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70-524: About 19, see article The Buteoninae are a subfamily of birds of prey which consists of medium to large, broad-winged species . They have large, powerful, hooked beaks for tearing flesh from their prey, strong legs, and powerful talons . They also have extremely keen eyesight to enable them to spot potential prey from a distance. This subfamily contains the buzzards (buteonine hawks) with great diversity in appearance and form and some appearing eagle-like, with at least 50 species included overall in

140-658: A lammergeier might have killed Aeschylus by accident. Many stories of Brazilian indigenous peoples speak about children mauled by Uiruuetê, the Harpy Eagle in Tupi language . Various large raptors like golden eagles are reported attacking human beings, but its unclear if they intend to eat them or if they have ever been successful in killing one. Some fossil evidence indicates large birds of prey occasionally preyed on prehistoric hominids. The Taung Child , an early human found in Africa,

210-569: A "siege". The word heron first appeared in the English language around 1300, originating from Old French hairon, eron (12th century), earlier hairo (11th century), from Frankish haigiro or from Proto-Germanic *haigrô , *hraigrô . Herons are also known as shitepokes / ˈ ʃ aɪ t p oʊ k / , or euphemistically as shikepokes or shypokes . Webster's Dictionary suggests that herons were given this name because of their habit of defecating when flushed. The 1971 Compact Edition of

280-454: A distance or during flight, strong feet with sharp talons for grasping or killing prey, and powerful, curved beaks for tearing off flesh. Although predatory birds primarily hunt live prey, many species (such as fish eagles , vultures and condors ) also scavenge and eat carrion . Although the term "bird of prey" could theoretically be taken to include all birds that actively hunt and eat other animals, ornithologists typically use

350-467: A great deal of data is required. Dimorphisms can also be the product of intrasexual selection between males and females. It appears that both sexes of the species play a role in the sexual dimorphism within raptors; females tend to compete with other females to find good places to nest and attract males, and males competing with other males for adequate hunting ground so they appear as the most healthy mate. It has also been proposed that sexual dimorphism

420-465: A larger image to be projected onto the retina. The visual acuity of some large raptors such as eagles and Old World vultures are the highest known among vertebrates; the wedge-tailed eagle has twice the visual acuity of a typical human and six times that of the common ostrich , the vertebrate with the largest eyes. There are two regions in the retina, called the deep and shallow fovea, that are specialized for acute vision. These regions contain

490-664: A royal decree of James VI (1566–1625) of Scotland . The OED speculates that shiterow is a corruption of shiteheron . Another former name was heronshaw or hernshaw , derived from Old French heronçeau . Corrupted to handsaw , this name appears in Shakespeare 's Hamlet . A possible further corruption took place in the Norfolk Broads, where the heron is often referred to as a harnser . The herons are medium- to large-sized birds with long legs and necks. They exhibit very little sexual dimorphism in size. The smallest species

560-510: A study of little egrets and cattle egrets in India, the majority of the colonies surveyed contained both species. Nesting is seasonal in temperate species; in tropical species, it may be seasonal (often coinciding with the rainy season) or year-round. Even in year-round breeders, nesting intensity varies throughout the year. Tropical herons typically have only one breeding season per year, unlike some other tropical birds which may raise up to three broods

630-583: A variety of live aquatic prey. Their diet includes a wide variety of aquatic animals, including fish, reptiles, amphibians , crustaceans , molluscs , and aquatic insects. Individual species may be generalists or specialize in certain prey types, such as the yellow-crowned night heron , which specializes in crustaceans, particularly crabs . Many species also opportunistically take larger prey, including birds and bird eggs, rodents, and more rarely carrion . Even more rarely, herons eating acorns, peas, and grains have been reported, but most vegetable matter consumed

700-522: A year. Courtship usually takes part on the nest . Males arrive first and begin the building of the nest, where they display to attract females. During courtship, the male employs a stretch display and uses erectile neck feathers; the neck area may swell. The female risks an aggressive attack if she approaches too soon and may have to wait up to four days. In colonial species, displays involve visual cues, which can include adopting postures or ritual displays, whereas in solitary species, auditory cues, such as

770-582: Is a deprecated superorder within Raptores, formerly composed of the orders Falconiformes and Strigiformes. The clade was invalidated after 2012. Falconiformes is now placed in Eufalconimorphae , while Strigiformes is placed in Afroaves . The order Accipitriformes is believed to have originated 44 million years ago when it split from the common ancestor of the secretarybird ( Sagittarius serpentarius ) and

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840-729: Is a simplified phylogeny of Telluraves which is the clade where the birds of prey belong to along with passerines and several near-passerine lineages. The orders in bold text are birds of prey orders; this is to show the paraphyly of the group as well as their relationships to other birds. Accipitriformes (hawks, vultures, and relatives) [REDACTED] Strigiformes (owls) [REDACTED] Coraciimorphae (woodpeckers, rollers, hornbills, etc.) [REDACTED] Cariamiformes (seriemas) [REDACTED] Falconiformes (falcons) [REDACTED] Psittacopasserae (parrots and songbirds) [REDACTED] A recent phylogenomic study from Wu et al. (2024) has found an alternative phylogeny for

910-411: Is absent. Generally, herons lay between three and seven eggs . Larger clutches are reported in the smaller bitterns and more rarely some of the larger day herons, and single-egg clutches are reported for some of the tiger herons. Clutch size varies by latitude within species, with individuals in temperate climates laying more eggs than tropical ones. On the whole, the eggs are glossy blue or white, with

980-402: Is accidental. The most common hunting technique is for the bird to sit motionless on the edge of or standing in shallow water and to wait until prey comes within range. Birds may either do this from an upright posture, giving them a wider field of view for seeing prey or from a crouched position, which is more cryptic and means the bill is closer to the prey when it is located. Having seen prey,

1050-492: Is an order of telluravian birds consisting of the living seriemas and extinct terror birds . Jarvis et al. 2014 suggested including them in the category of birds of prey, and McClure et al. 2019 considered seriemas to be birds of prey. The Peregrine Fund also considers seriemas to be birds of prey. Like most birds of prey, seriemas and terror birds prey on vertebrates . However, seriemas were not traditionally considered birds of prey. There were traditionally classified in

1120-538: Is believed to have been killed by an eagle-like bird similar to the crowned eagle . The Haast's eagle may have preyed on early humans in New Zealand , and this conclusion would be consistent with Maori folklore . Leptoptilos robustus might have preyed on both Homo floresiensis and anatomically modern humans, and the Malagasy crowned eagle , teratorns , Woodward's eagle and Caracara major are similar in size to

1190-445: Is commonly believed that the dimorphisms found in raptors occur due to sexual selection or environmental factors. In general, hypotheses in favor of ecological factors being the cause for sexual dimorphism in raptors are rejected. This is because the ecological model is less parsimonious , meaning that its explanation is more complex than that of the sexual selection model. Additionally, ecological models are much harder to test because

1260-507: Is derived from the Latin word rapio , meaning "to seize or take by force". The common names for various birds of prey are based on structure, but many of the traditional names do not reflect the evolutionary relationships between the groups. Many of these English language group names originally referred to particular species encountered in Britain . As English-speaking people travelled further,

1330-521: Is generally long and harpoon-like. It can vary from extremely narrow, as in the agami heron , to wider as in the grey heron . The most atypical heron bill is owned by the boat-billed heron , which has a broad, thick bill. Herons' bills and other bare parts of the body are usually yellow, black, or brown in colour, although this can vary during the breeding season. The wings are broad and long, exhibiting 10 or 11 primary feathers (the boat-billed heron has only nine), 15–20 secondaries, and 12 rectrices (10 in

1400-411: Is merely the product of disruptive selection , and is merely a stepping stone in the process of speciation, especially if the traits that define gender are independent across a species. Sexual dimorphism can be viewed as something that can accelerate the rate of speciation. In non-predatory birds, males are typically larger than females. However, in birds of prey, the opposite is the case. For instance,

1470-495: Is now regarded as a member of the Ardeidae. Although herons resemble birds in some other families, such as the storks , ibises , spoonbills , and cranes , they differ from these in flying with their necks retracted, not outstretched. They are also one of the bird groups that have powder down . Some members of this group nest colonially in trees, while others, notably the bitterns, use reed beds . A group of herons has been called

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1540-623: Is unclear at the moment, and molecular studies have so far suffered from studying only a small number of taxa. Especially among the subfamily Ardeinae, the relationships are very inadequately resolved. The arrangement presented here should be considered provisional. A 2008 study suggests that this family belongs to the Pelecaniformes . In response to these findings, the International Ornithological Congress reclassified Ardeidae and their sister taxa Threskiornithidae under

1610-400: Is usually considered the dwarf bittern , which measures 25–30 cm (10–12 in) in length, although all the species in the genus Ixobrychus are small and many broadly overlap in size. The largest species of heron is the goliath heron , which stands up to 152 cm (60 in) tall. All herons can retract their necks by folding them into a tight S-shape, due to the modified shape of

1680-506: The Pacific reef heron , both dark and light colour morphs exist, and the percentage of each morph varies geographically; its white morphs only occur in areas with coral beaches. The herons are a widespread family with a cosmopolitan distribution . They exist on all continents except Antarctica and are present in most habitats except the coldest extremes of the Arctic, extremely high mountains, and

1750-640: The grey heron is mostly sedentary in Britain, but mostly migratory in Scandinavia. Birds are particularly inclined to disperse widely after breeding, but before the annual migration, where the species is colonial, searching out new feeding areas and reducing the pressures on feeding grounds near the colony. The migration typically occurs at night, usually as individuals or in small groups. The herons, egrets and bitterns are carnivorous . The members of this family are mostly associated with wetlands and water and feed on

1820-497: The kestrel is a type of falcon in which males are the primary providers, and the females are responsible for nurturing the young. In this species, the smaller the kestrels are, the less food is needed and thus, they can survive in environments that are harsher. This is particularly true in the male kestrels. It has become more energetically favorable for male kestrels to remain smaller than their female counterparts because smaller males have an agility advantage when it comes to defending

1890-641: The merlin ( Falco columbarius ). The taxonomy of Carl Linnaeus grouped birds (class Aves) into orders, genera, and species, with no formal ranks between genus and order. He placed all birds of prey into a single order, Accipitres , subdividing this into four genera: Vultur (vultures), Falco (eagles, hawks, falcons, etc.), Strix (owls), and Lanius (shrikes). This approach was followed by subsequent authors such as Gmelin , Latham and Turton . Louis Pierre Vieillot used additional ranks: order, tribe, family, genus, species. Birds of prey (order Accipitres) were divided into diurnal and nocturnal tribes;

1960-489: The paleognath Lithornis vulturinus . In Buddhism, a heron symbolizes purity, transformation and the wisdom of the Buddha. In addition, as a bird that transcends elements – on the earth, in the water and in the air – the heron symbolizes the expansion of awareness and the ubiquity of consciousness. In some Native American cultures, this bird symbolizes renewal, rejuvenation and rebirth – an ever present reminder that we are all

2030-509: The stripe-backed bittern ( Ixobrychus involucris ) were nested with members of the genus Botaurus . Hruska and collaborators resurrected the genus Calherodius Peters , 1931 to contain two night herons (the white-backed night heron and the white-eared night heron ) that were previously placed in Gorsachius . The western cattle egret ( Bubulcus ibis ) was embedded in the genus Ardea . The eastern cattle egret ( Bubulcus coromandus )

2100-471: The Haast's eagle, implying that they similarly could pose a threat to a human being. Birds of prey have incredible vision and rely heavily on it for a number of tasks. They utilize their high visual acuity to obtain food, navigate their surroundings, distinguish and flee from predators, mating, nest construction, and much more. They accomplish these tasks with a large eye in relation to their skull, which allows for

2170-510: The IOC lists 74 heron species, divided into 18 genera. Other prehistoric and fossil species are included in the respective genus accounts. In addition, Proherodius is a disputed fossil which was variously considered a heron or one of the extinct long-legged waterfowl , the Presbyornithidae . It is only known from a sternum ; a tarsometatarsus that had been assigned to it actually belongs to

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2240-539: The Oxford English Dictionary describes the use of shitepoke for the small green heron of North America ( Butorides virescens ) as originating in the United States, citing a published example from 1853. The OED also observes that shiterow or shederow are terms used for herons, and also applied as derogatory terms meaning a thin, weakly person . This name for a heron is found in a list of game birds in

2310-489: The accipitrid species. The phylogeny of Accipitriformes is complex and difficult to unravel. Widespread paraphylies were observed in many phylogenetic studies. More recent and detailed studies show similar results. However, according to the findings of a 2014 study, the sister relationship between larger clades of Accipitriformes was well supported (e.g. relationship of Harpagus kites to buzzards and sea eagles and these latter two with Accipiter hawks are sister taxa of

2380-691: The bills of birds of prey shown by hunters. In Britain, kites and buzzards were seen as destroyers of game and killed, for instance in 1684–5 alone as many as 100 kites were killed. Rewards for their killing were also in force in the Netherlands from 1756. From 1705 to 1800, it has been estimated that 624087 birds of prey were killed in a part of Germany that included Hannover, Luneburg, Lauenburg and Bremen with 14125 claws deposited just in 1796–97. Many species also develop lead poisoning after accidental consumption of lead shot when feeding on animals that had been shot by hunters. Lead pellets from direct shooting that

2450-522: The biogeographic realms of the Southern Hemisphere. The appearance of migratory behaviour occurred in the tropics parallel with the range expansion of migratory species to temperate habitats. Similar results of southern origin in other taxonomic groups can be found in the literature. Distribution and biogeographic history highly determine the origin of migration in birds of prey. Based on some comparative analyses, diet breadth also has an effect on

2520-581: The birds have escaped from also cause reduced fitness and premature deaths. Some evidence supports the contention that the African crowned eagle occasionally views human children as prey, with a witness account of one attack (in which the victim, a seven-year-old boy, survived and the eagle was killed), and the discovery of part of a human child skull in a nest. This would make it the only living bird known to prey on humans, although other birds such as ostriches and cassowaries have killed humans in self-defense and

2590-407: The bitterns). The feathers of the herons are soft and the plumage is usually blue, black, brown, grey, or white, and can often be strikingly complex. Amongst the day herons, little sexual dimorphism in plumage is seen (except in the pond-herons); however, for the night herons and smaller bitterns, plumage differences between the sexes are the rule. Many species also have different colour morphs . In

2660-526: The cervical vertebrae, of which they have 20 or 21; the neck is retracted during flight, unlike most other long-necked birds. The neck is longer in the day herons than the night herons and bitterns. The legs are long and strong, and in almost every species are unfeathered from the lower part of the tibia (the exception is the zigzag heron ). In flight, the legs and feet are generally held in a horizontal position, pointing backwards. Toes are long and thin, with three pointing forwards and one backwards. The bill

2730-516: The clade containing Aquilinae and Harpiinae). The diurnal birds of prey are formally classified into six families of two different orders (Accipitriformes and Falconiformes). These families were traditionally grouped together in a single order Falconiformes but are now split into two orders, the Falconiformes and Accipitriformes . The Cathartidae are sometimes placed in a separate order Cathartiformes . Formerly, they were sometimes placed in

2800-517: The deep booming of the bitterns, are important. The exception to this is the boat-billed heron, which pairs up away from the nesting site. Having paired, they continue to build the nest in almost all species, although in the little bittern and least bittern , only the male works on the nest. Some ornithologists have reported observing female herons attaching themselves to impotent mates, then seeking sexual gratification elsewhere. The nests of herons are usually found near or above water. Although

2870-433: The driest deserts. Almost all species are associated with water; they are essentially non-swimming waterbirds that feed on the margins of lakes, rivers, swamps, ponds, and the sea. They are predominantly found in lowland areas, although some species live in alpine areas, and the majority of species occur in the tropics. The herons are a highly mobile family, with most species being at least partially migratory ; for example,

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2940-439: The evolution of migratory behaviour in this group, but its relevance needs further investigation. The evolution of migration in animals seems to be a complex and difficult topic with many unanswered questions. A recent study discovered new connections between migration and the ecology, life history of raptors. A brief overview from abstract of the published paper shows that "clutch size and hunting strategies have been proved to be

3010-403: The exception being the large bitterns, which lay olive-brown eggs. Analyses of skeletons, mainly skulls , suggested that the Ardeidae could be split into a diurnal and a crepuscular / nocturnal group which included the bitterns. From DNA studies, and from skeletal analyses that focussed more on bones of the body and limbs, that two-group division has been revealed to be incorrect. Rather,

3080-462: The existing literature combining anatomical, genetic, and behavioural studies showed that, in general, raptors have functional olfactory systems that they are likely to use in a range of different contexts. Birds of prey have been historically persecuted both directly and indirectly. In the Danish Faroe Islands , there were rewards Naebbetold (by royal decree from 1741) given in return for

3150-493: The familiar names were applied to new birds with similar characteristics. Names that have generalised this way include: kite ( Milvus milvus ), sparrowhawk or sparhawk ( Accipiter nisus ), goshawk ( Accipiter gentilis ), kestrel ( Falco tinninculus ), hobby ( Falco subbuteo ), harrier (simplified from "hen-harrier", Circus cyaneus ), buzzard ( Buteo buteo ). Some names have not generalised, and refer to single species (or groups of closely related (sub)species), such as

3220-586: The family Ardeidae , with 74 recognised species , some of which are referred to as egrets or bitterns rather than herons. Members of the genus Botaurus are referred to as bitterns, and, together with the zigzag heron , or zigzag bittern, in the monotypic genus Zebrilus , form a monophyletic group within the Ardeidae. Egrets do not form a biologically distinct group from herons, and tend to be named differently because they are mainly white or have decorative plumes in breeding plumage. Herons, by evolutionary adaptation, have long beaks. The classification of

3290-468: The feet are used to flush out hidden prey. The wings may be used to frighten prey (or possibly attract it to shade) or to reduce glare; the most extreme example of this is exhibited by the black heron , which forms a full canopy with its wings over its body. Some species of heron, such as the little egret and grey heron, have been documented using bait to lure prey to within striking distance. Herons may use items already in place, or actively add items to

3360-424: The head is moved from side to side, so that the heron can calculate the position of the prey in the water and compensate for refraction , and then the bill is used to spear the prey. In addition to sitting and waiting, herons may feed more actively. They may walk slowly, around or less than 60 paces a minute, snatching prey when it is observed. Other active feeding behaviours include foot stirring and probing, where

3430-400: The highest density of photoreceptors, and provide the highest points of visual acuity. The deep fovea points forward at an approximate 45° angle, while the shallow fovea points approximately 15° to the right or left of the head axis. Several raptor species repeatedly cock their heads into three distinct positions while observing an object. First, is straight ahead with their head pointed towards

3500-410: The highly acute deep fovea. Like all birds, raptors possess tetrachromacy , however, due to their emphasis on visual acuity, many diurnal birds of prey have little ability to see ultraviolet light as this produces chromatic aberration which decreases the clarity of vision. Heron 18 extant, see text Cochlearidae Herons are long-legged, long-necked, freshwater and coastal birds in

3570-474: The individual heron/egret species is fraught with difficulty, and no clear consensus exists about the correct placement of many species into either of the two major genera, Ardea and Egretta . Similarly, the relationships of the genera in the family are not completely resolved. However, one species formerly considered to constitute a separate monotypic family, the Cochlearidae or the boat-billed heron ,

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3640-618: The many lineages of Accipitridae which seem to have radiated out of Africa or South Asia, the Buteoninae clearly originated in the Americas based on fossil records and current species distributions (more than 75% of the extant raptors from this lineage are found in the Americas). A genus level cladogram of the Buteoninae is shown below. It is based on a molecular phylogenetic study of the family Accipitridae by Therese Catanach and collaborators that

3710-583: The most important variables in shaping distribution areas, and also the geographic dissimilarities may mask important relationships between life history traits and migratory behaviours. The West Palearctic-Afrotropical and the North-South American migratory systems are fundamentally different from the East Palearctic-Indomalayan system, owing to the presence versus absence of ecological barriers." Maximum entropy modelling can help in answering

3780-821: The narrower definition followed in this page, excluding many piscivorous predators such as storks , cranes , herons , gulls , skuas , penguins , and kingfishers , as well as many primarily insectivorous birds such as passerines (e.g. shrikes ), nightjars , frogmouths , songbirds such as crows and ravens , alongside opportunistic predators from predominantly frugivorous or herbivorous ratites such as cassowaries and rheas . Some extinct predatory telluravian birds had talons similar to those of modern birds of prey, including mousebird relatives ( Sandcoleidae ), and Messelasturidae indicating possible common descent . Some Enantiornithes also had such talons, indicating possible convergent evolution , as enanthiornithines weren't even modern birds . The term raptor

3850-444: The nest and hunting. Larger females are favored because they can incubate larger numbers of offspring, while also being able to brood a larger clutch size. It is a long-standing belief that birds lack any sense of smell, but it has become clear that many birds do have functional olfactory systems . Despite this, most raptors are still considered to primarily rely on vision, with raptor vision being extensively studied. A 2020 review of

3920-450: The nests of a few species have been found on the ground where suitable trees or shrubs are unavailable, they are typically placed in vegetation. Trees are used by many species, and here they may be placed high up from the ground, whereas species living in reed beds may nest very close to the ground. Though the majority of nesting of herons is seen in or immediately around water, colonies commonly occur in several cities when human persecution

3990-629: The object. Second and third are sideways to the right or left of the object, with their head axis positioned approximately 40° adjacent to the object. This movement is believed to be associated with lining up the incoming image to fall on the deep fovea. Raptors will choose which head position to use depending on the distance to the object. At distances as close as 8m, they used primarily binocular vision. At distances greater than 21m, they spent more time using monocular vision. At distances greater than 40m, they spent 80% or more time using their monocular vision. This suggests that raptors tilt their head to rely on

4060-464: The oldest dates published so far in the case of birds of prey. For example, a previous reconstruction of migratory behaviour in one Buteo clade with a result of the origin of migration around 5 million years ago was also supported by that study. Migratory species of raptors may have had a southern origin because it seems that all of the major lineages within Accipitridae had an origin in one of

4130-412: The order Ciconiiformes . The secretary bird and/or osprey are sometimes listed as subfamilies of Acciptridae: Sagittariinae and Pandioninae, respectively. Australia's letter-winged kite is a member of the family Accipitridae , although it is a nocturnal bird. The nocturnal birds of prey—the owls —are classified separately as members of two extant families of the order Strigiformes : Below

4200-503: The order Gruiformes . And they are still not considered birds of prey in general parlance. Their bodies are also shaped completely differently from birds of prey. They have long legs and long necks. While secretarybirds also have long legs, they otherwise resemble raptors. Seriemas do not. Their beak is hooked, but too long. Migratory behaviour evolved multiple times within accipitrid raptors. The earliest event occurred nearly 14 to 12 million years ago. This result seems to be one of

4270-601: The order Pelecaniformes instead of the previous order of Ciconiiformes . The cladogram shown below is based on a molecular phylogenetic study of the Ardeidae by Jack Hruska and collaborators published in 2023. For several species these results conflict with the taxonomy published online in July 2023 by Frank Gill , Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC). The least bittern ( Ixobrychus exilis ) and

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4340-808: The owls remained monogeneric (family Ægolii, genus Strix ), whilst the diurnal raptors were divided into three families: Vulturini, Gypaëti, and Accipitrini. Thus Vieillot's families were similar to the Linnaean genera, with the difference that shrikes were no longer included amongst the birds of prey. In addition to the original Vultur and Falco (now reduced in scope), Vieillot adopted four genera from Savigny: Phene , Haliæetus , Pandion , and Elanus . He also introduced five new genera of vultures ( Gypagus , Catharista , Daptrius , Ibycter , Polyborus ) and eleven new genera of accipitrines ( Aquila , Circaëtus , Circus , Buteo , Milvus , Ictinia , Physeta , Harpia , Spizaëtus , Asturina , Sparvius ). Falconimorphae

4410-820: The placement of the birds of prey. Their analysis has found support in a clade consisting of the Strigiformes and Accipitriformes in new clade Hieraves . Hieraves was also recovered to be the sister clade to Australaves (which it includes the Cariamiformes and Falconiformes along with Psittacopasserae ). Below is their phylogeny from the study. Coraciimorphae (woodpeckers, rollers, hornbills, etc.) [REDACTED] Strigiformes (owls) [REDACTED] Accipitriformes (hawks, vultures, and relatives) [REDACTED] Cariamiformes (seriemas) [REDACTED] Falconiformes (falcons) [REDACTED] Psittacopasserae (parrots and songbirds) [REDACTED] Cariamiformes

4480-464: The question: why species winters at one location while the others are elsewhere. Temperature and precipitation related factors differ in the limitation of species distributions. "This suggests that the migratory behaviours differ among the three main migratory routes for these species" which may have important conservational consequences in the protection of migratory raptors. Birds of prey (raptors) are known to display patterns of sexual dimorphism . It

4550-452: The similarities in skull morphology among certain herons reflect convergent evolution to cope with the different challenges of daytime and nighttime feeding. Today, it is believed that three major groups can be distinguished, which are: The night herons may still warrant separation from the day herons and egrets (as subfamily Nycticoracinae, as it was traditionally done). However, the position of some genera (e.g. Butorides or Syrigma )

4620-481: The subfamily. At one time, several types were grouped, including large assemblages such as booted eagles, but modern studies using mitochondrial DNA clarified that this subfamily was smaller than formerly classified. The subfamily Buteoninae was introduced (as "Buteonina") by the Irish zoologist Nicholas Vigors in 1825 with Buteo as the type genus . The subfamily includes about 79 currently recognized species. Unlike

4690-439: The success rate of prey capture increased 3.6 times over solitary foraging. While the family exhibits a range of breeding strategies, overall, the herons are monogamous and mostly colonial . Most day herons and night herons are colonial, or partly colonial depending on circumstances, whereas the bitterns and tiger herons are mostly solitary nesters. Colonies may contain several species, as well as other species of waterbirds. In

4760-617: The water to attract fish such as the banded killifish . Items used may be man-made, such as bread; alternatively, striated herons in the Amazon have been watched repeatedly dropping seeds, insects, flowers, and leaves into the water to catch fish. Three species, the black-headed heron , whistling heron , and especially the cattle egret , are less tied to watery environments and may feed far away from water. Cattle egrets improve their foraging success by following large grazing animals, catching insects flushed by their movement. One study found that

4830-1119: Was not sampled. The placement of the forest bittern ( Zonerodius heliosylus ) was ambiguous, but the results suggest that it is probably closely related to members of the genus Ardeola rather than to the subfamily Tigriornithinae . Tigriornis – white-crested tiger heron Tigrisoma – tiger herons (3 species) Cochlearius – boat-billed heron Agamia – agami heron Zebrilus – zigzag heron Botaurus – bitterns (14 species of which 1 extinct, includes species formerly placed in Ixobrychus ) Gorsachius – night herons (2 species) Calherodius – white-backed night heron Oroanassa – white-eared night heron Pilherodius – capped heron Syrigma – whistling heron Egretta – herons and egrets (13 species) Nyctanassa – night herons (2 species of which 1 extinct) Nycticorax – night herons (6 species of which 4 extinct) Butorides – herons (3 species) Ardeola – pond herons (6 species) Ardea – herons and egrets (16 species, including cattle egrets) As of August 2024

4900-1148: Was published in 2024. Milvus – kites (3 species - with split of yellow-billed) Haliastur – kites (2 species) Haliaeetus – sea and fish eagles (4 species) Icthyophaga – sea and fish eagles (6 species) Butastur – buzzards (4 species) Ictinia – kites (2 species) Geranospiza – crane hawk Busarellus – black-collared hawk Rostrhamus – snail kite Helicolestes – slender-billed kite Morphnarchus – barred hawk Cryptoleucopteryx – plumbeous hawk Buteogallus – hawks and eagles (9 species) Rupornis – roadside hawk (placement uncertain) Parabuteo – hawks (2 species) Geranoaetus – hawks (3 species) Pseudastur – hawks (3 species) Leucopternis – hawks (3 species) Buteo – hawks and buzzards (28 species) Birds of prey Birds of prey or predatory birds , also known as raptors , are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals , reptiles and other smaller birds). In addition to speed and strength, these predators have keen eyesight for detecting prey from

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