, red, or purple on the back and on the top of the head, with silvery sides and white ventral surfaces. It has black spots on its tail and the upper half of its body. Although spots are seen on the tail in pink salmon and silver on the tail in coho and chum salmon , Chinook are unique among the Pacific salmon in combining black spots and silver on
11160-424: Is measured, by surface area, either Lake Superior or Lake Michigan–Huron is the second-largest lake in the world and the largest freshwater lake. Lake Michigan is the largest lake, by surface area, that is entirely within one country, the United States. The Great Lakes began to form at the end of the Last Glacial Period around 14,000 years ago, as retreating ice sheets exposed the basins they had carved into
11346-403: Is no easy way to tell whether related geographic or temporal forms belong to the same or different species. Species gaps can be verified only locally and at a point of time. One is forced to admit that Darwin's insight is correct: any local reality or integrity of species is greatly reduced over large geographic ranges and time periods. The botanist Brent Mishler argued that the species concept
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#173285231291211532-468: Is not valid, notably because gene flux decreases gradually rather than in discrete steps, which hampers objective delimitation of species. Indeed, complex and unstable patterns of gene flux have been observed in cichlid teleosts of the East African Great Lakes . Wilkins argued that "if we were being true to evolution and the consequent phylogenetic approach to taxa, we should replace it with
11718-761: Is safe for use in food and that has been favourably reported on by the British Humane Slaughter Association . In recognition of the sustainable, environmentally conscious practices, the New Zealand salmon farming industry has been acknowledged as the world's greenest by the Global Aquaculture Performance Index. Chile is the only country other than New Zealand currently producing significant quantities of farmed Chinook salmon. The United States has not produced farmed Chinook in commercial quantities since 1994. In Canada, most commercial Chinook salmon farming ceased by 2009. Fisheries in
11904-586: Is similar to Mayr's Biological Species Concept, but stresses genetic rather than reproductive isolation. In the 21st century, a genetic species could be established by comparing DNA sequences. Earlier, other methods were available, such as comparing karyotypes (sets of chromosomes ) and allozymes ( enzyme variants). An evolutionarily significant unit (ESU) or "wildlife species" is a population of organisms considered distinct for purposes of conservation. In palaeontology , with only comparative anatomy (morphology) and histology from fossils as evidence,
12090-525: Is similar to how other tribes relied more on buffalo for food, and have many legends and spiritual ties to them. Chinook salmon were described and enthusiastically eaten by the Lewis and Clark Expedition . Lewis wrote that, when fresh, they tasted better than any other fish he had ever eaten. They did not particularly like dried or "pounded" salmon. Lewis and Clark knew about Pacific salmon but had never seen one. The Western world had known about Pacific salmon since
12276-433: Is sometimes an important source of genetic variation. Viruses can transfer genes between species. Bacteria can exchange plasmids with bacteria of other species, including some apparently distantly related ones in different phylogenetic domains , making analysis of their relationships difficult, and weakening the concept of a bacterial species. Great Lakes The Great Lakes ( French : Grands Lacs ), also called
12462-510: Is the only one of the Great Lakes that is entirely within the United States; the others form a water boundary between the United States and Canada. The lakes are divided among the jurisdictions of the Canadian province of Ontario and the U.S. states of Michigan , Wisconsin , Minnesota , Illinois , Indiana , Ohio , Pennsylvania , and New York . Both the province of Ontario and the state of Michigan include in their boundaries portions of four of
12648-594: Is threatened by hybridisation, but this can be selected against once a pair of populations have incompatible alleles of the same gene, as described in the Bateson–Dobzhansky–Muller model . A different mechanism, phyletic speciation, involves one lineage gradually changing over time into a new and distinct form (a chronospecies ), without increasing the number of resultant species. Horizontal gene transfer between organisms of different species, either through hybridisation , antigenic shift , or reassortment ,
12834-572: Is vitally important for the fish to reach the oceans to grow into healthy adult fish to sustain the species without being impeded by man-made structures such as dams. The bodies of water for salmon habitat must be clean and oxygenated. One sign of high productivity and growth rate in the oceans is the level of algae . Increased algal levels lead to higher levels of carbon dioxide in the water, which transfers into living organisms, fostering underwater plants and small organisms, which salmon eat. Algae can filter high levels of toxins and pollutants. Thus, it
13020-514: The California drought made them even more vulnerable. A study was done specifically on the California Delta over three years, and it was discovered that the Chinook salmon had a low survival rate for different reasons, and as a result, the Chinook salmon population here has been on a decline. Some of the factors affecting the populations include the route used during migration, drought conditions,
13206-944: The Chicago Sanitary and Ship Canal to the Gulf of Mexico via the Illinois River (from the Chicago River ) and the Mississippi River. An alternate track is via the Illinois River (from Chicago), to the Mississippi, up the Ohio, and then through the Tennessee–Tombigbee Waterway (a combination of a series of rivers and lakes and canals), to Mobile Bay and the Gulf of Mexico. Commercial tug -and- barge traffic on these waterways
13392-832: The Door Peninsula , the Peninsulas of Michigan , and the Ontario Peninsula . Some of these peninsulas even contain smaller peninsulas, such as the Bruce , Keweenaw , Leelanau , Niagara , and Thumb peninsulas. Population centers on the peninsulas include Grand Rapids , Flint , and Detroit in Michigan along with London , Hamilton , Brantford , and Toronto in Ontario. Although the Saint Lawrence Seaway and Great Lakes Waterway make
13578-516: The Environmental Protection Agency have shown that when the mercury comes in contact with many of the bacteria and compounds in the fresh water, it forms the compound methyl mercury , which has a much greater impact on human health than elemental mercury due to a higher propensity for absorption. This form of mercury is not detrimental to a majority of fish types, but is very detrimental to people and other wildlife animals who consume
13764-669: The Finger Lakes region of New York, as well as in Prince Edward County, Ontario , on Lake Ontario's northeast shore. The Great Lakes have been observed to help intensify storms, such as Hurricane Hazel in 1954, and the 2011 Goderich, Ontario tornado , which moved onshore as a tornadic waterspout . In 1996, a rare tropical or subtropical storm was observed forming in Lake Huron, dubbed the 1996 Lake Huron cyclone . Rather large severe thunderstorms covering wide areas are well known in
13950-516: The Great Lakes of North America , are a series of large interconnected freshwater lakes spanning the Canada–United States border . The five lakes are Superior , Michigan , Huron , Erie , and Ontario . (Hydrologically, Michigan and Huron are a single body of water, as they are joined by Straits of Mackinac .) The Great Lakes Waterway enables modern travel and shipping by water among
14136-631: The Guadalupe River watershed in California, the southernmost major metropolitan area hosting salmon runs in the United States. Populations have disappeared from large areas where they once flourished, however, or shrunk by as much as 40 percent. In some regions, their inland range has been cut off, mainly by dams and habitat alterations: in Southern California , in some areas east of the Coast Ranges of California and Oregon , and in large areas in
14322-523: The ICZN for animals and the ICN for plants, do not make rules for defining the boundaries of the species. Research can change the boundaries, also known as circumscription, based on new evidence. Species may then need to be distinguished by the boundary definitions used, and in such cases the names may be qualified with sensu stricto ("in the narrow sense") to denote usage in the exact meaning given by an author such as
14508-489: The Klamath River , four dams on the Klamath ( Iron Gate Dam , Copco No 2 Dam , Copco No 1 Dam , and John C. Boyle Dam ) will be removed by the end of 2024. Klamath River Renewal Corporation (KRRC), who is in charge of the operation, cited the adverse effects of the dams on fish survivability and the blockage of fish habitat as major reasons for removal. In June 2023, Copco 2 Dam was torn down. In addition to dam removal, KRRC
14694-520: The Klamath tribes and the Yurok Tribe , have lived along the Klamath river , and the Chinook salmon have been an important part of their lives, spiritually and physically. An Indian legend of a tribe on the Klamath river describes how the construction of the dam has hurt the fish population and that the impact on them has gone unnoticed, and the destruction of the dam is what has brought back their food supply and made them happy again. The Klamath tribe had
14880-513: The Niagara Escarpment precludes all natural navigation, the four upper lakes are commonly called the "upper great lakes". This designation is not universal. Those living on the shore of Lake Superior often refer to all the other lakes as "the lower lakes", because they are farther south. Sailors of bulk freighters transferring cargoes from Lake Superior and northern Lake Michigan and Lake Huron to ports on Lake Erie or Ontario commonly refer to
15066-645: The North Pacific Ocean and the river systems of western North America, ranging from California to Alaska , as well as Asian rivers ranging from northern Japan to the Palyavaam River in Arctic northeast Siberia. They have been introduced to other parts of the world, including New Zealand and Patagonia . Introduced Chinook salmon are thriving in Lake Michigan and Michigan 's western rivers. A large Chinook
15252-635: The Snake River and upper Columbia River drainage basins . In certain areas such as California's Sacramento–San Joaquin River Delta , it was revealed that extremely low numbers of juvenile Chinook salmon (less than 1%) were surviving. In the western Pacific, the distribution ranges from northern Japan ( Hokkaido ) in the south to the Arctic Ocean as far as the East Siberian Sea and Palyavaam River in
15438-516: The alewife , an invasive species of nuisance fish from the Atlantic Ocean. In the 1960s, alewives constituted 90% of the biota in these lakes. Coho salmon had been introduced the year before, and the program was successful. Chinook and Coho salmon thrived on the alewives and spawned in the lakes' tributaries. After this success, Chinook were introduced into the other Great Lakes , where sport fishermen prize them for their aggressive behaviour on
15624-497: The biosphere . Since the 19th century, an estimated 160 new species have found their way into the Great Lakes ecosystem; many have become invasive; the overseas ship ballast and ship hull parasitism are causing severe economic and ecological impacts. According to the Inland Seas Education Association, on average a new species enters the Great Lakes every eight months. Introductions into the Great Lakes include
15810-399: The fly agaric . Natural hybridisation presents a challenge to the concept of a reproductively isolated species, as fertile hybrids permit gene flow between two populations. For example, the carrion crow Corvus corone and the hooded crow Corvus cornix appear and are classified as separate species, yet they can hybridise where their geographical ranges overlap. A ring species is
15996-498: The jaguar ( Panthera onca ) of Latin America or the leopard ( Panthera pardus ) of Africa and Asia. In contrast, the scientific names of species are chosen to be unique and universal (except for some inter-code homonyms ); they are in two parts used together : the genus as in Puma , and the specific epithet as in concolor . A species is given a taxonomic name when a type specimen
16182-414: The zebra mussel , which was first discovered in 1988, and quagga mussel in 1989. Since 2000, the invasive quagga mussel has smothered the bottom of Lake Michigan almost from shore to shore, and their numbers are estimated at 900 trillion. The mollusks are efficient filter feeders, competing with native mussels and reducing available food and spawning grounds for fish. In addition, the mussels may be
16368-579: The " snowbelts " receive regular snow fall from this localized weather pattern, especially along the eastern shores of the lakes. Snowbelts are found in Wisconsin, Michigan, Ohio, Pennsylvania, New York, and Ontario. Related to the lake effect is the regular occurrence of fog, particularly along the shorelines of the lakes. This is most noticeable along Lake Superior's shores. The lakes tend to moderate seasonal temperatures to some degree but not with as large an influence as do large oceans; they absorb heat and cool
16554-406: The "cynical species concept", and arguing that far from being cynical, it usefully leads to an empirical taxonomy for any given group, based on taxonomists' experience. Other biologists have gone further and argued that we should abandon species entirely, and refer to the "Least Inclusive Taxonomic Units" (LITUs), a view that would be coherent with current evolutionary theory. The species concept
16740-401: The 2008 estimate was 66,000 fish. Scientists from universities and federal, state, and tribal agencies concluded the 2004 and 2005 broods were harmed by poor ocean conditions in 2005 and 2006, in addition to "a long-term, steady degradation of the freshwater and estuarine environment." Such conditions included weak upwelling, warm sea surface temperatures , and low densities of food. In Oregon,
16926-678: The 2010 spring Chinook run was forecast to increase by up to 150% over 2009 populations, growing from 200,000 to over 500,000, making this the largest run in recorded history. Lower temperatures in 2008 North Pacific waters brought in fatter plankton, which, along with greater outflows of Columbia River water, fed the resurgent populations. The Oregon Department of Fish and Wildlife estimated 80% were hatchery-born. Chinook runs in other habitats have not recovered proportionately. In April 2016, Coleman National Fish Hatchery outside of Red Bluff, California , released 12 million juvenile Chinook salmon, with many salmon being tagged for monitoring. The release
17112-560: The Canadian side, Toronto , Hamilton and Mississauga . Though the five lakes lie in separate basins, they form a single, naturally interconnected body of fresh water, within the Great Lakes Basin . As a chain of lakes and rivers, they connect the east-central interior of North America to the Atlantic Ocean. From the interior to the outlet at the Saint Lawrence River, water flows from Superior to Huron and Michigan, southward to Erie, and finally northward to Lake Ontario. The lakes drain
17298-489: The Chicago Sanitary and Ship Canal in order to keep several species of invasive Asian carp out of the lakes. These fast-growing planktivorous fish have heavily colonized the Mississippi and Illinois river systems. Invasive species, particularly zebra and quagga mussels, may be at least partially responsible for the collapse of the deepwater demersal fish community in Lake Huron, as well as drastic unprecedented changes in
17484-647: The Chinook are in highly impacts their survival rates, as some, like the Chinook in the Fraser River, only have a 30% survival rate. More studies and actions are needed for there to be an impact on the survival rates of the Chinook. Due to many of these reasons, the National Wildlife Federation has listed Chinook populations as endangered or threatened. The Chinook salmon is spiritually and culturally prized among certain First Nations peoples. For tribes on
17670-458: The Great Lakes accessible to ocean-going vessels, shifts in shipping to wider ocean-going container ships —which do not fit through the locks on these routes—have limited container shipping on the lakes. Most Great Lakes trade is of bulk material, and bulk freighters of Seawaymax -size or less can move throughout the entire lakes and out to the Atlantic. Larger ships are confined to working within
17856-459: The Great Lakes during mid-summer; these Mesoscale convective complexes or MCCs can cause damage to wide swaths of forest and shatter glass in city buildings. These storms mainly occur during the night, and the systems sometimes have small embedded tornadoes, but more often straight-line winds accompanied by intense lightning. Historically, the Great Lakes, in addition to their lake ecology , were surrounded by various forest ecoregions (except in
18042-441: The Great Lakes each year. Plastics in the water break up into very small particles known as microplastics . Microplastics can also come from synthetic clothing washed down our drains. Plastic waste found in the lakes include single-use plastics , plastics used in packaging, takeout containers as well as pre-production pellets produced by plastics industry . High concentrations of microplastics were discovered in 100 percent of
18228-438: The Great Lakes region include: Plant lists include: Logging of the extensive forests in the Great Lakes region removed riparian and adjacent tree cover over rivers and streams, which provide shade, moderating water temperatures in fish spawning grounds. Removal of trees also destabilized the soil, with greater volumes washed into stream beds causing siltation of gravel beds, and more frequent flooding. Running cut logs down
18414-550: The Great Lakes. A number of self-operating floating devices called Seabin, were put in the Great Lakes to capture plastic trash as part of the Great Lakes Plastic Cleanup project. The project captured 74,000 pieces of trash using this technology between 2020 and 2021; however, it does not claim to catch up with 22 million pounds (10.0 kt) of plastic that ends up in Great Lakes every year. The production, consumption, and throwing away of plastics seems to remain
18600-552: The Klamath , supported by the Karuk and Yurok Tribes, is one of the major campaigns advocating for the Klamath dam removals, which started deconstruction in 2023. Other tribes, including the Nuxalk , Kwakiutl , and Kyuquot , relied primarily on Chinook to eat. Known as the "king salmon" in Alaska for its large size and flavorful flesh, the Chinook is the state fish of this state, and of Oregon. Species A species ( pl. : species)
18786-457: The Northwest coast, salmon were an important part of their culture for spiritual reasons and food. Many celebrate the first spring Chinook caught each year with "first-salmon ceremonies." While salmon fishing in general remains important economically for many tribal communities, it is especially the Chinook harvest that is typically the most valuable. The relation to salmon for the tribes in this area
18972-622: The Origin of Species explained how species could arise by natural selection . That understanding was greatly extended in the 20th century through genetics and population ecology . Genetic variability arises from mutations and recombination , while organisms themselves are mobile, leading to geographical isolation and genetic drift with varying selection pressures . Genes can sometimes be exchanged between species by horizontal gene transfer ; new species can arise rapidly through hybridisation and polyploidy ; and species may become extinct for
19158-500: The Straits of Mackinac. The straits are five miles (8 km) wide and 120 feet (37 m) deep; the water levels rise and fall together, and the flow between Michigan and Huron frequently reverses direction. Dispersed throughout the Great Lakes are approximately 35,000 islands . The largest among them is Manitoulin Island in Lake Huron, the largest island in any inland body of water in
19344-427: The U.S. A variety of steps taken to process industrial and municipal pollution discharges into the system greatly improved water quality by the 1980s, and Lake Erie in particular is significantly cleaner. Discharge of toxic substances has been sharply reduced. Federal and state regulations control substances like PCBs . The first of 43 " Great Lakes Areas of Concern " to be formally "de-listed" through successful cleanup
19530-564: The U.S. and Canada are limited by impacts to weak and endangered salmon runs. Nine populations of Chinook salmon are listed under the U.S. Endangered Species Act (ESA) as either threatened or endangered. In the Snake River , Spring/Summer Chinook and Fall Chinook are ESA listed as Threatened. The fall and late-fall runs in the Central Valley population in California is a U.S. National Marine Fisheries Service (NMFS) species of concern . The Center for Biological Diversity and Pacific Rivers,
19716-405: The abbreviation "sp." in the singular or "spp." (standing for species pluralis , Latin for "multiple species") in the plural in place of the specific name or epithet (e.g. Canis sp.). This commonly occurs when authors are confident that some individuals belong to a particular genus but are not sure to which exact species they belong, as is common in paleontology . Authors may also use "spp." as
19902-495: The air in summer, then slowly radiate that heat in autumn. They protect against frost during transitional weather and keep the summertime temperatures cooler than further inland. This effect can be very localized and overridden by offshore wind patterns. This temperature buffering produces areas known as " fruit belts ", where fruit can be produced that is typically grown much farther south. For instance, western Michigan has apple orchards, and cherry orchards are cultivated adjacent to
20088-570: The amount of hybridisation is insufficient to completely mix their respective gene pools . A further development of the recognition concept is provided by the biosemiotic concept of species. In microbiology , genes can move freely even between distantly related bacteria, possibly extending to the whole bacterial domain. As a rule of thumb, microbiologists have assumed that members of Bacteria or Archaea with 16S ribosomal RNA gene sequences more similar than 97% to each other need to be checked by DNA–DNA hybridisation to decide if they belong to
20274-447: The amount of snowmelt, and infrastructure that affects the flow of water (such as dams and levees). Each of these factors has significantly impacted Chinook survival rates, as most have made it more challenging for Chinook to travel from their spawning grounds to the ocean and back. The fluctuation of water depth as well as temperature have made this more challenging, and as a result, Chinook populations are declining. Which rivers or streams
20460-459: The anti-depressant drugs pass out of human bodies and through sanitation systems into the Great Lakes, this has resulted in fish in the Great Lakes with twenty times the level of anti-depressants in their brains than what is in the water, leading to the fish being exceedingly happy and hence less risk-averse, to the extent of damaging the fish populations. Researchers have found that more than 22 million pounds (10.0 kt) of plastic end up in
20646-432: The arrival of Europeans, and many more are threatened or endangered. Meanwhile, exotic and invasive species have also been introduced. While the organisms living on the bottom of shallow waters are similar to those found in smaller lakes, the deep waters contain organisms found only in deep, cold lakes of the northern latitudes. These include the delicate opossum shrimp (order mysida ), the deepwater scud (a crustacean of
20832-574: The baby salmon are typically grown to the smolt stage (around six months of age) before they are transferred to the sea cages or ponds. Most sea cage farming occurs in the Marlborough Sounds , Stewart Island, and Akaroa Harbour, while freshwater operations in Canterbury , Otago , and Tasman use ponds, raceways, and hydro canals for grow-out operations. Low stocking densities, ranging between less than 1 kg/m and around 25 kg/m (depending on
21018-474: The biological species concept, "the several versions" of the phylogenetic species concept, and the idea that species are of the same kind as higher taxa are not suitable for biodiversity studies (with the intention of estimating the number of species accurately). They further suggested that the concept works for both asexual and sexually-reproducing species. A version of the concept is Kevin de Queiroz 's "General Lineage Concept of Species". An ecological species
21204-505: The biological species concept, a cladistic species does not rely on reproductive isolation – its criteria are independent of processes that are integral in other concepts. Therefore, it applies to asexual lineages. However, it does not always provide clear cut and intuitively satisfying boundaries between taxa, and may require multiple sources of evidence, such as more than one polymorphic locus, to give plausible results. An evolutionary species, suggested by George Gaylord Simpson in 1951,
21390-428: The boundaries between closely related species become unclear with hybridisation , in a species complex of hundreds of similar microspecies , and in a ring species . Also, among organisms that reproduce only asexually , the concept of a reproductive species breaks down, and each clone is potentially a microspecies. Although none of these are entirely satisfactory definitions, and while the concept of species may not be
21576-445: The chemical composition of the water the fish had lived in, just as a tree's growth rings provide hints about dry and wet years. The bone is built with the chemical signature of the environment that hosted the fish. Researchers were able to tell where different individuals of Chinook were born and lived in the first year of their lives. Testing was done by measuring the strontium in the bones. Strontium can accurately show researchers
21762-433: The concept of a chronospecies can be applied. During anagenesis (evolution, not necessarily involving branching), some palaeontologists seek to identify a sequence of species, each one derived from the phyletically extinct one before through continuous, slow and more or less uniform change. In such a time sequence, some palaeontologists assess how much change is required for a morphologically distinct form to be considered
21948-416: The concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses ) are given a two-part name , called a "binomial". The first part of a binomial is the genus to which the species belongs. The second part
22134-427: The core of Great Lakes trash problem. Algae such as diatoms , along with other phytoplankton , are photosynthetic primary producers supporting the food web of the Great Lakes, and have been affected by global warming . The changes in the size or in the function of the primary producers may have a direct or an indirect impact on the food web. Photosynthesis carried out by diatoms constitutes about one fifth of
22320-750: The cyanobacteria ... will produce toxins", said Michael McKay, of the University of Windsor. Water quality testing was underway in August 2019. Until 1970, mercury was not listed as a harmful chemical, according to the United States Federal Water Quality Administration. In the 21st century, mercury has become more apparent in water tests. Mercury compounds have been used in paper mills to prevent slime from forming during their production, and chemical companies have used mercury to separate chlorine from brine solutions. Studies conducted by
22506-545: The energy for migration. Later, they change physiologically to live in salt water. They rely on eelgrass and seaweeds for camouflage (protection from predators), shelter, and foraging habitat as they make their way to the open ocean. Adult fish need a rich, open ocean habitat to acquire the strength needed to travel back upstream, escape predators, and reproduce before dying. In his book King of Fish , David Montgomery writes, "The reserves of fish at sea are important to restocking rivers disturbed by natural catastrophes." Thus, it
22692-435: The evidence to support hypotheses about evolutionarily divergent lineages that have maintained their hereditary integrity through time and space. Molecular markers may be used to determine diagnostic genetic differences in the nuclear or mitochondrial DNA of various species. For example, in a study done on fungi , studying the nucleotide characters using cladistic species produced the most accurate results in recognising
22878-427: The exact location and time of a fish swimming in a river. The total North Pacific fisheries harvest of the Chinook salmon in 2010 was some 1.4 million fish, corresponding to 7,000 tonnes ; 1.1 million of the fish were captured in the United States, and others were divided by Canada and Russia. The share of Chinook salmon from the total commercial Pacific salmon harvest was less than 1% by weight and only about 0.3% of
23064-481: The fish has spent five years in the ocean) have declined from being up to 3–5% of the population to being almost none. Ocean-4 chinook are also seeing a rapid decline in their population. This means that Chinook are not living as long as they used to. This trend has mostly been seen in Alaska, but also Oregon and Washington. New trends have also been seen regarding the size of Ocean-1, 2, 3, 4, and 5 from 1975 to 2015. The size of Chinook who have spent one and two years in
23250-455: The fish that were studied by researchers from the Rochman Lab. About 50 million pounds (23 kt) of fish is harvested each year from Great Lakes which has raised concerns on how this might affect human health. Microscopic pieces of plastic have also been found in drinking water coming from Great Lakes. It is estimated that nearly 40 million people in the region rely on drinking water from
23436-443: The fish washing up on shore; estimates by various governments have placed the percentage of Lake Michigan's biomass which was made up of alewives in the early 1960s as high as 90%. In the late 1960s, the various state and federal governments began stocking several species of salmonids, including the native lake trout as well as non-native chinook and coho salmon; by the 1980s, alewife populations had dropped drastically. The ruffe ,
23622-443: The fish. Mercury has been known for health related problems such as birth defects in humans and animals, and the near extinction of eagles in the Great Lakes region. The amount of raw sewage dumped into the waters was the primary focus of both the first Great Lakes Water Quality Agreement and federal laws passed in both countries during the 1970s. Implementation of secondary treatment of municipal sewage by major cities greatly reduced
23808-417: The glaciers "rebounded" as it was uncovered. Since the glaciers covered some areas longer than others, this glacial rebound occurred at different rates. The Great Lakes have a humid continental climate, Köppen climate classification Dfa (in southern areas) and Dfb (in northern parts) with varying influences from air masses from other regions including dry, cold Arctic systems, mild Pacific air masses from
23994-402: The harvestable size of 3–4 kilograms (7–9 pounds). The broodstock for the farms is usually selected from existing farm stock or sometimes sourced from wild populations. Eggs and milt are stripped manually from sexually mature salmon and incubated under conditions replicating the streams and rivers where the salmon would spawn naturally (at around 10–12 °C or 50–54 °F). After hatching,
24180-410: The historical average level. Although "true tides—changes in water level caused by the gravitational forces of the sun and moon—do occur in a semi-diurnal (twice daily) pattern", such changes are quite small and generally obscured by other forces. The lake levels are affected primarily by changes in regional meteorology and climatology. The outflows from Lakes Superior and Ontario are regulated, while
24366-573: The hook. The species has also established itself in Patagonian waters in South America, where both introduced and escaped hatchery fish have colonized rivers and established stable spawning runs . Chinook salmon have been found spawning in headwater reaches of the Rio Santa Cruz , apparently having migrated over 1,000 km (620 mi) from the ocean. The population is thought to be derived from
24552-488: The ice sheet left behind a large amount of meltwater ( Lake Algonquin , Lake Chicago , Glacial Lake Iroquois , and Champlain Sea ) that filled up the basins that the glaciers had carved, thus creating the Great Lakes as they are today. Because of the uneven nature of glacier erosion , some higher hills became Great Lakes islands. The Niagara Escarpment follows the contour of the Great Lakes between New York and Wisconsin. Land below
24738-545: The key indicators even in the current era of sophisticated analyses and measuring instruments. According to the bi-national (U.S. and Canadian) resource book, The Great Lakes: An Environmental Atlas and Resource Book : "The largest Great Lakes fish harvests were recorded in 1889 and 1899 at some 67,000 tonnes (66,000 long tons; 74,000 short tons) [147 million pounds]." By 1801, the New York Legislature found it necessary to pass regulations curtailing obstructions to
24924-597: The lake and causing blooms," said Michael McKay, executive director of the Great Lakes Institute for Environmental Research (GLIER) at the University of Windsor . The largest Lake Erie bloom to date occurred in 2015, exceeding the severity index at 10.5 and in 2011 at a 10. In early August 2019, satellite images depicted a bloom stretching up to 1,300 square kilometres on Lake Erie, with the heaviest concentration near Toledo, Ohio . A large bloom does not necessarily mean
25110-634: The lake shore as far north as the Grand Traverse Bay . Near Collingwood, Ontario , commercial fruit orchards, including a few wineries, exist near the shoreline of southern Nottawasaga Bay . The eastern shore of Lake Michigan and the southern shore of Lake Erie have many successful wineries because of the lakes' moderating effects, as do the large commercial fruit and wine growing areas of the Niagara Peninsula located between Lake Erie and Lake Ontario. A similar phenomenon allows wineries to flourish in
25296-525: The lakes has been observed since records began in 1918. The water level of Lake Michigan–Huron had remained fairly constant over the 20th century. Recent lake levels include record low levels in 2013 in Lakes Superior, Erie, and Michigan-Huron, followed by record high levels in 2020 in the same lakes. The water level in Lake Ontario has remained relatively constant in the same time period, hovering around
25482-464: The lakes' levels: diversion (the transfer of water to other watersheds) and consumption (substantially done today by the use of lake water to power and cool electric generation plants, resulting in evaporation). Outflows through the Chicago Sanitary and Ship Canal is more than balanced by artificial inflows via the Ogoki River and Long Lake/ Kenogami River diversions. Fluctuation of the water levels in
25668-513: The lakes. Only barges can access the Illinois Waterway system providing access to the Gulf of Mexico via the Mississippi River. Despite their vast size, large sections of the Great Lakes freeze over in winter, interrupting most shipping from January to March. Some icebreakers ply the lakes, keeping the shipping lanes open through other periods of ice on the lakes. The Great Lakes are connected by
25854-585: The lakes. The lakes connect to the Atlantic Ocean via the Saint Lawrence River , and to the Mississippi River basin through the Illinois Waterway . The Great Lakes are the largest group of freshwater lakes on Earth by total area and the second-largest by total volume. They contain 21% of the world's surface fresh water by volume. The total surface is 94,250 square miles (244,106 km ), and
26040-415: The lakes. The province of Ontario does not border Lake Michigan, and the state of Michigan does not border Lake Ontario. New York and Wisconsin's jurisdictions extend into two lakes, and each of the remaining states into one of the lakes. As the surfaces of Lakes Superior, Huron, Michigan, and Erie are all approximately the same elevation above sea level, while Lake Ontario is significantly lower, and because
26226-588: The land, which then filled with meltwater. The lakes have been a major source for transportation, migration, trade, and fishing, serving as a habitat to many aquatic species in a region with much biodiversity . The surrounding region is called the Great Lakes region , which includes the Great Lakes Megalopolis . Major cities within the region include, on the American side, from east to west, Buffalo , Cleveland , Detroit , Chicago , and Milwaukee ; and, on
26412-702: The largest of the Pacific salmon. In the Kenai River of Alaska, mature Chinook averaged 16.8 kg (37 lb 1 oz). The current sport-caught world record, 44.1 kg ( 97 + 1 ⁄ 4 lb), was caught on May 17, 1985, in the Kenai River. The commercial catch world record is 57 kg (126 lb) caught near Rivers Inlet , British Columbia, in the late 1970s. Chinook, like many other species of salmon, are considered euryhaline , and thus live in both saltwater and freshwater environments throughout their life. Once hatching, salmon spend one to eight years in
26598-418: The late 18th century. Maritime fur traders and explorers, such as George Vancouver , frequently acquired salmon by trade with the indigenous people of the Northwest coast. Lewis and Clark first encountered Chinook salmon as a gift from Chief Cameahwait , on August 13, 1805, near Lemhi Pass . Tasting it convinced Lewis they had crossed the continental divide. In Oregon, many Native American tribes, including
26784-468: The latter as the lower lakes and Lakes Michigan, Huron, and Superior as the upper lakes. This corresponds to thinking of lakes Erie and Ontario as "down south" and the others as "up north". Vessels sailing north on Lake Michigan are considered "upbound" even though they are sailing toward its effluent current. Lakes Huron and Michigan are sometimes considered a single lake, called Lake Michigan–Huron, because they are one hydrological body of water connected by
26970-494: The launch of the bi-national Great Lakes Fishery Commission . Several species of exotic water fleas have accidentally been introduced into the Great Lakes, such as the spiny waterflea, Bythotrephes longimanus , and the fishhook waterflea, Cercopagis pengoi , potentially having an effect on the zooplankton population. Several species of crayfish have also been introduced that may contend with native crayfish populations. More recently an electric fence has been set up across
27156-492: The life stage of the salmon), and the absence of disease in the fish mean New Zealand farmers do not need to use antibiotics or vaccines to maintain the health of their salmon stocks. The salmon are fed food pellets of fish meal specially formulated for Chinook salmon (typical proportions of the feed are: 45% protein, 22% fat, and 14% carbohydrate plus ash and water) that contain no steroids or other growth enhancers. Regulations and monitoring programmes ensure salmon are farmed in
27342-554: The longest freshwater migration route of any salmon, over 3,000 km (1,900 mi) from its mouth in the Bering Sea to spawning grounds upstream of Whitehorse, Yukon . Since Chinook rely on fat reserves for energy upon re-entering fresh water, commercial fish caught here are highly prized for their unusually high levels of heart-healthy omega-3 fatty acids . However, the high costs of harvest and transport from this rural area limits its affordability. The highest elevation Chinook spawn
27528-478: The low count of Chinook salmon present because of the collapse of the Sacramento River run, one of the biggest south of the Columbia. In April 2009, California again canceled the season. The Pacific Fishery Management Council's goal for the Sacramento River run is an escapement total (fish that return to freshwater spawn areas and hatcheries) of 122,000–180,000 fish. The 2007 escapement was estimated at 88,000, and
27714-476: The natural migrations of Atlantic salmon from Lake Erie into their spawning channels. In the early 19th century, the government of Upper Canada found it necessary to introduce similar legislation prohibiting the use of weirs and nets at the mouths of Lake Ontario's tributaries. Other protective legislation was passed, but enforcement remained difficult. On both sides of the Canada–United States border,
27900-791: The north. Nevertheless, they are consistently present and the distribution is well known only in Kamchatka . Elsewhere, information is scarce, but they have a patchy presence in the Anadyr River basin and parts of the Chukchi Peninsula . Also, in parts of the northern Magadan Oblast near the Shelikhov Gulf and Penzhina Bay , stocks might persist but remain poorly studied. In 1967, the Michigan Department of Natural Resources introduced Chinook into Lake Michigan and Lake Huron to control
28086-445: The number of fish. The trend has been down in the captures compared to the period before 1990, when the total harvest had been around 25,000 tonnes. Global production has, however, remained at a stable level because of increased aquaculture. The world's largest producer and market supplier of Chinook salmon is New Zealand. In 2009, New Zealand exported 5,088 tonnes (5,609 short tons) of Chinook salmon, marketed as king salmon, equating to
28272-478: The numerous fungi species of all the concepts studied. Versions of the phylogenetic species concept that emphasise monophyly or diagnosability may lead to splitting of existing species, for example in Bovidae , by recognising old subspecies as species, despite the fact that there are no reproductive barriers, and populations may intergrade morphologically. Others have called this approach taxonomic inflation , diluting
28458-412: The ocean (averaging from three to four years) before returning to their home rivers to spawn. The salmon undergo radical morphological changes as they prepare for the spawning event ahead. Salmon lose the silvery blue they had as ocean fish, and their color darkens, sometimes with a radical change in hue. Salmon are sexually dimorphic , and the male salmon develop canine-like teeth, and their jaws develop
28644-429: The ocean has been rising, while the size of Chinook of three to five years has been declining. The size increase was seen mainly in hatchery fish, not wild, and hatchery fish were often larger than wild, but the decrease was seen in both types of populations. Factors have been discovered that have influenced the size of the Chinook. They include, but are not limited to, the years they spent in fresh water before migrating to
28830-530: The ocean, reaching age eight. More northerly populations tend to have longer lives. Salmon need suitable spawning habitat. Clean, cool, oxygenated, sediment-free fresh water is essential for egg development. Chinook use larger sediment (gravel) sizes for spawning than other Pacific salmon. Riparian vegetation and woody debris help juvenile salmon by providing cover and maintaining low water temperatures. Chinook also need healthy ocean habitats. Juvenile salmon grow in clean, productive estuarine environments and gain
29016-420: The ocean, the time of year they were caught, which season run they participated in, and where they were caught. However, what is causing these negative trends is still not fully known or researched. Some possibilities can be climate change, pollution, and fishing practices. In California specifically, Chinook populations in the rivers have been declining. Chinook that are migratory are already more vulnerable, and
29202-463: The order amphipoda ), two types of copepods , and the deepwater sculpin (a spiny, large-headed fish). The Great Lakes are an important source of fishing . Early European settlers were astounded by both the variety and quantity of fish; there were 150 different species in the Great Lakes. Throughout history, fish populations were the early indicator of the condition of the Lakes and have remained one of
29388-506: The outflows of Michigan-Huron and Erie are not regulated at all. Ontario is the most tightly regulated, with its outflow controlled by the Moses-Saunders Power Dam , which explains its consistent historical levels. The Great Lakes contain 21% of the world's surface fresh water: 5,472 cubic miles (22,810 km ), or 6.0×10 U.S. gallons, that is 6 quadrillion U.S. gallons, (2.3×10 liters). The lakes contain about 84% of
29574-585: The paper is accepted for publication. The type material is usually held in a permanent repository, often the research collection of a major museum or university, that allows independent verification and the means to compare specimens. Describers of new species are asked to choose names that, in the words of the International Code of Zoological Nomenclature , are "appropriate, compact, euphonious, memorable, and do not cause offence". Books and articles sometimes intentionally do not identify species fully, using
29760-674: The person who named the species, while the antonym sensu lato ("in the broad sense") denotes a wider usage, for instance including other subspecies. Other abbreviations such as "auct." ("author"), and qualifiers such as "non" ("not") may be used to further clarify the sense in which the specified authors delineated or described the species. Species are subject to change, whether by evolving into new species, exchanging genes with other species, merging with other species or by becoming extinct. The evolutionary process by which biological populations of sexually-reproducing organisms evolve to become distinct or reproductively isolated as species
29946-507: The proliferation of dams and impoundments have multiplied, necessitating more regulatory efforts. Concerns by the mid-19th century included obstructions in the rivers which prevented salmon and lake sturgeon from reaching their spawning grounds. The Wisconsin Fisheries Commission noted a reduction of roughly 25% in general fish harvests by 1875. The states have removed dams from rivers where necessary. Overfishing has been cited as
30132-487: The result of misclassification leading to questions on whether there really are any ring species. The commonly used names for kinds of organisms are often ambiguous: "cat" could mean the domestic cat, Felis catus , or the cat family, Felidae . Another problem with common names is that they often vary from place to place, so that puma, cougar, catamount, panther, painter and mountain lion all mean Puma concolor in various parts of America, while "panther" may also mean
30318-573: The ring. Ring species thus present a difficulty for any species concept that relies on reproductive isolation. However, ring species are at best rare. Proposed examples include the herring gull – lesser black-backed gull complex around the North pole, the Ensatina eschscholtzii group of 19 populations of salamanders in America, and the greenish warbler in Asia, but many so-called ring species have turned out to be
30504-835: The rivers of Canterbury and North Otago: Rangitata River , the Opihi River , the Ashburton River , the Rakaia River , the Waimakariri River , the Hurunui River , and the Waiau Uwha River . The success of the latter introductions is thought to be partly attributable to the use of ova from autumn-run populations as opposed to ova from spring-run populations used in the first attempts. Whilst other salmon have also been introduced into New Zealand, only Chinook salmon (or king salmon as it
30690-428: The routine discharge of untreated sewage during the 1970s and 1980s. The International Joint Commission in 2009 summarized the change: "Since the early 1970s, the level of treatment to reduce pollution from waste water discharges to the Great Lakes has improved considerably. This is a result of significant expenditures to date on both infrastructure and technology, and robust regulatory systems that have proven to be, on
30876-508: The same species. This concept was narrowed in 2006 to a similarity of 98.7%. The average nucleotide identity (ANI) method quantifies genetic distance between entire genomes , using regions of about 10,000 base pairs . With enough data from genomes of one genus, algorithms can be used to categorize species, as for Pseudomonas avellanae in 2013, and for all sequenced bacteria and archaea since 2020. Observed ANI values among sequences appear to have an "ANI gap" at 85–95%, suggesting that
31062-529: The same species. When two species names are discovered to apply to the same species, the older species name is given priority and usually retained, and the newer name considered as a junior synonym, a process called synonymy . Dividing a taxon into multiple, often new, taxa is called splitting . Taxonomists are often referred to as "lumpers" or "splitters" by their colleagues, depending on their personal approach to recognising differences or commonalities between organisms. The circumscription of taxa, considered
31248-436: The shoreline of all the lakes would cover a distance roughly equivalent to travelling half-way around the world at the equator. A notable modern phenomenon is the formation of ice volcanoes over the lakes during wintertime. Storm-generated waves carve the lakes' ice sheet and create conical mounds through the eruption of water and slush. The process is only well-documented in the Great Lakes, and has been credited with sparing
31434-583: The southern shorelines from worse rocky erosion. It has been estimated that the foundational geology that created the conditions shaping the present day upper Great Lakes was laid from 1.1 to 1.2 billion years ago, when two previously fused tectonic plates split apart and created the Midcontinent Rift , which crossed the Great Lakes Tectonic Zone . A valley was formed providing a basin that eventually became modern day Lake Superior. When
31620-502: The species concept and making taxonomy unstable. Yet others defend this approach, considering "taxonomic inflation" pejorative and labelling the opposing view as "taxonomic conservatism"; claiming it is politically expedient to split species and recognise smaller populations at the species level, because this means they can more easily be included as endangered in the IUCN red list and can attract conservation legislation and funding. Unlike
31806-542: The species in New Zealand began in the 1970s when hatcheries were initially set up to enhance and support wild fish stocks, with the first commercial operations starting in 1976. After some opposition against their establishment by societal groups, including anglers, the first sea cage farm was established in 1983 at Big Glory Bay in Stewart Island by British Petroleum NZ Ltd. Today, the salmon are hatched in land-based hatcheries (several of which exist) and transferred to sea cages or freshwater farms, where they are grown out to
31992-414: The surface freshwater of North America; if the water were evenly distributed over the entire continent's land area, it would reach a depth of 5 feet (1.5 meters). This is enough water to cover the 48 contiguous U.S. states to a uniform depth of 9.5 feet (2.9 m). Although the lakes contain a large percentage of the world's fresh water, the Great Lakes supply only a small portion of U.S. drinking water on
32178-436: The tail. Another distinctive feature is a black gum line that is present in both salt and fresh water. Adult fish typically range in size from 60 to 90 cm (24 to 36 in), but may be up to 150 cm (58 in) in length; they average 4.5 to 22.7 kg (10 to 50 lb), but may reach 59 kg (130 lb). The meat can be either pink or white, depending on what the salmon have been feeding on. Chinook salmon are
32364-440: The total volume (measured at the low water datum) is 5,439 cubic miles (22,671 km ), slightly less than the volume of Lake Baikal (5,666 cu mi or 23,615 km , 22–23% of the world's surface fresh water). Because of their sea-like characteristics, such as rolling waves, sustained winds, strong currents, great depths, and distant horizons, the five Great Lakes have long been called inland seas . Depending on how it
32550-448: The treatment plant, are temporarily overwhelmed by heavy rainstorms. Local sewage treatment authorities then must release untreated effluent, a mix of rainwater and sewage, into local water bodies. While enormous public investments such as the Deep Tunnel projects in Chicago and Milwaukee have greatly reduced the frequency and volume of these events, they have not been eliminated. The number of such overflow events in Ontario, for example,
32736-455: The tributary rivers into the Great Lakes also dislocated sediments. In 1884, the New York Fish Commission determined that the dumping of sawmill waste (chips and sawdust) had impacted fish populations. The first U.S. Clean Water Act , passed by a Congressional override after being vetoed by U.S. President Richard Nixon in 1972, was a key piece of legislation, along with the bi-national Great Lakes Water Quality Agreement signed by Canada and
32922-416: The warmer and shallower portions of the system such as Lake Erie, Saginaw Bay , Green Bay , and the southernmost portion of Lake Michigan. By the mid-1980s, most jurisdictions bordering the Great Lakes had controlled phosphate detergents . Blue-green algae, or cyanobacteria blooms, have been problematic on Lake Erie since 2011. "Not enough is being done to stop fertilizer and phosphorus from getting into
33108-430: The west pick up the air and moisture from the lake surface, which is slightly warmer in relation to the cold surface winds above. As the slightly warmer, moist air passes over the colder land surface, the moisture often produces concentrated, heavy snowfall that sets up in bands or "streamers". This is similar to the effect of warmer air dropping snow as it passes over mountain ranges. During freezing weather with high winds,
33294-427: The west, and warm, wet tropical systems from the south and the Gulf of Mexico. The lakes have a moderating effect on the climate; they can also increase precipitation totals and produce lake effect snowfall . The Great Lakes can have an effect on regional weather called lake-effect snow , which is sometimes very localized. Even late in winter, the lakes often have no icepack in the middle. The prevailing winds from
33480-480: The whole, quite effective." The commission reported that all urban sewage treatment systems on the U.S. side of the lakes had implemented secondary treatment, as had all on the Canadian side except for five small systems. Though contrary to federal laws in both countries, those treatment system upgrades have not yet eliminated combined sewer overflow events. This describes when older sewerage systems, which combine storm water with sewage into single sewers heading to
33666-494: The wild. It is difficult to define a species in a way that applies to all organisms. The debate about species concepts is called the species problem. The problem was recognised even in 1859, when Darwin wrote in On the Origin of Species : I was much struck how entirely vague and arbitrary is the distinction between species and varieties. He went on to write: No one definition has satisfied all naturalists; yet every naturalist knows vaguely what he means when he speaks of
33852-436: The world. The second-largest island is Isle Royale in Lake Superior. Both of these islands are large enough to contain multiple lakes themselves—for instance, Manitoulin Island's Lake Manitou is the world's largest lake on a freshwater island. Some of these lakes even have their own islands, like Treasure Island in Lake Mindemoya in Manitoulin Island. The Great Lakes also have several peninsulas between them, including
34038-523: The zooplankton community of the lake. Scientists understand that the micro-aquatic life of the lakes is abundant but know very little about some of the most plentiful microbes and their environmental effects in the Great Lakes. Although a drop of lake water may contain 1 million bacteria cells and 10 million viruses , only since 2012 has there been a long-term study of the lakes' micro-organisms. Between 2012 and 2019 more than 160 new species have been discovered. Native habitats and ecoregions in
34224-438: Was Ontario's Collingwood Harbour in 1994; Ontario's Severn Sound followed in 2003. Presque Isle Bay in Pennsylvania is formally listed as in recovery, as is Ontario's Spanish Harbour. Dozens of other Areas of Concern have received partial cleanups such as the Rouge River (Michigan) and Waukegan Harbor (Illinois). Phosphate detergents were historically a major source of nutrient to the Great Lakes algae blooms in particular in
34410-411: Was done in hopes of helping restore the salmon population of Battle Creek . In June 2021, the California State Water Resources Control Board approved a plan by the United States Bureau of Reclamation to release water from Lake Shasta for irrigation use, which "significantly" increased the risk of extinction of winter-run Chinook in the Sacramento River. In an effort to restore the fish habitat of
34596-517: Was eliminated as the mussels were harvested for use as buttons by early Great Lakes entrepreneurs. The Great Lakes: An Environmental Atlas and Resource Book (1972) notes: "Only pockets remain of the once large commercial fishery." Water quality improvements realized during the 1970s and 1980s, combined with successful salmonid stocking programs, have enabled the growth of a large recreational fishery. The last commercial fisherman left Milwaukee in 2011 because of overfishing and anthropogenic changes to
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