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105-515: See also Classification of the Cactaceae A cactus ( pl. : cacti , cactuses , or less commonly, cactus ) is a member of the plant family Cactaceae ( / k æ k ˈ t eɪ s i . iː , - ˌ aɪ / ), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word cactus derives, through Latin, from the Ancient Greek word κάκτος ( káktos ),

210-453: A pericarpel . Tissue derived from the petals and sepals continues the pericarpel, forming a composite tube—the whole may be called a floral tube, although strictly speaking only the part furthest from the base is floral in origin. The outside of the tubular structure often has areoles that produce wool and spines. Typically, the tube also has small scale-like bracts , which gradually change into sepal-like and then petal-like structures, so

315-463: A cactus. Stem shapes vary considerably among cacti. The cylindrical shape of columnar cacti and the spherical shape of globular cacti produce a low surface area-to-volume ratio, thus reducing water loss, as well as minimizing the heating effects of sunlight. The ribbed or fluted stems of many cacti allow the stem to shrink during periods of drought and then swell as it fills with water during periods of availability. A mature saguaro ( Carnegiea gigantea )

420-546: A cluster may share a common root. Other cacti have a quite different appearance. In tropical regions, some grow as forest climbers and epiphytes . Their stems are typically flattened and almost leaf-like in appearance, with few or even no spines. Climbing cacti can be very large; a specimen of Hylocereus was reported as 100 meters (330 ft) long from root to the most distant stem. Epiphytic cacti, such as species of Rhipsalis or Schlumbergera , often hang downwards, forming dense clumps where they grow in trees high above

525-452: A common ancestor), so that Pereskioideae is not monophyletic although the three other subfamilies are. The Bayesian consensus cladogram from a 2005 study is shown below: Pereskia Clade A Pereskia Clade B Opuntioideae Maihuenioideae Cactoideae Five tribes have been recognized within the subfamily Opuntioideae : Tephrocacteae , Pterocacteae , Austrocylindropuntieae , Cylindropuntieae and Opuntieae . All but

630-462: A continuous supply of CO 2 during photosynthesis means the stomata must be open, so water vapor is continuously being lost. Plants using the C 3 mechanism lose as much as 97% of the water taken up through their roots in this way. A further problem is that as temperatures rise, the enzyme that captures CO 2 starts to capture more and more oxygen instead, reducing the efficiency of photosynthesis by up to 25%. Crassulacean acid metabolism (CAM)

735-496: A costly undertaking, conservation efforts often concentrate on maximizing the efficiency of the existing system. This may include chiselling compacted soils, creating furrow dikes to prevent runoff, and using soil moisture and rainfall sensors to optimize irrigation schedules. Usually large gains in efficiency are possible through measurement and more effective management of the existing irrigation system. The 2011 UNEP Green Economy Report notes that "[i]mproved soil organic matter from

840-417: A decrease in groundwater levels and if continued it can exhaust the resource. Ground and surface waters are connected and overuse of groundwater can reduce and, in extreme examples, diminish the water supply of lakes, rivers, and streams. In coastal regions, over pumping groundwater can increase saltwater intrusion which results in the contamination of groundwater water supply. Sustainable use of groundwater

945-409: A drought. The concentration of salts in the root cells of cacti is relatively high. All these adaptations enable cacti to absorb water rapidly during periods of brief or light rainfall. Thus, Ferocactus cylindraceus reportedly can take up a significant amount of water within 12 hours from as little as 7 mm (0.3 in) of rainfall, becoming fully hydrated in a few days. Although in most cacti,

1050-413: A greater volume than the body. Taproots may aid in stabilizing the larger columnar cacti. Climbing, creeping and epiphytic cacti may have only adventitious roots , produced along the stems where these come into contact with a rooting medium. Like their spines, cactus flowers are variable. Typically, the ovary is surrounded by material derived from stem or receptacle tissue, forming a structure called

1155-414: A high surface area-to-volume ratio, at maturity they contain little or no water, being composed of fibers made up of dead cells. Spines provide protection from herbivores and camouflage in some species, and assist in water conservation in several ways. They trap air near the surface of the cactus, creating a moister layer that reduces evaporation and transpiration . They can provide some shade, which lowers

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1260-411: A management plan to conserve that system and is often used for ensuring the right management plan to be put into action. The conservation of water is extremely important in order to preserve wildlife habitats. There are many organisms in temperate regions who are affected by shortages in water. Additionally, many freshwater organisms are increasingly feeling the impacts of water pollution as it disrupts

1365-513: A name originally used by Theophrastus for a spiny plant whose identity is now not certain. Cacti occur in a wide range of shapes and sizes. They are native to the Americas, ranging from Patagonia in the south to parts of western Canada in the north, with the exception of Rhipsalis baccifera , which is also found in Africa and Sri Lanka . Cacti are adapted to live in very dry environments, including

1470-418: A normal shoot, nodes bearing leaves or flowers would be separated by lengths of stem (internodes). In an areole, the nodes are so close together, they form a single structure. The areole may be circular, elongated into an oval shape, or even separated into two parts; the two parts may be visibly connected in some way (e.g. by a groove in the stem) or appear entirely separate (a dimorphic areole). The part nearer

1575-407: A number of tribes and subfamilies. The ICSG classification of the family recognizes four subfamilies: Pereskioideae (consisting only of the genus Pereskia ), Opuntioideae , Maihuenioideae (consisting only of the genus Maihuenia ) and Cactoideae . Molecular phylogenetic studies suggest that Pereskia is not monophyletic (i.e. its species are not the complete set of descendants of

1680-465: A particular plant is an example of, say, Mammillaria mammillaris , they should be able to compare it with the type specimen to which this name is permanently attached. Type specimens are normally prepared by compression and drying, after which they are stored in herbaria to act as definitive references. However, cacti are very difficult to preserve in this way; they have evolved to resist drying and their bodies do not easily compress. A further difficulty

1785-447: A rainstorm. A few species differ significantly in appearance from most of the family. At least superficially, plants of the genera Leuenbergeria , Rhodocactus and Pereskia resemble other trees and shrubs growing around them. They have persistent leaves, and when older, bark-covered stems. Their areoles identify them as cacti, and in spite of their appearance, they, too, have many adaptations for water conservation . Leuenbergeria

1890-657: A single genus, Calymmanthium . Of the remaining eight, only two ( Cacteae and Rhipsalideae ) have been shown to be monophyletic. A summary of the cladograms for the Cactoideae presented in a 2011 paper is shown below (ICSG tribes in bold). Blossfeldia Cacteae Calymmantheae (i.e. Calymmanthium ), species of Copiapoa and Frailea including members of Pachycereeae , Hylocereeae and Browningieae Rhipsalideae other core Cactoideae II, including members of Notocacteae , Trichocereeae , Cereeae and Browningieae The classification of

1995-619: A specimen, which, according to David Hunt , ensured he "left a trail of nomenclatural chaos that will probably vex cactus taxonomists for centuries." In 1984, it was decided that the Cactaceae Section of the International Organization for Succulent Plant Study should set up a working party, now called the International Cactaceae Systematics Group (ICSG), to produce consensus classifications down to

2100-592: A toilet are trying to make a difference in water consumption and waste. It is also possible to use/"pollute" the water in stages (keeping use in flush toilets for last), hereby allowing more use of the water for various tasks within a same cycle (before it needs to be purified again, which can also be done in-situ). Earthships often use such a setup. Also available are modern shower heads that help reduce wasting water: Old shower heads are said to use 5-10 gallons per minute, while new fixtures available use 2.5 gallons per minute and offer equal water coverage. Another method

2205-416: Is "water-use efficiency". Water use is considered inefficient if the same purpose of its use can be accomplished with less water. Technical efficiency derives from engineering practice where it is typically used to describe the ratio of output to input and is useful in comparing various products and processes. For example, one showerhead would be considered more efficient than another if it could accomplish

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2310-433: Is a mechanism adopted by cacti and other succulents to avoid the problems of the C 3 mechanism. In full CAM, the stomata open only at night, when temperatures and water loss are lowest. CO 2 enters the plant and is captured in the form of organic acids stored inside cells (in vacuoles ). The stomata remain closed throughout the day, and photosynthesis uses only this stored CO 2 . CAM uses water much more efficiently at

2415-832: Is also considered wasteful. According to EPA estimate, household leaks in the US can waste approximately 900 billion gallons (3.4 billion cubic meters) of water annually nationwide. Generally, water management agencies are reluctant or unwilling to give a concrete definition to a relatively vague concept of water waste . However, definition of water waste is often given in local drought emergency ordinances. One example refers to any acts or omissions, whether willful or negligent, that are "causing or permitting water to leak, discharge, flow or run to waste into any gutter, sanitary sewer, watercourse or public or private storm drain, or to any adjacent property, from any tap, hose, faucet, pipe, sprinkler, pond, pool, waterway, fountain or nozzle." In this example,

2520-449: Is an essential part of irrigation. Plants always take a lot of ground water thus ground water should be replenished. For crop irrigation, optimal water efficiency means minimizing losses due to evaporation , runoff , or subsurface drainage while maximizing production. An evaporation pan in combination with specific crop correction factors can be used to determine how much water is needed to satisfy plant requirements. Flood irrigation ,

2625-450: Is another method used to treat water. Recent studies show that sand filtration needs further improvements, but it is approaching optimization with its effectiveness at removing pathogens from water. Sand filtration is very effective at removing protozoa and bacteria, but struggles with removing viruses. Large-scale sand filtration facilities also require large surface areas to accommodate them. The removal of pathogens from recycled water

2730-558: Is available. Similar restrictions are in effect in the State of Victoria, Australia. Temporary water use bans (also known as "hosepipe bans") are used in England, Scotland, Wales and Northern Ireland. Strictly speaking, water that is discharged into the sewer, or directly to the environment is not wasted or lost. It remains within the hydrologic cycle and returns to the land surface and surface water bodies as precipitation. However, in many cases,

2835-419: Is beneficial reduction in water loss , use and waste of resources. Another is avoiding any damage to water quality . A third is improving water management practices that reduce the use or enhance the beneficial use of water. Technology solutions exist for households, commercial and agricultural applications to reduce the . Water conservation programs involved in social solutions are typically initiated at

2940-399: Is close to the ancestral species from which all cacti evolved) does have long-lasting leaves, which are, however, thickened and succulent in many species. Other species of cactus with long-lasting leaves, such as the opuntioid Pereskiopsis , also have succulent leaves. A key issue in retaining water is the ratio of surface area to volume. Water loss is proportional to surface area, whereas

3045-546: Is considered close to the ancestral species from which all cacti evolved. In tropical regions, other cacti grow as forest climbers and epiphytes (plants that grow on trees). Their stems are typically flattened, almost leaf-like in appearance, with fewer or even no spines, such as the well-known Christmas cactus or Thanksgiving cactus (in the genus Schlumbergera ). Cacti have a variety of uses: many species are used as ornamental plants, others are grown for fodder or forage, and others for food (particularly their fruit). Cochineal

3150-480: Is derived through Latin from the Ancient Greek κάκτος ( kaktos ), a name used by Theophrastus for a spiny plant, which may have been the cardoon ( Cynara cardunculus ). Later botanists, such as Philip Miller in 1754, divided cacti into several genera, which, in 1789, Antoine Laurent de Jussieu placed in his newly created family Cactaceae. By the early 20th century, botanists came to feel Linnaeus's name Cactus had become so confused as to its meaning (was it

3255-429: Is essential in water conservation. A fundamental component to water conservation strategy is communication and education outreach of different water programs. Developing communication that educates science to land managers, policy makers, farmers, and the general public is another important strategy utilized in water conservation. Communication of the science of how water systems work is an important aspect when creating

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3360-446: Is important to consider implementing water-conserving changes to industrial and commercial application use. It was found that high-income countries use roughly 59% of their water for industrial usage while low-income countries use 8% for industrial usage. One big change that industrial and commercial companies can implement are to improve the assessment and maintenance of water systems. It is easy to add water-efficient applications but it

3465-453: Is increasingly affordable, especially for the home gardener and in light of rising water rates. Using drip irrigation methods can save up to 30,000 gallons of water per year when replacing irrigation systems that spray in all directions. There are also cheap effective methods similar to drip irrigation such as the use of soaking hoses that can even be submerged in the growing medium to eliminate evaporation. As changing irrigation systems can be

3570-439: Is not unnecessarily being lost and there is no excess money being spent on utility bills. A third change that industrial and commercial companies can implement is installing a rain sensor . This sensor should be able to detect when precipitation is occurring and stop the program which would normally irrigate the land. After the rain ends, the sensor should turn the program back on and resume to its normal watering cycle. Water

3675-556: Is of high priority because wastewater always contains pathogens capable of infecting humans. The levels of pathogenic viruses have to be reduced to a certain level in order for recycled water to not pose a threat to human populations. Further research is necessary to determine more accurate methods of assessing the level of pathogenic viruses in treated wastewater. Wasting water is the flip side of water conservation, and in household applications, it means causing or permitting water discharge without any practical purpose. Inefficient water use

3780-833: Is particularly true of tree-living cacti, such as Rhipsalis and Schlumbergera , but also of some ground-living cacti, such as Ariocarpus . The spines of cacti are often useful in identification, since they vary greatly between species in number, color, size, shape and hardness, as well as in whether all the spines produced by an areole are similar or whether they are of distinct kinds. Most spines are straight or at most slightly curved, and are described as hair-like, bristle-like, needle-like or awl-like, depending on their length and thickness. Some cacti have flattened spines (e.g. Sclerocactus papyracanthus ). Other cacti have hooked spines. Sometimes, one or more central spines are hooked, while outer spines are straight (e.g., Mammillaria rekoi ). In addition to normal-length spines, members of

3885-470: Is present in Pereskia species. By studying the ratio of C to C incorporated into a plant—its isotopic signature —it is possible to deduce how much CO 2 is taken up at night and how much in the daytime. Using this approach, most of the Pereskia species investigated exhibit some degree of CAM-cycling, suggesting this ability was present in the ancestor of all cacti. Pereskia leaves are claimed to only have

3990-446: Is really a thin sheet of plastic that is placed over the soil. There are holes in the plastic for the plants to grow through. Some studies have shown that plastic mulch conserves water by reducing the evaporation of soil moisture, however, there haven't been enough applied studies to determine the total water savings that this practice may bring about. Water shortage has become an increasingly difficult problem to manage. More than 40% of

4095-414: Is said to be able to absorb as much as 200 U.S. gallons (760 L; 170 imp gal) of water during a rainstorm. The outer layer of the stem usually has a tough cuticle , reinforced with waxy layers, which reduce water loss. These layers are responsible for the grayish or bluish tinge to the stem color of many cacti. The stems of most cacti have adaptations to allow them to conduct photosynthesis in

4200-461: Is that many cacti were given names by growers and horticulturalists rather than botanists; as a result, the provisions of the International Code of Nomenclature for algae, fungi, and plants (which governs the names of cacti, as well as other plants) were often ignored. Curt Backeberg , in particular, is said to have named or renamed 1,200 species without one of his names ever being attached to

4305-720: Is the product of an insect that lives on some cacti. Many succulent plants in both the Old and New World – such as some Euphorbiaceae (euphorbias) – are also spiny stem succulents and because of this are sometimes incorrectly referred to as "cactus". The 1,500 to 1,800 species of cacti mostly fall into one of two groups of "core cacti": opuntias (subfamily Opuntioideae ) and "cactoids" (subfamily Cactoideae ). Most members of these two groups are easily recognizable as cacti. They have fleshy succulent stems that are major organs of photosynthesis . They have absent, small, or transient leaves . They have flowers with ovaries that lie below

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4410-405: Is the proper maintenance and inspection of it which will lead to long-term changes. A water conservation plan can be created, including adding various goals and benchmarks for both the employees and the company. Another change that industrial and commercial companies can make are to check water-consuming systems at regular intervals for any leaks or problems. By doing this, it will ensure that water

4515-522: Is to recycle the water of the shower directly, by means a semi-closed system which features a pump and filter. Such a setup (called a " water recycling shower ") has also been employed at the VIRTUe LINQ house. Besides recycling water, it also reuses the heat of the water (which would otherwise be lost). Contrary to the popular view that the most effective way to save water is to curtail water-using behavior (e.g., by taking shorter showers), experts suggest

4620-412: Is useful to supply water to the needy. Harvested and filtered rain water can be used for toilets, home gardening, lawn irrigation, and small scale agriculture . Another strategy in water conservation is protecting groundwater resources. When precipitation occurs, some infiltrates the soil and goes underground. Water in this saturation zone is called groundwater. Contamination of groundwater causes

4725-606: The Atacama Desert , one of the driest places on Earth. Because of this, cacti show many adaptations to conserve water. For example, almost all cacti are succulents , meaning they have thickened, fleshy parts adapted to store water. Unlike many other succulents, the stem is the only part of most cacti where this vital process takes place. Most species of cacti have lost true leaves, retaining only spines , which are highly modified leaves. As well as defending against herbivores , spines help prevent water loss by reducing air flow close to

4830-399: The C 3 mechanism with CAM restricted to stems. More recent studies show that "it is highly unlikely that significant carbon assimilation occurs in the stem"; Pereskia species are described as having "C 3 with inducible CAM." Leafless cacti carry out all their photosynthesis in the stem, using full CAM. As of February 2012, it is not clear whether stem-based CAM evolved once only in

4935-440: The monophyly of three of these subfamilies (not Pereskioideae), but have not supported all of the tribes or even genera below this level; indeed, a 2011 study found only 39% of the genera in the subfamily Cactoideae sampled in the research were monophyletic . Classification of the cacti currently remains uncertain and is likely to change. A 2005 study suggested the genus Pereskia as then circumscribed ( Pereskia sensu lato)

5040-440: The sepals and petals , often deeply sunken into a fleshy receptacle (the part of the stem from which the flower parts grow). All cacti have areoles —highly specialized short shoots with extremely short internodes that produce spines , normal shoots, and flowers. The remaining cacti fall into only two groups: three tree-like genera, Leuenbergeria , Pereskia and Rhodocactus (all formerly placed in Pereskia ), and

5145-483: The "columns" may be horizontal rather than vertical. Thus, Stenocereus eruca can be described as columnar even though it has stems growing along the ground, rooting at intervals. Cacti whose stems are even smaller may be described as globular (or globose). They consist of shorter, more ball-shaped stems than columnar cacti. Globular cacti may be solitary, such as Ferocactus latispinus , or their stems may form clusters that can create large mounds. All or some stems in

5250-535: The Cactaceae The classification of the family Cactaceae remains uncertain as of August 2023 . Since the mid-1990s, the system produced by the International Cactaceae Systematics Group (ICSG) of the International Organization for Succulent Plant Study has been used as the basis of many published classifications. Detailed treatments produced in the 21st century have divided the family into around 125–130 genera and 1,400–1,500 species, which are then arranged into

5355-542: The Cactaceae thus remains subject to change, from the genus level upwards. In 2016, David Hunt wrote that "the advent of molecular systematic studies in the past two decades will hopefully have a stabilizing effect on classification in due course, but has so far had the opposite effect". In 2010, Nyffler and Eggli produced a classification based on an explicit phylogeny. The broad outlines of their cladogram agree with that of Hern et al. (2011), shown above, although some details are different. Their classification corresponds to

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5460-448: The ICSG classification of the subfamily Cactoideae (as presented by Anderson in 2001 ) with a 2002 classification of the subfamily Opuntioideae. There are differences in the genera recognized in the systems; where the older system does not recognize the genus, "–" is shown. Unless otherwise indicated, the subfamily placement of the genera listed here is based on Anderson's 2001 presentation of

5565-456: The ICSG classification, as is the tribal placement of the genera of the subfamily Cactoideae. The division of the subfamily Opuntioideae into tribes is additional to the ICSG system. Water conservation Water conservation aims to sustainably manage the natural resource of fresh water , protect the hydrosphere , and meet current and future human demand . Water conservation makes it possible to avoid water scarcity . It covers all

5670-551: The US Environmental Protection Agency estimates that metering alone can reduce consumption by 20 to 40 percent. In addition to raising consumer awareness of their water use, metering is also an important way to identify and localize water leakage . Water metering might benefit society by providing a financial incentive to avoid waste in water use. Some researchers have suggested that water conservation efforts should be primarily directed at farmers, in light of

5775-402: The absence of leaves. This is discussed further below under Metabolism . Many cacti have roots that spread out widely, but only penetrate a short distance into the soil. In one case, a young saguaro only 12 cm (4.7 in) tall had a root system with a diameter of 2 m (7 ft), but no more than 10 cm (4 in) deep. Cacti can also form new roots quickly when rain falls after

5880-413: The amount of water present is proportional to volume. Structures with a high surface area-to-volume ratio, such as thin leaves, necessarily lose water at a higher rate than structures with a low area-to-volume ratio, such as thickened stems. Spines , which are modified leaves, are present on even those cacti with true leaves, showing the evolution of spines preceded the loss of leaves. Although spines have

5985-573: The cactus and providing some shade. In the absence of true leaves, cacti's enlarged stems carry out photosynthesis . Cactus spines are produced from specialized structures called areoles , a kind of highly reduced branch. Areoles are an identifying feature of cacti. As well as spines, areoles give rise to flowers , which are usually tubular and multipetaled. Many cacti have short growing seasons and long dormancies and are able to react quickly to any rainfall, helped by an extensive but relatively shallow root system that quickly absorbs any water reaching

6090-874: The city code also clarifies that "in the case of washing, "discharge," "flow" or "run to waste" means that water in excess of that necessary to wash, wet or clean the dirty or dusty object, such as an automobile, sidewalk, or parking area, flows to waste. Water utilities (and other media sources) often provide listings of wasteful water-use practices and prohibitions of wasteful uses. Examples include utilities in San Antonio, Texas. Las Vegas, Nevada, California Water Service company in California, and City of San Diego, California. The City of Palo Alto in California enforces permanent water use restrictions on wasteful practices such as leaks, runoff, irrigating during and immediately after rainfall, and use of potable water when non-potable water

6195-478: The cladogram shown below. Pereskia A Pereskia B Maihuenia incertae sedis Cylindropuntieae Opuntieae Blossfeldia ( Blossfeldieae ) Cacteae incertae sedis Phyllocacteae Rhipsalideae Notocacteae Cereeae The table below shows how the genera recognized by Nyffler and Eggli are placed into their subfamilies and tribes (they also use some subtribes, not shown here). The column headed "Older tribe (if different)" shows

6300-598: The core cacti, or separately in the opuntias and cactoids; CAM is known to have evolved convergently many times. To carry out photosynthesis, cactus stems have undergone many adaptations. Early in their evolutionary history, the ancestors of modern cacti (other than Leuenbergeria species) developed stomata on their stems and began to delay developing bark. However, this alone was not sufficient; cacti with only these adaptations appear to do very little photosynthesis in their stems. Stems needed to develop structures similar to those normally found only in leaves. Immediately below

6405-426: The desalination of fresh water. Despite this, many seawater desalination plants have been built in response to water shortages around the world. This makes it necessary to evaluate the impacts of seawater desalination and to find ways to improve desalination technology. Current research involves the use of experiments to determine the most effective and least energy intensive methods of desalination. Sand filtration

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6510-496: The ecosystem. " World Water Day " is celebrated on 22 March. Water conservation programs involved in social solutions are typically initiated at the local level, by either municipal water utilities or regional governments. Common strategies include public outreach campaigns, tiered water rates (charging progressively higher prices as water use increases), or restrictions on outdoor water use such as lawn watering and car washing. Cities in dry climates often require or encourage

6615-667: The end into more than one stigma . The stamens usually arise from all over the inner surface of the upper part of the floral tube, although in some cacti, the stamens are produced in one or more distinct "series" in more specific areas of the inside of the floral tube. The flower as a whole is usually radially symmetrical ( actinomorphic ), but may be bilaterally symmetrical ( zygomorphic ) in some species. Flower colors range from white through yellow and red to magenta. All cacti have some adaptations to promote efficient water use. Most cacti— opuntias and cactoids —specialize in surviving in hot and dry environments (i.e. are xerophytes ), but

6720-489: The ends of stems, which are still growing and forming new areoles. In Pereskia , a genus close to the ancestor of cacti, areoles remain active for much longer; this is also the case in Opuntia and Neoraimondia . The great majority of cacti have no visible leaves ; photosynthesis takes place in the stems (which may be flattened and leaflike in some species). Exceptions occur in three (taxonomically, four) groups of cacti. All

6825-401: The expense of water supply provision and leaves more fresh water in lakes, rivers and aquifers for other users and also for supporting ecosystems. For example, we should not treat toilet as a trash can. If we flush cigarette butts or tissues in it, we are wasting gallons of water. Because the process of recycling water cannot be accomplished. A concept that is closely related to water wasting

6930-472: The fact that crop irrigation accounts for 70% of the world's fresh water use. The agricultural sector of most countries is important both economically and politically, and water subsidies are common. Conservation advocates have urged removal of all subsidies to force farmers to grow more water-efficient crops and adopt less wasteful irrigation techniques. New technology poses a few new options for consumers, features such as full flush and half flush when using

7035-403: The first ancestors of modern cacti were already adapted to periods of intermittent drought. A small number of cactus species in the tribes Hylocereeae and Rhipsalideae have become adapted to life as climbers or epiphytes , often in tropical forests, where water conservation is less important. The absence of visible leaves is one of the most striking features of most cacti. Pereskia (which

7140-972: The first, Tephrocacteae , were shown to be "essentially monophyletic" in molecular phylogenetic study in 2009. A summary of the consensus Bayesian tree is shown below (tribes are bold; the number of species in the study is shown in parentheses). Puna (1), Maihueniopsis (3), Cumulopuntia (1) =  Tephrocacteae (part) Pterocactus (6) =  Pterocacteae Maihueniopsis (1), Puna (1), Tephrocactus (7) =  Tephrocacteae (part) Maihueniopsis (1), Cumulopuntia (5), Puna (1), Austrocylindropuntia (4) ≈  Austrocylindropuntieae Quiabentia (1), Pereskiopsis (2), Micropuntia (2), Cylindropuntia (12), Grusonia s.s. (1), Corynopuntia (6) ≈  Cylindropuntieae Miqueliopuntia (1), Tunilla (2), Maihueniopsis (1) Opuntia (2) Brasiliopuntia (1), Opuntia (1), Tacinga (2) Opuntia (28), Consolea (6), Nopalea (7) Many of

7245-570: The genera Leuenbergeria , Pereskia and Rhodocactus , the branches are covered with leaves, so the species of these genera may not be recognized as cacti. In most other cacti, the branches are more typically cactus-like, bare of leaves and bark and covered with spines, as in Pachycereus pringlei or the larger opuntias . Some cacti may become tree-sized but without branches, such as larger specimens of Echinocactus platyacanthus . Cacti may also be described as shrubby , with several stems coming from

7350-573: The genera within the Opuntioideae are not monophyletic. Maihueniopsis is highly polyphyletic , appearing in four separate lineages. The two largest genera within the subfamily, Opuntia and Cylindropuntia are also not monophyletic. The classification of the Opuntioideae is thus uncertain as of March 2012 ; Griffith and Porter say that changes in classification will require "broad information (of multiple data types) regarding all species of opuntioid cacti". The ICSG classification divides

7455-872: The genus Mammillaria and outgrowths almost like leaves in Ariocarpus species. The stem may also be ribbed or fluted in shape. The prominence of these ribs depends on how much water the stem is storing: when full (up to 90% of the mass of a cactus may be water), the ribs may be almost invisible on the swollen stem, whereas when the cactus is short of water and the stems shrink, the ribs may be very visible. The stems of most cacti are some shade of green, often bluish or brownish green. Such stems contain chlorophyll and are able to carry out photosynthesis; they also have stomata (small structures that can open and close to allow passage of gases). Cactus stems are often visibly waxy. Areoles are structures unique to cacti. Although variable, they typically appear as woolly or hairy areas on

7560-528: The genus or the family?) that it should not be used as a genus name. The 1905 Vienna botanical congress rejected the name Cactus and instead declared Mammillaria was the type genus of the family Cactaceae. It did, however, conserve the name Cactaceae, leading to the unusual situation in which the family Cactaceae no longer contains the genus after which it was named. The difficulties continued, partly because giving plants scientific names relies on " type specimens ". Ultimately, if botanists want to know whether

7665-557: The ground or from branches very low down, such as in Stenocereus thurberi . Smaller cacti may be described as columnar. They consist of erect, cylinder-shaped stems, which may or may not branch, without a very clear division into trunk and branches. The boundary between columnar forms and tree-like or shrubby forms is difficult to define. Smaller and younger specimens of Cephalocereus senilis , for example, are columnar, whereas older and larger specimens may become tree-like. In some cases,

7770-552: The ground surface. Cactus stems are often ribbed or fluted with a number of ribs which corresponds to a number in the Fibonacci numbers (2, 3, 5, 8, 13, 21, 34 etc.). This allows them to expand and contract easily for quick water absorption after rain, followed by retention over long drought periods. Like other succulent plants, most cacti employ a special mechanism called " crassulacean acid metabolism " (CAM) as part of photosynthesis. Transpiration , during which carbon dioxide enters

7875-500: The ground. The leafless, spiny stem is the characteristic feature of the majority of cacti (all belonging to the largest subfamily, the Cactoideae ). The stem is typically succulent, meaning it is adapted to store water. The surface of the stem may be smooth (as in some species of Opuntia ) or covered with protuberances of various kinds, which are usually called tubercles. These vary from small "bumps" to prominent, nipple-like shapes in

7980-426: The groundwater water supply to not be able to be used as a resource of fresh drinking water and the natural regeneration of contaminated groundwater can take years to replenish. Some examples of potential sources of groundwater contamination include storage tanks, septic systems, uncontrolled hazardous waste , landfills , atmospheric contaminants , chemicals, and road salts . Contamination of groundwater decreases

8085-438: The growing season and then lost (as in many species of Opuntia ). The small genus Maihuenia also relies on leaves for photosynthesis. The structure of the leaves varies somewhat between these groups. Opuntioids and Maihuenia have leaves that appear to consist only of a midrib. Even those cacti without visible photosynthetic leaves do usually have very small leaves, less than 0.5 mm (0.02 in) long in about half of

8190-507: The highest possible volume for water storage with the lowest possible surface area for water loss from transpiration . The tallest free-standing cactus is Pachycereus pringlei , with a maximum recorded height of 19.2 m (63 ft), and the smallest is Blossfeldia liliputiana , only about 1 cm (0.4 in) in diameter at maturity. A fully grown saguaro ( Carnegiea gigantea ) is said to be able to absorb as much as 200 U.S. gallons (760 L; 170 imp gal) of water during

8295-606: The installation of xeriscaping or natural landscaping in new homes to reduce outdoor water usage. Most urban outdoor water use in California is residential, illustrating a reason for outreach to households as well as businesses. One fundamental conservation goal is universal water metering . The prevalence of residential water metering varies significantly worldwide. Recent studies have estimated that water supplies are metered in less than 30% of UK households. Although individual water meters have often been considered impractical in homes with private wells or in multifamily buildings,

8400-479: The level of genera. Their system has been used as the basis of subsequent classifications. Detailed treatments published in the 21st century have divided the family into around 125–130 genera and 1,400–1,500 species, which are then arranged into a number of tribes and subfamilies. The ICSG classification of the cactus family recognized four subfamilies, the largest of which was divided into nine tribes. The subfamilies were: Molecular phylogenetic studies have supported

8505-532: The local level, by either municipal water utilities or regional governments. The Aims of water conservation efforts include: The key activities to conserve water are as follows: One of the strategies in water conservation is rainwater harvesting . Digging ponds, lakes, canals, expanding the water reservoir, and installing rain water catching ducts and filtration systems on homes are different methods of harvesting rain water. Many people in many countries keep clean containers so they can boil it and drink it, which

8610-959: The most efficient way is replacing toilets and retrofitting washers; as demonstrated by two household end use logging studies in the US. Water-saving technology for the home includes: Smart water meters are also a promising technology for reducing household water usage. A study conducted in Valencia, Spain, shows the potential that smart meter-based water consumption feedback has for conserving water in households. The findings showed that households that were equipped with smart water meters increased their water savings. This technology works to show people how much water they were using in their household, suggest ways they can reduce water usage, and incentivize water savings with physical rewards. Many water-saving devices (such as low-flush toilets) that are useful in homes can also be useful for business water saving. Other water-saving technology for businesses includes: It

8715-454: The much smaller Maihuenia . These two groups are rather different from other cacti, which means any description of cacti as a whole must frequently make exceptions for them. Species of the first three genera superficially resemble other tropical forest trees. When mature, they have woody stems that may be covered with bark and long-lasting leaves that provide the main means of photosynthesis. Their flowers may have superior ovaries (i.e., above

8820-484: The oldest and most common type, is often very uneven in distribution, as parts of a field may receive excess water in order to deliver sufficient quantities to other parts. Overhead irrigation , using center-pivot or lateral-moving sprinklers, has the potential for a much more equal and controlled distribution pattern. Drip irrigation is the most expensive and least-used type, but offers the ability to deliver water to plant roots with minimal losses. However, drip irrigation

8925-421: The outer epidermis, a hypodermal layer developed made up of cells with thickened walls, offering mechanical support. Air spaces were needed between the cells to allow carbon dioxide to diffuse inwards. The center of the stem, the cortex, developed " chlorenchyma " – a plant tissue made up of relatively unspecialized cells containing chloroplasts , arranged into a "spongy layer" and a " palisade layer " where most of

9030-448: The photosynthesis occurs. Naming and classifying cacti has been both difficult and controversial since the first cacti were discovered for science. The difficulties began with Carl Linnaeus . In 1737, he placed the cacti he knew into two genera, Cactus and Pereskia . However, when he published Species Plantarum in 1753—the starting point for modern botanical nomenclature—he relegated them all to one genus, Cactus . The word "cactus"

9135-442: The plant and water escapes, does not take place during the day at the same time as photosynthesis, but instead occurs at night. The plant stores the carbon dioxide it takes in as malic acid , retaining it until daylight returns, and only then using it in photosynthesis. Because transpiration takes place during the cooler, more humid night hours, water loss is significantly reduced. Many smaller cacti have globe-shaped stems, combining

9240-452: The points of attachment of the sepals and petals) and areoles that produce further leaves. The two species of Maihuenia have succulent but non-photosynthetic stems and prominent succulent leaves. Cacti show a wide variety of growth habits , which are difficult to divide into clear, simple categories. Cacti can be tree-like (arborescent), meaning they typically have a single more-or-less woody trunk topped by several to many branches . In

9345-461: The policies, strategies and activities to reach these aims. Population, household size and growth and affluence all affect how much water is used. Climate change and other factors have increased pressure on natural water resources . This is especially the case in manufacturing and agricultural irrigation . Many countries have successfully implemented policies to conserve water conservation. There are several key activities to conserve water. One

9450-452: The price of limiting the amount of carbon fixed from the atmosphere and thus available for growth. CAM-cycling is a less water-efficient system whereby stomata open in the day, just as in plants using the C 3 mechanism. At night, or when the plant is short of water, the stomata close and the CAM mechanism is used to store CO 2 produced by respiration for use later in photosynthesis. CAM-cycling

9555-406: The replenishment of available freshwater so taking preventative measures by protecting groundwater resources from contamination is an important aspect of water conservation. An additional strategy to water conservation is practicing sustainable methods of utilizing groundwater resources. Groundwater flows due to gravity and eventually discharges into streams. Excess pumping of groundwater leads to

9660-427: The sepals and petals cannot be clearly differentiated (and hence are often called " tepals "). Some cacti produce floral tubes without wool or spines (e.g. Gymnocalycium ) or completely devoid of any external structures (e.g. Mammillaria ). Unlike the flowers of most other cacti, Pereskia flowers may be borne in clusters. Cactus flowers usually have many stamens , but only a single style , which may branch at

9765-424: The source of the water is at a significant distance from the return point and may be in a different catchment. The separation between extraction point and return point can represent significant environmental degradation in the watercourse and riparian strip. What is "wasted" is the community's supply of water that was captured, stored, transported and treated to drinking quality standards. Efficient use of water saves

9870-481: The species of Leuenbergeria , Pereskia and Rhodocactus are superficially like normal trees or shrubs and have numerous leaves with a midrib and a flattened blade (lamina) on either side. This group is paraphyletic , forming two taxonomic clades . Many cacti in the opuntia group (subfamily Opuntioideae ) also have visible leaves, which may be long-lasting (as in Pereskiopsis species) or produced only during

9975-665: The species studied and almost always less than 1.5 mm (0.06 in) long. The function of such leaves cannot be photosynthesis; a role in the production of plant hormones, such as auxin , and in defining axillary buds has been suggested. Botanically, " spines " are distinguished from "thorns": spines are modified leaves, and thorns are modified branches. Cacti produce spines, always from areoles as noted above. Spines are present even in those cacti with leaves, such as Pereskia , Pereskiopsis and Maihuenia , so they clearly evolved before complete leaflessness. Some cacti only have spines when young, possibly only when seedlings. This

10080-510: The stem acts as the main organ for storing water, some cacti have in addition large taproots . These may be several times the length of the above-ground body in the case of species such as Copiapoa atacamensis , which grows in one of the driest places in the world, the Atacama Desert in northern Chile. Photosynthesis requires plants to take in carbon dioxide gas (CO 2 ). As they do so, they lose water through transpiration . Like other types of succulents , cacti reduce this water loss by

10185-432: The stems from which spines emerge. Flowers are also produced from areoles. In the genus Leuenbergeria , believed similar to the ancestor of all cacti, the areoles occur in the axils of leaves (i.e. in the angle between the leaf stalk and the stem). In leafless cacti, areoles are often borne on raised areas on the stem where leaf bases would have been. Areoles are highly specialized and very condensed shoots or branches. In

10290-428: The subfamily Cactoideae into nine tribes. However, phylogenetic research has not supported most of these tribes, nor even the genera of which they are composed. A 2011 study found that "an extraordinarily high proportion of genera" were not monophyletic, including 22 (61%) of the 36 genera in the subfamily Cactoideae sampled in the research. Of the nine tribes recognized within Cactoideae, one, Calymmantheae , comprises

10395-492: The subfamily Opuntioideae have relatively short spines, called glochids , that are barbed along their length and easily shed. These enter the skin and are difficult to remove due to being very fine and easily broken, causing long-lasting irritation. Most ground-living cacti have only fine roots , which spread out around the base of the plant for varying distances, close to the surface. Some cacti have taproots ; in genera such as Ariocarpus , these are considerably larger and of

10500-409: The temperature of the surface of the cactus, also reducing water loss. When sufficiently moist air is present, such as during fog or early morning mist, spines can condense moisture, which then drips onto the ground and is absorbed by the roots. The majority of cacti are stem succulents , i.e., plants in which the stem is the main organ used to store water. Water may form up to 90% of the total mass of

10605-421: The top of the stem then produces flowers, the other part spines. Areoles often have multicellular hairs ( trichomes ) that give the areole a hairy or woolly appearance, sometimes of a distinct color such as yellow or brown. In most cacti, the areoles produce new spines or flowers only for a few years and then become inactive. This results in a relatively fixed number of spines, with flowers being produced only from

10710-449: The use of green manures, mulching, and recycling of crop residues and animal manure increases the water holding capacity of soils and their ability to absorb water during torrential rains", which is a way to optimize the use of rainfall and irrigation during dry periods in the season. As seen in China, plastic mulch also has the potential to conserve water in agricultural practices. The "mulch"

10815-460: The way in which they carry out photosynthesis. "Normal" leafy plants use the C 3 mechanism : during daylight hours, CO 2 is continually drawn out of the air present in spaces inside leaves and converted first into a compound containing three carbon atoms ( 3-phosphoglycerate ) and then into products such as carbohydrates . The access of air to internal spaces within a plant is controlled by stomata , which are able to open and close. The need for

10920-476: The world's population live in a region where the demand for water exceeds its supply. The imbalance between supply and demand, along with persisting issues such as climate change and population growth , has made water reuse a necessary method for conserving water. There are a variety of methods used in the treatment of waste water to ensure that it is safe to use for irrigation of food crops and/or drinking water. Seawater desalination requires more energy than

11025-430: Was basal within the Cactaceae, but confirmed earlier suggestions it was not monophyletic , i.e., did not include all the descendants of a common ancestor. The Bayesian consensus cladogram from this study is shown below with subsequent generic changes added. Pereskia s.l. Clade A → Leuenbergeria Pereskia s.l. Clade B → Rhodocactus + Pereskia s.s. Opuntioideae Maihuenia Classification of

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