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128-566: A bumblebee (or bumble bee , bumble-bee , or humble-bee ) is any of over 250 species in the genus Bombus , part of Apidae , one of the bee families. This genus is the only extant group in the tribe Bombini , though a few extinct related genera (e.g., Calyptapis ) are known from fossils . They are found primarily in higher altitudes or latitudes in the Northern Hemisphere , although they are also found in South America, where

256-460: A de facto official language of Namibia after the end of German colonial rule alongside English and Afrikaans , and had de jure co-official status from 1984 until its independence from South Africa in 1990. However, the Namibian government perceived Afrikaans and German as symbols of apartheid and colonialism, and decided English would be the sole official language upon independence, stating that it

384-402: A second language , and 75–100   million as a foreign language . This would imply the existence of approximately 175–220   million German speakers worldwide. German sociolinguist Ulrich Ammon estimated a number of 289 million German foreign language speakers without clarifying the criteria by which he classified a speaker. As of 2012 , about 90   million people, or 16% of

512-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

640-587: A "commonly used" language and the Pan South African Language Board is obligated to promote and ensure respect for it. Cameroon was also a colony of the German Empire from the same period (1884 to 1916). However, German was replaced by French and English, the languages of the two successor colonial powers, after its loss in World War I . Nevertheless, since the 21st century, German has become

768-560: A bare shiny area surrounded by a fringe of hairs used to transport pollen , whereas in cuckoo bumblebees, the hind leg is hairy all round, and they never carry pollen. Like their relatives the honeybees, bumblebees feed on nectar , using their long hairy tongues to lap up the liquid; the proboscis is folded under the head during flight. Bumblebees gather nectar to add to the stores in the nest, and pollen to feed their young. They forage using colour and spatial relationships to identify flowers to feed from. Some bumblebees steal nectar , making

896-546: A buzzing insect such as a bumblebee or cockchafer , "dumble" probably imitating the sound of these insects, while " dor " meant "beetle". In On the Origin of Species (1859), Charles Darwin speculated about "humble-bees" and their interactions with other species: I have [...] reason to believe that humble-bees are indispensable to the fertilisation of the heartsease ( Viola tricolor ), for other bees do not visit this flower. From experiments which I have tried, I have found that

1024-1009: A degree of complexity criticised by Williams (2008). The cuckoo bumblebees Psithyrus have sometimes been treated as a separate genus but are now considered to be part of Bombus , in one or more subgenera. Examples of Bombus species include Bombus pauloensis , Bombus dahlbomii , Bombus fervidus , Bombus lapidarius , Bombus ruderatus , and Bombus rupestris . Mendacibombus , 12 species Bombias , 3 species Kallobombus , 1 species Orientalibombus , 3 species Subterraneobombus , 10 species Megabombus , 22 species Thoracobombus , 50 species Psithyrus , 30 species Pyrobombus , 50 species Alpinobombus , 5 species Bombus (subgenus) , 5 species Alpigenobombus , 7 species Melanobombus , 17 species Sibiricobombus , 7 species Cullumanobombus , 23 species Bumblebees vary in appearance, but are generally plump and densely furry. They are larger, broader and stouter-bodied than honeybees, and their abdomen tip

1152-407: A few lowland tropical species exist. A few species ( B. polaris and B. alpinus ) range into very cold climates where other bees might not be found; B. polaris occurs in northern Ellesmere Island in the high Arctic, along with another bumblebee B. hyperboreus , which parasitises its nest. This is the northernmost occurrence of any eusocial insect. One reason for their presence in cold places

1280-512: A few lowland tropical species have been identified. European bumblebees have also been introduced to New Zealand and Tasmania . Female bumblebees can sting repeatedly, but generally ignore humans and other animals. Most bumblebees are social insects that form colonies with a single queen. The colonies are smaller than those of honey bees , growing to as few as 50 individuals in a nest. Cuckoo bumblebees are brood parasitic and do not make nests or form colonies; their queens aggressively invade

1408-556: A greater need for regularity in written conventions. While the major changes of the MHG period were socio-cultural, High German was still undergoing significant linguistic changes in syntax, phonetics, and morphology as well (e.g. diphthongization of certain vowel sounds: hus (OHG & MHG "house") → haus (regionally in later MHG)→ Haus (NHG), and weakening of unstressed short vowels to schwa [ə]: taga (OHG "days")→ tage (MHG)). A great wealth of texts survives from

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1536-576: A hole near the base of a flower to access the nectar while avoiding pollen transfer. Bumblebees are important agricultural pollinators , so their decline in Europe, North America, and Asia is a cause for concern. The decline has been caused by habitat loss , the mechanisation of agriculture, and pesticides . The word "bumblebee" is a compound of "bumble" and "bee"—'bumble' meaning to hum, buzz, drone, or move ineptly or flounderingly. The generic name Bombus , assigned by Pierre André Latreille in 1802,

1664-643: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

1792-621: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

1920-733: A magnet, indicating the importance of social information. The bees did not copy one another exactly: in fact, the study suggested that the bees were instead attempting to emulate one another's goals . Nest size depends on species of bumblebee. Most form colonies of between 50 and 400 individuals, but colonies have been documented as small as ~20 individuals and as large as 1700. These nests are small compared to honeybee hives, which hold about 50,000 bees. Many species nest underground, choosing old rodent burrows or sheltered places, and avoiding places that receive direct sunlight that could result in overheating. Other species make nests above ground, whether in thick grass or in holes in trees. A bumblebee nest

2048-453: A native tongue today, mostly descendants of German colonial settlers . The period of German colonialism in Namibia also led to the evolution of a Standard German-based pidgin language called " Namibian Black German ", which became a second language for parts of the indigenous population. Although it is nearly extinct today, some older Namibians still have some knowledge of it. German remained

2176-679: A popular foreign language among pupils and students, with 300,000 people learning or speaking German in Cameroon in 2010 and over 230,000 in 2020. Today Cameroon is one of the African countries outside Namibia with the highest number of people learning German. In the United States, German is the fifth most spoken language in terms of native and second language speakers after English, Spanish , French , and Chinese (with figures for Cantonese and Mandarin combined), with over 1 million total speakers. In

2304-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

2432-463: A region often resemble each other in mutually protective Müllerian mimicry . Harmless insects such as hoverflies often derive protection from resembling bumblebees, in Batesian mimicry , and may be confused with them. Nest-making bumblebees can be distinguished from similarly large, fuzzy cuckoo bumblebees by the form of the female hind leg. In nesting bumblebees, it is modified to form a pollen basket ,

2560-475: A result, the surviving texts are written in highly disparate regional dialects and exhibit significant Latin influence, particularly in vocabulary. At this point monasteries, where most written works were produced, were dominated by Latin, and German saw only occasional use in official and ecclesiastical writing. While there is no complete agreement over the dates of the Middle High German (MHG) period, it

2688-562: A suction cup and during lapping, nectar may be drawn up the proboscis by capillary action . When at rest or flying, the proboscis is kept folded under the head. The longer the tongue, the deeper the bumblebee can probe into a flower and bees probably learn from experience which flower source is best-suited to their tongue length. Bees with shorter proboscides, like Bombus bifarius , have a more difficult time foraging nectar relative to other bumblebees with longer proboscides; to overcome this disadvantage, B. bifarius workers were observed to lick

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2816-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

2944-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

3072-580: Is a recognized minority language in the following countries: In France, the High German varieties of Alsatian and Moselle Franconian are identified as " regional languages ", but the European Charter for Regional or Minority Languages of 1998 has not yet been ratified by the government. Namibia also was a colony of the German Empire, from 1884 to 1915. About 30,000 people still speak German as

3200-418: Is added, they are more likely to go out to forage. Bumblebees have been observed to partake in social learning . In a 2017 study involving Bombus terrestris , bees were taught to complete an unnatural task of moving large objects to obtain a reward. Bees who first observed another bee complete the task were significantly more successful in learning the task than bees who observed the same action performed by

3328-547: Is also notable for its broad spectrum of dialects , with many varieties existing in Europe and other parts of the world. Some of these non-standard varieties have become recognized and protected by regional or national governments. Since 2004, heads of state of the German-speaking countries have met every year, and the Council for German Orthography has been the main international body regulating German orthography . German

3456-546: Is an Indo-European language that belongs to the West Germanic group of the Germanic languages . The Germanic languages are traditionally subdivided into three branches: North Germanic , East Germanic , and West Germanic . The first of these branches survives in modern Danish , Swedish , Norwegian , Faroese , and Icelandic , all of which are descended from Old Norse . The East Germanic languages are now extinct, and Gothic

3584-472: Is called the "German Sprachraum ". German is the official language of the following countries: German is a co-official language of the following countries: Although expulsions and (forced) assimilation after the two World wars greatly diminished them, minority communities of mostly bilingual German native speakers exist in areas both adjacent to and detached from the Sprachraum. Within Europe, German

3712-430: Is complicated by the existence of several varieties whose status as separate "languages" or "dialects" is disputed for political and linguistic reasons, including quantitatively strong varieties like certain forms of Alemannic and Low German . With the inclusion or exclusion of certain varieties, it is estimated that approximately 90–95 million people speak German as a first language , 10–25   million speak it as

3840-745: Is derived from the Latin word for a buzzing or humming sound, borrowed from Ancient Greek βόμβος ( bómbos ). According to the Oxford English Dictionary (OED), the term "bumblebee" was first recorded as having been used in the English language in the 1530 work Lesclarcissement by John Palsgrave , "I bomme, as a bombyll bee dothe." However the OED also states that the term "humblebee" predates it, having first been used in 1450 in Fysshynge wyth Angle , "In Juyll

3968-596: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

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4096-520: Is generally seen as ending when the 1346–53 Black Death decimated Europe's population. Modern High German begins with the Early New High German (ENHG) period, which Wilhelm Scherer dates 1350–1650, terminating with the end of the Thirty Years' War . This period saw the further displacement of Latin by German as the primary language of courtly proceedings and, increasingly, of literature in

4224-595: Is generally seen as lasting from 1050 to 1350. This was a period of significant expansion of the geographical territory occupied by Germanic tribes, and consequently of the number of German speakers. Whereas during the Old High German period the Germanic tribes extended only as far east as the Elbe and Saale rivers, the MHG period saw a number of these tribes expanding beyond this eastern boundary into Slavic territory (known as

4352-559: Is less closely related to languages based on Low Franconian dialects (e.g., Dutch and Afrikaans), Low German or Low Saxon dialects (spoken in northern Germany and southern Denmark ), neither of which underwent the High German consonant shift. As has been noted, the former of these dialect types is Istvaeonic and the latter Ingvaeonic, whereas the High German dialects are all Irminonic; the differences between these languages and standard German are therefore considerable. Also related to German are

4480-461: Is more rounded. Many species have broad bands of colour, the patterns helping to distinguish different species. Whereas honeybees have short tongues and therefore mainly pollinate open flowers, some bumblebee species have long tongues and collect nectar from flowers that are closed into a tube. Bumblebees have fewer stripes (or none), and usually have part of the body covered in black fur, while honeybees have many stripes including several grey stripes on

4608-505: Is not organised into hexagonal combs like that of a honeybee; the cells are instead clustered together untidily. The workers remove dead bees or larvae from the nest and deposit them outside the nest entrance, helping to prevent disease. Nests in temperate regions last only for a single season and do not survive the winter. In the early spring, the queen comes out of diapause and finds a suitable place to create her colony. Then she builds wax cells in which to lay her eggs which were fertilised

4736-435: Is one of the major languages of the world . German is the second-most widely spoken Germanic language , after English, both as a first and as a second language . German is also widely taught as a foreign language , especially in continental Europe (where it is the third most taught foreign language after English and French), and in the United States. Overall, German is the fourth most commonly learned second language, and

4864-587: Is one of the three biggest newspapers in Namibia and the only German-language daily in Africa. An estimated 12,000 people speak German or a German variety as a first language in South Africa, mostly originating from different waves of immigration during the 19th and 20th centuries. One of the largest communities consists of the speakers of "Nataler Deutsch", a variety of Low German concentrated in and around Wartburg . The South African constitution identifies German as

4992-401: Is partly derived from Latin and Greek , along with fewer words borrowed from French and Modern English . English, however, is the main source of more recent loanwords . German is a pluricentric language ; the three standardized variants are German , Austrian , and Swiss Standard German . Standard German is sometimes called High German , which refers to its regional origin. German

5120-677: Is present from the fossil record. An additional three species, "Bombus" luianus , "Bombus" anacolus and "Bombus" dilectus have been attributed to Bombus from the Middle Miocene Shanwang formation of China by Zhang, ( 1990 ) and Zhang et al ( 1994 ). Due to not being able to study Zhang's type specimens , but only illustrations of the fossils, Dehon et al did not place the three species within any specific subgenera, and considered all three as "species inquirenda", needing fuller re-examination. Two other species were not examined at all by Dehon et al , Bombus? crassipes of

5248-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

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5376-555: Is that bumblebees can regulate their body temperature , via solar radiation , internal mechanisms of "shivering" (called heterothermy ), and countercurrent exchange to retain heat. Other bees have similar physiology , but the mechanisms seem best developed and have been most studied in bumblebees. They adapt to higher elevations by extending their wing stroke amplitude. Bumblebees have a largely cosmopolitan distribution but are absent from Australia (apart from Tasmania where they have been introduced) and are found in Africa only north of

5504-510: Is the Sachsenspiegel , the first book of laws written in Middle Low German ( c.  1220 ). The abundance and especially the secular character of the literature of the MHG period demonstrate the beginnings of a standardized written form of German, as well as the desire of poets and authors to be understood by individuals on supra-dialectal terms. The Middle High German period

5632-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

5760-750: Is the most spoken native language within the European Union . It is the most widely spoken and official (or co-official) language in Germany , Austria , Switzerland , Liechtenstein , and the Italian autonomous province of South Tyrol . It is also an official language of Luxembourg , Belgium and the Italian autonomous region of Friuli-Venezia Giulia , as well as a recognized national language in Namibia . There are also notable German-speaking communities in France ( Alsace ),

5888-566: Is the only language in this branch which survives in written texts. The West Germanic languages, however, have undergone extensive dialectal subdivision and are now represented in modern languages such as English, German, Dutch , Yiddish , Afrikaans , and others. Within the West Germanic language dialect continuum, the Benrath and Uerdingen lines (running through Düsseldorf - Benrath and Krefeld - Uerdingen , respectively) serve to distinguish

6016-481: Is understood in all areas where German is spoken. Approximate distribution of native German speakers (assuming a rounded total of 95 million) worldwide: As a result of the German diaspora , as well as the popularity of German taught as a foreign language , the geographical distribution of German speakers (or "Germanophones") spans all inhabited continents. However, an exact, global number of native German speakers

6144-673: The Ostsiedlung ). With the increasing wealth and geographic spread of the Germanic groups came greater use of German in the courts of nobles as the standard language of official proceedings and literature. A clear example of this is the mittelhochdeutsche Dichtersprache employed in the Hohenstaufen court in Swabia as a standardized supra-dialectal written language. While these efforts were still regionally bound, German began to be used in place of Latin for certain official purposes, leading to

6272-627: The Alamanni , Bavarian, and Thuringian groups, all belonging to the Elbe Germanic group ( Irminones ), which had settled in what is now southern-central Germany and Austria between the second and sixth centuries, during the great migration. In general, the surviving texts of Old High German (OHG) show a wide range of dialectal diversity with very little written uniformity. The early written tradition of OHG survived mostly through monasteries and scriptoria as local translations of Latin originals; as

6400-557: The Czech Republic ( North Bohemia ), Poland ( Upper Silesia ), Slovakia ( Košice Region , Spiš , and Hauerland ), Denmark ( North Schleswig ), Romania and Hungary ( Sopron ). Overseas, sizeable communities of German-speakers are found in Brazil ( Blumenau and Pomerode ), South Africa ( Kroondal ), Namibia , among others, some communities have decidedly Austrian German or Swiss German characters (e.g. Pozuzo , Peru). German

6528-522: The Early Middle Ages . German is an inflected language , with four cases for nouns, pronouns, and adjectives (nominative, accusative, genitive, dative); three genders (masculine, feminine, neuter) and two numbers (singular, plural). It has strong and weak verbs . The majority of its vocabulary derives from the ancient Germanic branch of the Indo-European language family, while a smaller share

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6656-417: The European Union 's population, spoke German as their mother tongue, making it the second most widely spoken language on the continent after Russian and the second biggest language in terms of overall speakers (after English), as well as the most spoken native language. The area in central Europe where the majority of the population speaks German as a first language and has German as a (co-)official language

6784-514: The German states . While these states were still part of the Holy Roman Empire , and far from any form of unification, the desire for a cohesive written language that would be understandable across the many German-speaking principalities and kingdoms was stronger than ever. As a spoken language German remained highly fractured throughout this period, with a vast number of often mutually incomprehensible regional dialects being spoken throughout

6912-724: The Holy Roman Emperor Maximilian I , and the other being Meißner Deutsch , used in the Electorate of Saxony in the Duchy of Saxe-Wittenberg . Alongside these courtly written standards, the invention of the printing press led to the development of a number of printers' languages ( Druckersprachen ) aimed at making printed material readable and understandable across as many diverse dialects of German as possible. The greater ease of production and increased availability of written texts brought about increased standardisation in

7040-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

7168-799: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

7296-644: The Old High German language in several Elder Futhark inscriptions from as early as the sixth century AD (such as the Pforzen buckle ), the Old High German period is generally seen as beginning with the Abrogans (written c.  765–775 ), a Latin-German glossary supplying over 3,000 Old High German words with their Latin equivalents. After the Abrogans , the first coherent works written in Old High German appear in

7424-640: The Sprachraum in Europe. German is used in a wide variety of spheres throughout the country, especially in business, tourism, and public signage, as well as in education, churches (most notably the German-speaking Evangelical Lutheran Church in Namibia (GELK) ), other cultural spheres such as music, and media (such as German language radio programs by the Namibian Broadcasting Corporation ). The Allgemeine Zeitung

7552-536: The Standard German language in its written form, and the Duden Handbook was declared its standard definition. Punctuation and compound spelling (joined or isolated compounds) were not standardized in the process. The Deutsche Bühnensprache ( lit.   ' German stage language ' ) by Theodor Siebs had established conventions for German pronunciation in theatres , three years earlier; however, this

7680-503: The Upper German dialects spoken in the southern German-speaking countries , such as Swiss German ( Alemannic dialects ) and the various Germanic dialects spoken in the French region of Grand Est , such as Alsatian (mainly Alemannic, but also Central–and   Upper Franconian dialects) and Lorraine Franconian (Central Franconian). After these High German dialects, standard German

7808-462: The pagan Germanic tradition. Of particular interest to scholars, however, has been the Hildebrandslied , a secular epic poem telling the tale of an estranged father and son unknowingly meeting each other in battle. Linguistically, this text is highly interesting due to the mixed use of Old Saxon and Old High German dialects in its composition. The written works of this period stem mainly from

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7936-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

8064-714: The Empire. Its use indicated that the speaker was a merchant or someone from an urban area, regardless of nationality. Prague (German: Prag ) and Budapest ( Buda , German: Ofen ), to name two examples, were gradually Germanized in the years after their incorporation into the Habsburg domain; others, like Pressburg ( Pozsony , now Bratislava), were originally settled during the Habsburg period and were primarily German at that time. Prague, Budapest, Bratislava, and cities like Zagreb (German: Agram ) or Ljubljana (German: Laibach ), contained significant German minorities. In

8192-420: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

8320-718: The Frisian languages— North Frisian (spoken in Nordfriesland ), Saterland Frisian (spoken in Saterland ), and West Frisian (spoken in Friesland )—as well as the Anglic languages of English and Scots. These Anglo-Frisian dialects did not take part in the High German consonant shift, and the Anglic languages also adopted much vocabulary from both Old Norse and the Norman language . The history of

8448-524: The German language begins with the High German consonant shift during the Migration Period , which separated Old High German dialects from Old Saxon . This sound shift involved a drastic change in the pronunciation of both voiced and voiceless stop consonants ( b , d , g , and p , t , k , respectively). The primary effects of the shift were the following below. While there is written evidence of

8576-447: The German states; the invention of the printing press c.  1440 and the publication of Luther's vernacular translation of the Bible in 1534, however, had an immense effect on standardizing German as a supra-dialectal written language. The ENHG period saw the rise of several important cross-regional forms of chancery German, one being gemeine tiutsch , used in the court of

8704-675: The Germanic dialects that were affected by the High German consonant shift (south of Benrath) from those that were not (north of Uerdingen). The various regional dialects spoken south of these lines are grouped as High German dialects, while those spoken to the north comprise the Low German and Low Franconian dialects. As members of the West Germanic language family, High German, Low German, and Low Franconian have been proposed to be further distinguished historically as Irminonic , Ingvaeonic , and Istvaeonic , respectively. This classification indicates their historical descent from dialects spoken by

8832-646: The Irminones (also known as the Elbe group), Ingvaeones (or North Sea Germanic group), and Istvaeones (or Weser–Rhine group). Standard German is based on a combination of Thuringian - Upper Saxon and Upper Franconian dialects, which are Central German and Upper German dialects belonging to the High German dialect group. German is therefore closely related to the other languages based on High German dialects, such as Luxembourgish (based on Central Franconian dialects ) and Yiddish . Also closely related to Standard German are

8960-616: The Late Miocene Krottensee deposits in the Czech Republic, and Bombus proavus from the Middle Miocene Latah Formation , USA. The genus Bombus , the only one extant genus in the tribe Bombini, comprises over 250 species; for an overview of the differences between bumblebees and other bees and wasps, see characteristics of common wasps and bees . The genus has been divided variously into up to 49 subgenera,

9088-507: The MHG period. Significantly, these texts include a number of impressive secular works, such as the Nibelungenlied , an epic poem telling the story of the dragon -slayer Siegfried ( c.  thirteenth century ), and the Iwein , an Arthurian verse poem by Hartmann von Aue ( c.  1203 ), lyric poems , and courtly romances such as Parzival and Tristan . Also noteworthy

9216-566: The Meliponini, which have somewhat more advanced eusocial behaviour. Sophie Cardinal and Bryan Danforth comment that "While remarkable, a hypothesis of dual origins of advanced eusociality is congruent with early studies on corbiculate morphology and social behavior." Their analysis, combining molecular , morphological and behavioural data, gives the following cladogram : Apini (honeybees) Euglossini (orchid bees) Bombini (bumblebees) Meliponini (stingless bees) On this hypothesis,

9344-448: The Sahara. More than a hundred years ago they were also introduced to New Zealand, where they play an important role as efficient pollinators. The bumblebee tongue (the proboscis ) is a long, hairy structure that extends from a sheath-like modified maxilla. The primary action of the tongue is lapping, that is, repeated dipping of the tongue into liquid. The tip of the tongue probably acts as

9472-582: The abdomen. Sizes are very variable even within species; the largest British species, B. terrestris , has queens up to 22 mm (0.9 in) long, males up to 16 mm (0.6 in) long, and workers between 11 and 17 mm (0.4–0.7 in) long. The largest bumblebee species in the world is B. dahlbomii of Chile, up to about 40 mm (1.6 in) long, and described as "flying mice" and "a monstrous fluffy ginger beast". Bumblebees are typically found in temperate climates , and are often found at higher latitudes and altitudes than other bees, although

9600-407: The back of spurs on the nectar duct, which resulted in a small reward. The exoskeleton of the abdomen is divided into plates called dorsal tergites and ventral sternites . Wax is secreted from glands on the abdomen and extruded between the sternites where it resembles flakes of dandruff . It is secreted by the queen when she starts a nest and by young workers. It is scraped from the abdomen by

9728-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

9856-734: The bees in the group with a selective advantage. In addition, parasitic (cuckoo) bumblebees resemble their hosts more closely than would be expected by chance, at least in areas like Europe where parasite-host co-speciation is common; but this too may be explained as Müllerian mimicry, rather than requiring the parasite's coloration to deceive the host ( aggressive mimicry ). Bumblebees are active under conditions during which honeybees and other smaller bees stay at home, and can readily absorb heat from even weak sunshine. The thick pile created by long setae (bristles) acts as insulation to keep bumblebees warm in cold weather; species from cold climates have longer setae (and thus thicker insulation) than those from

9984-515: The bumble bee." Since World War II "humblebee" has fallen into near-total disuse. The bumblebee tribe Bombini is one of four groups of corbiculate bees (those with pollen baskets) in the Apidae , the others being the Apini (honey bees), Euglossini ( orchid bees ), and Meliponini (stingless bees). The corbiculate bees are a monophyletic group. Advanced eusocial behaviour appears to have evolved twice in

10112-411: The cold. Bumblebees do not have ears, and it is not known whether or how well they can hear. However, they are sensitive to the vibrations made by sound travelling through wood or other materials. Bumblebees do not exhibit the " bee dances " used by honeybees to tell other workers the locations of food sources. Instead, when they return from a successful foraging expedition, they run excitedly around in

10240-482: The corbiculate bees. They find that Bombini is in fact sister to Meliponini, corroborating that previous finding from Sophie Cardinal and Bryan Danforth (2011). However, Romiguier et al. (2015) shows that Bombini, Meliponini, and Apini form a monophyletic group, where Apini shares a most recent common ancestor with the Bombini and Meliponini clade, while Euglossini is most distantly related to all three, since it does not share

10368-512: The cycle begins in February, reproduction starts in July or August, and ends in the winter months. The queen remains in hibernation until spring of the following year in order to optimize conditions to search for a nest. In fertilised queens, the ovaries only become active when the queen starts to lay. An egg passes along the oviduct to the vagina where there is a chamber called the spermatheca , in which

10496-430: The development of non-local forms of language and exposed all speakers to forms of German from outside their own area. With Luther's rendering of the Bible in the vernacular, German asserted itself against the dominance of Latin as a legitimate language for courtly, literary, and now ecclesiastical subject-matter. His Bible was ubiquitous in the German states: nearly every household possessed a copy. Nevertheless, even with

10624-452: The dialect so as to make the work as natural and accessible to German speakers as possible. Copies of Luther's Bible featured a long list of glosses for each region, translating words which were unknown in the region into the regional dialect. Luther said the following concerning his translation method: One who would talk German does not ask the Latin how he shall do it; he must ask the mother in

10752-511: The different bumblebees in a region resemble each other, so that a young predator need only learn to avoid any of them once. For example, in California a group of bumblebees consists of largely black species including B. californicus , B. caliginosus , B. vandykei , B. vosnesenskii , B. insularis and B. fernaldae . Other bees in California include a group of species all banded black and yellow. In each case, Müllerian mimicry provides

10880-507: The eastern provinces of Banat , Bukovina , and Transylvania (German: Banat, Buchenland, Siebenbürgen ), German was the predominant language not only in the larger towns—like Temeschburg ( Timișoara ), Hermannstadt ( Sibiu ), and Kronstadt ( Brașov )—but also in many smaller localities in the surrounding areas. In 1901, the Second Orthographic Conference ended with a (nearly) complete standardization of

11008-434: The families Syrphidae (hoverflies), Asilidae (robber flies), Tabanidae (horseflies), Oestridae (bot or warble flies) and Bombyliidae (bee flies, such as Bombylius major ) all include Batesian mimics of bumblebees, resembling them closely enough to deceive at least some predators. Many species of Bombus , including the group sometimes called Psithyrus (cuckoo bumblebees), have evolved Müllerian mimicry , where

11136-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

11264-727: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

11392-469: The greshop & the humbylbee in the medow." The latter term was used in A Midsummer Night's Dream ( c.  1600 ) by William Shakespeare , "The honie-bags steale from the humble Bees." Similar terms are used in other Germanic languages, such as the German Hummel ( Old High German humbala ), Dutch hommel or Swedish humla . An old provincial name, "dumbledor", also denoted

11520-406: The group, giving rise to controversy, now largely settled, as to the phylogenetic origins of the four tribes; it had been supposed that eusocial behaviour had evolved only once, requiring the Apini to be close to the Meliponini, which they do not resemble. It is now thought that the Apini (with advanced societies) and Euglossini are closely related, while the primitively eusocial Bombini are close to

11648-435: The home, the children on the streets, the common man in the market-place and note carefully how they talk, then translate accordingly. They will then understand what is said to them because it is German. When Christ says ' ex abundantia cordis os loquitur ,' I would translate, if I followed the papists, aus dem Überflusz des Herzens redet der Mund . But tell me is this talking German? What German understands such stuff? No,

11776-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

11904-474: The influence of Luther's Bible as an unofficial written standard, a widely accepted standard for written German did not appear until the middle of the eighteenth century. German was the language of commerce and government in the Habsburg Empire , which encompassed a large area of Central and Eastern Europe . Until the mid-nineteenth century, it was essentially the language of townspeople throughout most of

12032-628: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

12160-624: The late Eocene Calyptapis florissantensis from the Florissant Formation , USA, and Oligobombus cuspidatus from the Bembridge Marls of the Isle of Wight . Two species of Bombus have been described from the Oligocene of Beşkonak, Bucak Turkey: Bombus (Mendacibombus) beskonakensis and Bombus (Paraelectrobombus) patriciae . Both species were originally placed in genera considered at

12288-480: The legs, moulded until malleable and used in the construction of honeypots, to cover the eggs, to line empty cocoons for use as storage containers and sometimes to cover the exterior of the nest. The brightly coloured pile of the bumblebee is an aposematic (warning) signal, given that females can inflict a painful sting. Depending on the species and morph , the warning colours range from entirely black, to bright yellow, red, orange, white, and pink. Dipteran flies in

12416-504: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. German language German (German: Deutsch , pronounced [dɔʏtʃ] ) is a West Germanic language in the Indo-European language family , mainly spoken in Western and Central Europe . It

12544-402: The lowest chill-coma temperature. Of the bumblebees Bombus bimaculatus has the lowest at 7 °C (45 °F). However, bumblebees have been seen to fly in colder ambient temperatures. This discrepancy is likely because the chill-coma temperature was determined by tests done in a laboratory setting. However, bumblebees live in insulated shelters and can shiver to warm up before venturing into

12672-399: The molecular data suggest that the Bombini are 25 to 40 million years old, while the Meliponini (and thus the clade that includes the Bombini and Meliponini) are 81 to 96 million years old, about the same age as the corbiculate group. However, a more recent phylogeny using transcriptome data from 3,647 genes of ten corbiculate bee species supports the single origin of eusociality hypothesis in

12800-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

12928-402: The mother in the home and the plain man would say, Wesz das Herz voll ist, des gehet der Mund über . Luther's translation of the Bible into High German was also decisive for the German language and its evolution from Early New High German to modern Standard German. The publication of Luther's Bible was a decisive moment in the spread of literacy in early modern Germany , and promoted

13056-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

13184-404: The nest for several minutes before going out to forage once more. These bees may be offering some form of communication based on the buzzing sounds made by their wings, which may stimulate other bees to start foraging. Another stimulant to foraging activity is the level of food reserves in the colony. Bees monitor the amount of honey in the honeypots, and when little is left or when high-quality food

13312-512: The nests of other bumblebee species, kill the resident queens and then lay their own eggs, which are cared for by the resident workers. Cuckoo bumblebees were previously classified as a separate genus, but are now usually treated as members of Bombus . Bumblebees have round bodies covered in soft hair (long branched setae ) called 'pile', making them appear and feel fuzzy. They have aposematic (warning) coloration , often consisting of contrasting bands of colour, and different species of bumblebee in

13440-630: The ninth century, chief among them being the Muspilli , Merseburg charms , and Hildebrandslied , and other religious texts (the Georgslied , Ludwigslied , Evangelienbuch , and translated hymns and prayers). The Muspilli is a Christian poem written in a Bavarian dialect offering an account of the soul after the Last Judgment , and the Merseburg charms are transcriptions of spells and charms from

13568-465: The placement of genus Paraelectrobombus as Bombus subgenus Paraelectrobombus , rather than as a genus in Electrobombini . The subgenus Cullumanobombus was expanded to include not only Bombus trophonius but also Bombus randeckensis which was moved from subgenus Bombus and Bombus pristinus , first described by Unger ( 1867 ). Within the subgenus Melanobombus only Bombus cerdanyensis

13696-404: The previous year. The eggs that hatch develop into female workers, and in time, the queen populates the colony, with workers feeding the young and performing other duties similar to honeybee workers. In temperate zones , young queens ( gynes ) leave the nest in the autumn and mate , often more than once, with males ( drones ) that are forcibly driven out of the colony. The drones and workers die as

13824-462: The pronunciation of the ending -ig as [ɪk] instead of [ɪç]. In Northern Germany, High German was a foreign language to most inhabitants, whose native dialects were subsets of Low German. It was usually encountered only in writing or formal speech; in fact, most of High German was a written language, not identical to any spoken dialect, throughout the German-speaking area until well into the 19th century. However, wider standardization of pronunciation

13952-526: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

14080-427: The same most recent common ancestor as Bombini, Meliponini, and Apini. Thus, their analysis supports the single origin of eusociality hypothesis within the corbiculate bees, where eusociality evolved in the common ancestor of Bombini, Apini, and Meliponini. The fossil record for bees is limited, with around 14 species that might possibly be Bombini having been described by 2019. The only Bombus relatives in Bombini are

14208-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

14336-606: The sperm from the mating is stored. Depending on need, she may allow her egg to be fertilised. Unfertilised eggs become haploid males; fertilised eggs grow into diploid females and queens. The hormones that stimulate the development of the ovaries are suppressed in female worker bees, while the queen remains dominant. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

14464-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

14592-541: The states of North Dakota and South Dakota , German is the most common language spoken at home after English. As a legacy of significant German immigration to the country , German geographical names can be found throughout the Midwest region , such as New Ulm and Bismarck (North Dakota's state capital), plus many other regions. A number of German varieties have developed in the country and are still spoken today, such as Pennsylvania Dutch and Texas German . In Brazil,

14720-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

14848-496: The third most commonly learned second language in the United States in K-12 education. The language has been influential in the fields of philosophy, theology, science, and technology. It is the second most commonly used language in science and the third most widely used language on websites . The German-speaking countries are ranked fifth in terms of annual publication of new books, with one-tenth of all books (including e-books) in

14976-521: The time of description as outside of Bombus , being initially named Oligoapis beskonakensis and Paraelectrobombus patriciae respectively, however reexaminiation of the fore-wings lead to both being considered as Bombus species In 2012 a fossil bumblebee from the Miocene was found in Germany's Randeck Maar and classified as Bombus (Bombus) randeckensis . In 2014, another species, Bombus cerdanyensis ,

15104-512: The tropics. The temperature of the flight muscles, which occupy much of the thorax, needs to be at least 30 °C (86 °F) before flight can take place. The muscle temperature can be raised by shivering. It takes about five minutes for the muscles to reach this temperature at an air temperature of 13 °C (55 °F). The chill-coma temperature in relation to flying insects is the temperature at which flight muscles cannot be activated. Compared to honey bees and carpenter bees, bumblebees have

15232-566: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

15360-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

15488-465: The visits of bees, if not indispensable, are at least highly beneficial to the fertilisation of our clovers; but humble-bees alone visit the common red clover ( Trifolium pratense ), as other bees cannot reach the nectar. However, "bumblebee" remained in use, for example in The Tale of Mrs. Tittlemouse (1910) by Beatrix Potter , "Suddenly round a corner, she met Babbitty Bumble--"Zizz, Bizz, Bizzz!" said

15616-454: The weather turns colder; the young queens feed intensively to build up stores of fat for the winter. They survive in a resting state (diapause), generally below ground, until the weather warms up in the spring with the early bumblebee being the species that is among the first to emerge. Many species of bumblebee follow this general trend within the year. Bombus pensylvanicus is a species that follows this type of colony cycle. For this species

15744-543: The world being published in German. German is most closely related to other West Germanic languages, namely Afrikaans , Dutch , English , the Frisian languages , and Scots . It also contains close similarities in vocabulary to some languages in the North Germanic group , such as Danish , Norwegian , and Swedish . Modern German gradually developed from Old High German , which in turn developed from Proto-Germanic during

15872-579: The written form of German. One of the central events in the development of ENHG was the publication of Luther's translation of the Bible into High German (the New Testament was published in 1522; the Old Testament was published in parts and completed in 1534). Luther based his translation primarily on the Meißner Deutsch of Saxony , spending much time among the population of Saxony researching

16000-429: Was a "neutral" language as there were virtually no English native speakers in Namibia at that time. German, Afrikaans, and several indigenous languages thus became "national languages" by law, identifying them as elements of the cultural heritage of the nation and ensuring that the state acknowledged and supported their presence in the country. Today, Namibia is considered to be the only German-speaking country outside of

16128-530: Was an artificial standard that did not correspond to any traditional spoken dialect. Rather, it was based on the pronunciation of German in Northern Germany, although it was subsequently regarded often as a general prescriptive norm, despite differing pronunciation traditions especially in the Upper-German-speaking regions that still characterise the dialect of the area today – especially

16256-502: Was described from Late Miocene lacustrine beds of La Cerdanya, Spain, but not initially placed into any subgenus, The species Bombus trophonius was described in October 2017 and placed in Bombus subgenus Cullumanobombus . A redescription of the Bombini fossil record by Dehon et al ( 2019 ) resulted in the synonymization of the genus Oligoapis with Bombus subgenus Mendacibombus , and

16384-485: Was established on the basis of public speaking in theatres and the media during the 20th century and documented in pronouncing dictionaries. Official revisions of some of the rules from 1901 were not issued until the controversial German orthography reform of 1996 was made the official standard by governments of all German-speaking countries. Media and written works are now almost all produced in Standard German which

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